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<mods:title id="BC24F3BAE457422B21A276F90D93E406">Review of the genus Holostaspis Kolenati, 1858 (Acari: Laelapidae)</mods:title>
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<paragraph id="8BDA36AAFF86423B3D8A17BA9E006C86" blockId="23.[151,614,1446,1506]" box="[151,614,1446,1472]" pageId="23" pageNumber="324">
<heading id="D09281C6FF86423B3D8A17BA9E006C86" bold="true" box="[151,614,1446,1472]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<taxonomicName id="4C654D29FF86423B3D8A17BA9E006C86" authority="(Wasmann, 1897)" baseAuthorityName="Wasmann" baseAuthorityYear="1897" box="[151,614,1446,1472]" class="Arachnida" family="Laelapidae" genus="Holostaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophila">
<emphasis id="B911EAB8FF86423B3D8A17BA9E006C86" bold="true" box="[151,614,1446,1472]" pageId="23" pageNumber="324">
<emphasis id="B911EAB8FF86423B3D8A17BA9DE06C86" bold="true" box="[151,390,1446,1472]" italics="true" pageId="23" pageNumber="324">Holostaspis oophila</emphasis>
(
<bibRefCitation id="EFF44B5BFF86423B3C8917BA9E386C86" author="Wasmann, E." box="[404,606,1446,1472]" pageId="23" pageNumber="324" pagination="170 - 173" refId="ref26079" refString="Wasmann, E. (1897) Uber einige myrmecophile Acarinen. Zoologischer Anzeiger, 20, 170 - 173." type="journal article" year="1897">Wasmann, 1897</bibRefCitation>
)
</emphasis>
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<paragraph id="8BDA36AAFF86423B3D8A17D59D416CA4" blockId="23.[151,614,1446,1506]" box="[151,295,1481,1506]" pageId="23" pageNumber="324">
(
<figureCitation id="135E2A2FFF86423B3D8217D59D786CA4" box="[159,286,1481,1506]" captionStart-0="FIGURES 5762" captionStart-1="FIGURES 6366" captionStartId-0="25.[151,264,1940,1963]" captionStartId-1="26.[151,264,1589,1612]" captionTargetBox-0="[190,1408,203,1923]" captionTargetBox-1="[187,1390,254,1473]" captionTargetId-0="figure@25.[182,1412,193,1931]" captionTargetId-1="figure@26.[151,1436,193,1586]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." captionText-1="FIGURES 6366. Holostaspis oophila, female. 63. Leg I; 64. Leg II; 65. Leg III; 66. Leg IV." httpUri-0="https://zenodo.org/record/2653390/files/figure.png" httpUri-1="https://zenodo.org/record/2653392/files/figure.png" pageId="23" pageNumber="324">Figs 5766</figureCitation>
)
</paragraph>
<paragraph id="8BDA36AAFF86423B3D8A140D9F0B6F61" blockId="23.[151,1436,1553,1820]" box="[151,877,1553,1575]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB751184FF86423B3D8A140D9F0B6F61" box="[151,877,1553,1575]" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A140D9DFE6F61" authority="Moniez" authorityName="Moniez" box="[151,408,1553,1575]" class="Arachnida" family="Uropodidae" genus="Uropoda" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophila">
<emphasis id="B911EAB8FF86423B3D8A140D9D236F61" box="[151,325,1553,1575]" italics="true" pageId="23" pageNumber="324">Uropoda oophila</emphasis>
Moniez
</taxonomicName>
(
<emphasis id="B911EAB8FF86423B3CBA140E9E506F61" box="[423,566,1553,1575]" italics="true" pageId="23" pageNumber="324">nomen nudum</emphasis>
).—
<treatmentCitation id="0AC410BBFF86423B3F46140E9F506F61" author="Wasmann, E." box="[603,822,1553,1575]" page="197" pageId="23" pageNumber="324" year="1894">
<bibRefCitation id="EFF44B5BFF86423B3F46140E9F626F61" author="Wasmann, E." box="[603,772,1553,1575]" pageId="23" pageNumber="324" refId="ref26028" refString="Wasmann, E. (1894) Kritisches Verzeichniss der myrmekophilen und termitophilen Arthropoden. Mit Angabe der Lebensweise und mit Beschriebung neuer Arten. Verlag von Felix L. Dames, Berlin, 231 pp. https: // doi. org / 10.5962 / bhl. title. 122977" type="book" year="1894">Wasmann, 1894</bibRefCitation>
: 197
</treatmentCitation>
, 199.
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</paragraph>
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<paragraph id="8BDA36AAFF86423B3D8A142C9E4E6F00" blockId="23.[151,1436,1553,1820]" box="[151,552,1584,1606]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB751184FF86423B3D8A142C9E4E6F00" box="[151,552,1584,1606]" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A142C9E426F00" authority="Wasmann, 1897: 172" authorityName="Wasmann" authorityPageNumber="172" authorityYear="1897" box="[151,548,1584,1606]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophilus">
<emphasis id="B911EAB8FF86423B3D8A142C9D226F00" box="[151,324,1584,1606]" italics="true" pageId="23" pageNumber="324">Laelaps oophilus</emphasis>
<treatmentCitation id="0AC410BBFF86423B3C56142D9E426F00" author="Wasmann, E." box="[331,548,1584,1606]" page="172" pageId="23" pageNumber="324" year="1897">
<bibRefCitation id="EFF44B5BFF86423B3C56142D9D976F00" author="Wasmann, E." box="[331,497,1584,1606]" pageId="23" pageNumber="324" pagination="170 - 173" refId="ref26079" refString="Wasmann, E. (1897) Uber einige myrmecophile Acarinen. Zoologischer Anzeiger, 20, 170 - 173." type="journal article" year="1897">Wasmann, 1897</bibRefCitation>
: 172
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BDA36AAFF86423B3D8A145398436F22" blockId="23.[151,1436,1553,1820]" box="[151,1061,1614,1636]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB751184FF86423B3D8A145398436F22" box="[151,1061,1614,1636]" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A14539D336F22" box="[151,341,1614,1636]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophila">
<emphasis id="B911EAB8FF86423B3D8A14539D336F22" box="[151,341,1614,1636]" italics="true" pageId="23" pageNumber="324">Hypoaspis oophila</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC410BBFF86423B3C6E14529E2F6F22" author="Oudemans, A. C." box="[371,585,1614,1636]" page="24" pageId="23" pageNumber="324" year="1902">
<bibRefCitation id="EFF44B5BFF86423B3C6E14529E426F22" author="Oudemans, A. C." box="[371,548,1614,1636]" pageId="23" pageNumber="324" pagination="1 - 52" refId="ref24954" refString="Oudemans, A. C. (1902) New list of Dutch Acari. Second part. With remarks on known and descriptions of a new subfamily, new genera and species. Tijdschrift voor Entomologie, 45 (1 / 2), 1 - 52." type="journal article" year="1902">Oudemans, 1902</bibRefCitation>
: 24
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B3F4914529F506F22" author="Oudemans, A. C." box="[596,822,1614,1636]" page="130" pageId="23" pageNumber="324" year="1903">
<bibRefCitation id="EFF44B5BFF86423B3F4914529F656F22" author="Oudemans, A. C." box="[596,771,1614,1636]" pageId="23" pageNumber="324" pagination="123 - 150" refId="ref25003" refString="Oudemans, A. C. (1903) Notes on Acari. Fifth series. Tijdschrift voor Entomologie, 45 (3 / 4), 123 - 150." type="journal article" year="1903">Oudemans, 1903</bibRefCitation>
: 130
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B3E5D145298476F22" author="Buitendijk, A. M." box="[832,1057,1614,1636]" page="296" pageId="23" pageNumber="324" year="1945">
<bibRefCitation id="EFF44B5BFF86423B3E5D14529F886F22" author="Buitendijk, A. M." box="[832,1006,1614,1636]" pageId="23" pageNumber="324" pagination="281 - 391" refId="ref21362" refString="Buitendijk, A. M. (1945) Voorloopige catalogus van de Acari in de collectie-Oudemans. Zoologische Mededeelingen, 24, 281 - 391." type="journal article" year="1945">Buitendijk, 1945</bibRefCitation>
: 296
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BDA36AAFF86423B3D8A14719EC36FC5" blockId="23.[151,1436,1553,1820]" box="[151,677,1645,1667]" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A14719DA76FC5" authorityName="Wasmann" authorityYear="1897" box="[151,449,1645,1667]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophilus" subGenus="Eulaelaps">
<emphasis id="B911EAB8FF86423B3D8A14719C816FC5" box="[151,231,1645,1667]" italics="true" pageId="23" pageNumber="324">Laelaps</emphasis>
(
<emphasis id="B911EAB8FF86423B3DEB14719D3D6FC5" box="[246,347,1645,1667]" italics="true" pageId="23" pageNumber="324">Eulaelaps</emphasis>
)
<emphasis id="B911EAB8FF86423B3C7714719DA76FC5" box="[362,449,1645,1667]" italics="true" pageId="23" pageNumber="324">oophilus</emphasis>
</taxonomicName>
.—Berlese, 1903b: 13.
</paragraph>
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<paragraph id="8BDA36AAFF86423B3D8A14909ED06FE4" blockId="23.[151,1436,1553,1820]" box="[151,694,1676,1698]" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A14909DA56FE4" authorityName="Wasmann" authorityYear="1897" box="[151,451,1676,1698]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophilus" subGenus="Oolaelaps">
<emphasis id="B911EAB8FF86423B3D8A14909C816FE4" box="[151,231,1676,1698]" italics="true" pageId="23" pageNumber="324">Laelaps</emphasis>
(
<emphasis id="B911EAB8FF86423B3DEB14909D386FE4" box="[246,350,1676,1698]" italics="true" pageId="23" pageNumber="324">Oolaelaps</emphasis>
)
<emphasis id="B911EAB8FF86423B3C7114909DA56FE4" box="[364,451,1676,1698]" italics="true" pageId="23" pageNumber="324">oophilus</emphasis>
</taxonomicName>
.—Berlese, 1904b: 428.
</paragraph>
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<taxonomicName id="4C654D29FF86423B3D8A14B69D3A6F86" box="[151,348,1706,1728]" class="Arachnida" genus="Oolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophilus">
<emphasis id="B911EAB8FF86423B3D8A14B69D3A6F86" box="[151,348,1706,1728]" italics="true" pageId="23" pageNumber="324">Oolaelaps oophilus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC410BBFF86423B3C6714B69E376F86" author="Oudemans, A. C." box="[378,593,1706,1728]" page="68" pageId="23" pageNumber="324" year="1914">
<bibRefCitation id="EFF44B5BFF86423B3C6714B69E4A6F86" author="Oudemans, A. C." box="[378,556,1706,1728]" pageId="23" pageNumber="324" pagination="65 - 73" refId="ref25034" refString="Oudemans, A. C. (1914) Acarologische Aanteekeningen, LII. Entomologische Berichten, 76, 65 - 73." type="journal article" year="1914">Oudemans, 1914</bibRefCitation>
: 68
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B3F4614B69E976F86" author="Hull, R. J." box="[603,753,1706,1728]" page="69" pageId="23" pageNumber="324" year="1918">
<bibRefCitation id="EFF44B5BFF86423B3F4614B69EAD6F86" author="Hull, R. J." box="[603,715,1706,1728]" pageId="23" pageNumber="324" pagination="13 - 88" refId="ref22865" refString="Hull, R. J. (1918) Terrestrial Acari of the Tyne Province. Transactions of the Natural History Society of Northumberland, 5, 13 - 88." type="journal article" year="1918">Hull, 1918</bibRefCitation>
: 69
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B3FE114B6980C6F86" author="Castagnoli, M. &amp; Pegazzano, F." box="[764,1130,1706,1728]" page="286" pageId="23" pageNumber="324" year="1985">
<bibRefCitation id="EFF44B5BFF86423B3FE114B6985E6F86" author="Castagnoli, M. &amp; Pegazzano, F." box="[764,1080,1706,1728]" pageId="23" pageNumber="324" refId="ref21536" refString="Castagnoli, M. &amp; Pegazzano, F. (1985) Catalogue of the Berlese Acaroteca. Istituto Sperimentale per la Zoologia Agraria, Firenze, Italy, 490 pp." type="book" year="1985">Castagnoli &amp; Pegazzano, 1985</bibRefCitation>
: 286
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.
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</paragraph>
<paragraph id="8BDA36AAFF86423B3D8A14D69D526FB8" blockId="23.[151,1436,1553,1820]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB751184FF86423B3D8A14D69D526FB8" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A14D69D8E6F99" box="[151,488,1737,1759]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophila" subGenus="Holostaspis">
<emphasis id="B911EAB8FF86423B3D8A14D69D646F99" box="[151,258,1738,1759]" italics="true" pageId="23" pageNumber="324">Hypoaspis</emphasis>
(
<emphasis id="B911EAB8FF86423B3C0F14D59DEC6F99" box="[274,394,1737,1759]" italics="true" pageId="23" pageNumber="324">Holostaspis</emphasis>
)
<emphasis id="B911EAB8FF86423B3C8614D59D8E6F99" box="[411,488,1737,1759]" italics="true" pageId="23" pageNumber="324">oophila</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC410BBFF86423B3F1B14D69F606F99" author="Evans, G. O. &amp; Till, W. M." box="[518,774,1737,1759]" page="159" pageId="23" pageNumber="324" year="1966">
<bibRefCitation id="EFF44B5BFF86423B3F1B14D69EB46F99" author="Evans, G. O. &amp; Till, W. M." box="[518,722,1737,1759]" pageId="23" pageNumber="324" pagination="107 - 370" refId="ref22029" refString="Evans, G. O. &amp; Till, W. M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History), Zoology, 14, 107 - 370." type="journal article" year="1966">Evans &amp; Till, 1966</bibRefCitation>
: 159
</treatmentCitation>
, 206;
<treatmentCitation id="0AC410BBFF86423B3E5514D69F956F99" author="Karg, W." box="[840,1011,1737,1759]" page="176" pageId="23" pageNumber="324" year="1971">
<bibRefCitation id="EFF44B5BFF86423B3E5514D69FD96F99" author="Karg, W." box="[840,959,1737,1759]" pageId="23" pageNumber="324" pagination="1 - 475" refId="ref23625" refString="Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1 - 475." type="journal article" year="1971">Karg, 1971</bibRefCitation>
: 176
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B391D14D698F96F99" author="Karg, W." box="[1024,1183,1737,1759]" page="97" pageId="23" pageNumber="324" year="1979">
<bibRefCitation id="EFF44B5BFF86423B391D14D698116F99" author="Karg, W." box="[1024,1143,1737,1759]" pageId="23" pageNumber="324" pagination="65 - 104" refId="ref23707" refString="Karg, W. (1979) Die Gattung Hypoaspis Canestrini 1884 (Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere, 106, 65 - 104." type="journal article" year="1979">Karg, 1979</bibRefCitation>
: 97
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B39B614D699316F99" author="Karg, W." box="[1195,1367,1737,1759]" page="247" pageId="23" pageNumber="324" year="1982">
<bibRefCitation id="EFF44B5BFF86423B39B614D699446F99" author="Karg, W." box="[1195,1314,1737,1759]" pageId="23" pageNumber="324" pagination="233 - 256" refId="ref23742" refString="Karg, W. (1982) Zur Kenntnis der Raubmilbeng Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 58, 233 - 256." type="journal article" year="1982">Karg, 1982</bibRefCitation>
: 247
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B387E14D69D566FB8" author="Karg, W." page="157" pageId="23" pageNumber="324" year="1993">
<bibRefCitation id="EFF44B5BFF86423B387E14D69C9B6FB8" author="Karg, W." pageId="23" pageNumber="324" pagination="1 - 523" refId="ref23832" refString="Karg, W. (1993) Acari (Acarina), Milben. Parasitiformes (Anactinochaeta). Cohors Gamasina Leach. Raubmilben. 2. Uberarbeitete Auflage. Die Tierwelt Deutschlands, 59, 1 - 523." type="journal article" year="1993">Karg, 1993</bibRefCitation>
: 157
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDA36AAFF86423B3D8A151A9F3C6E5A" blockId="23.[151,1436,1553,1820]" box="[151,858,1798,1820]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="AB751184FF86423B3D8A151A9F3C6E5A" box="[151,858,1798,1820]" pageId="23" pageNumber="324">
<taxonomicName id="4C654D29FF86423B3D8A151A9D076E5A" baseAuthorityName="Wasmann" baseAuthorityYear="1897" box="[151,353,1798,1820]" class="Arachnida" family="Laelapidae" genus="Holostaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophila">
<emphasis id="B911EAB8FF86423B3D8A151A9D076E5A" box="[151,353,1798,1820]" italics="true" pageId="23" pageNumber="324">Holostaspis oophila</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC410BBFF86423B3C9D151B9E396E5A" author="Bregetova, N. G." box="[384,607,1798,1820]" page="549" pageId="23" pageNumber="324" year="1977">
<bibRefCitation id="EFF44B5BFF86423B3C9D151B9E4A6E5A" author="Bregetova, N. G." box="[384,556,1798,1820]" pageId="23" pageNumber="324" pagination="486 - 556" refId="ref21303" refString="Bregetova, N. G. (1977) Family Laelaptidae Berlese, 1882. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil - Inhabiting Mites. Mesostigmata. Nauka, Leningrad, pp. 486 - 556. [in Russian]" type="book chapter" year="1977">Bregetova, 1977</bibRefCitation>
: 549
</treatmentCitation>
;
<treatmentCitation id="0AC410BBFF86423B3F77151B9F306E5A" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[618,854,1798,1820]" page="490" pageId="23" pageNumber="324" year="2017">
<bibRefCitation id="EFF44B5BFF86423B3F77151B9F456E5A" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[618,803,1798,1820]" pageId="23" pageNumber="324" pagination="485 - 505" refId="ref23930" refString="Keum, E., Jung, C. &amp; Joharchi, O. (2017) New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea. Zootaxa, 4353, 485 - 505. https: // doi. org / 10.11646 / zootaxa. 4353.3.5" type="journal article" year="2017">
Keum
<emphasis id="B911EAB8FF86423B3FB3151B9EBC6E5A" box="[686,730,1798,1820]" italics="true" pageId="23" pageNumber="324">et al</emphasis>
., 2017
</bibRefCitation>
: 490
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDA36AAFF86423B3D8A15559D5D6E88" blockId="23.[151,1436,1865,1998]" pageId="23" pageNumber="324">
<emphasis id="B911EAB8FF86423B3D8A15559DFE6E24" bold="true" box="[151,408,1865,1890]" pageId="23" pageNumber="324">Specimens examined.</emphasis>
Eight females—SW
<collectingCountry id="F372763AFF86423B3F9215559E926E24" box="[655,756,1865,1890]" name="Slovakia" pageId="23" pageNumber="324">Slovakia</collectingCountry>
, Podunajská Rovina Lowland, Svätý Jur Town, Šúr Nature Reserve (
<geoCoordinate id="EE51506DFF86423B3C1C15729DE96EC3" box="[257,399,1902,1926]" direction="north" orientation="latitude" pageId="23" pageNumber="324" precision="15" value="48.225834">48°13'33&quot; N</geoCoordinate>
,
<geoCoordinate id="EE51506DFF86423B3C8615729E406EC0" box="[411,550,1902,1926]" direction="east" orientation="longitude" pageId="23" pageNumber="324" precision="15" value="17.208055">17°12'29&quot; E</geoCoordinate>
), forest-steppe, in anthill of
<taxonomicName id="4C654D29FF86423B3E79157298576EC3" ID-CoL="6JGKH" authorityName="Retzius" authorityYear="1783" box="[868,1073,1902,1925]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="pratensis">
<emphasis id="B911EAB8FF86423B3E79157298576EC3" box="[868,1073,1902,1925]" italics="true" pageId="23" pageNumber="324">Formica pratensis</emphasis>
</taxonomicName>
, altitude
<quantity id="4C9D9B4FFF86423B3986157298836EC3" box="[1179,1253,1902,1926]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="324" unit="m" value="130.0">130 m</quantity>
,
<date id="FFDB106AFF86423B39ED157199F16EC0" box="[1264,1431,1901,1926]" pageId="23" pageNumber="324" value="1991-04-29">April 29, 1991</date>
, coll. P. Mašán. These specimens were previously published under the name
<taxonomicName id="4C654D29FF86423B391F158E99106EEF" box="[1026,1398,1937,1961]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="oophila" subGenus="Holostaspis">
<emphasis id="B911EAB8FF86423B391F158E981E6EEF" box="[1026,1144,1938,1961]" italics="true" pageId="23" pageNumber="324">Hypoaspis</emphasis>
(
<emphasis id="B911EAB8FF86423B3997158D99686EEF" box="[1162,1294,1937,1961]" italics="true" pageId="23" pageNumber="324">Holostaspis</emphasis>
)
<emphasis id="B911EAB8FF86423B383C158D99106EEF" box="[1313,1398,1937,1961]" italics="true" pageId="23" pageNumber="324">oophila</emphasis>
</taxonomicName>
by Mašán (2001).
</paragraph>
</subSubSection>
<subSubSection id="C37F6521FF8942343DDA128B9DCC68B3" pageId="24" pageNumber="325" type="diagnosis">
<paragraph id="8BDA36AAFF8942343DDA128B9DCC68B3" blockId="24.[151,1437,151,1941]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343DDA128B9DD669F6" bold="true" box="[199,432,151,176]" pageId="24" pageNumber="325">Diagnosis (female).</emphasis>
Dorsal shield suboval, more broadly rounded posteriorly and slightly more tapered anteriorly, smooth anteromedially, reticulate in posterior area, bearing 39 pairs of setae and one unpaired accessory seta between
<emphasis id="B911EAB8FF8942343C3012FC9D1669BE" box="[301,368,224,248]" italics="true" pageId="24" pageNumber="325">J3J4</emphasis>
; dorsal setae mostly short, setae r-series slightly longer than other podonotals, in posterior third of shield setae are also progressively longer, with
<emphasis id="B911EAB8FF8942343FC413189F7A685D" box="[729,796,260,284]" italics="true" pageId="24" pageNumber="325">J4J5</emphasis>
,
<emphasis id="B911EAB8FF8942343E3113189F10685D" box="[812,886,260,284]" italics="true" pageId="24" pageNumber="325">Z3Z5</emphasis>
and
<emphasis id="B911EAB8FF8942343EAD13189F90685D" box="[944,1014,260,284]" italics="true" pageId="24" pageNumber="325">S3S5</emphasis>
setae 35 times longer than anterior setae, and slightly pilose; vertical setae (
<emphasis id="B911EAB8FF8942343F9F13349EFE6806" box="[642,664,296,320]" italics="true" pageId="24" pageNumber="325">j1</emphasis>
) longer than most other, podonotal setae, but shorter than most opisthonotal setae; setae
<emphasis id="B911EAB8FF8942343CB013509DA06822" box="[429,454,332,356]" italics="true" pageId="24" pageNumber="325">z1</emphasis>
slightly shorter than most other podonotal setae. Sternal shield slightly wider than long, smooth. Anal shield subtriangular, elongate, with anterior margin concave and anterolateral angles pointed; postanal seta thickened and approximately 1.31.5 times longer than adanals. Peritremes anteriorly reaching level of
<emphasis id="B911EAB8FF8942343DAB13A49CA96896" box="[182,207,440,464]" italics="true" pageId="24" pageNumber="325">s1</emphasis>
. Epistome with anterior margin rounded. Fixed digit of chelicera with two teeth, movable digit with a small rounded subdistal bulge.
</paragraph>
</subSubSection>
<subSubSection id="C37F6521FF8942343DDA13E39E836ED3" pageId="24" pageNumber="325" type="description">
<paragraph id="8BDA36AAFF8942343DDA13E39D916B5E" blockId="24.[151,1437,151,1941]" box="[199,503,511,536]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343DDA13E39D3F6B5E" bold="true" box="[199,345,511,536]" pageId="24" pageNumber="325">Description.</emphasis>
Female (n=4)
</paragraph>
<paragraph id="8BDA36AAFF8942343DDA10389E0F6A7E" blockId="24.[151,1437,151,1941]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343DDA10389DE46B7A" box="[199,386,548,572]" italics="true" pageId="24" pageNumber="325">Dorsal idiosoma</emphasis>
(
<figureCitation id="135E2A2FFF8942343C8C10389D836B7B" box="[401,485,548,573]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 57</figureCitation>
). Idiosoma
<quantity id="4C9D9B4FFF8942343F7510389E906B7B" box="[616,758,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="6.01" metricValueMax="6.06" metricValueMin="5.96" pageId="24" pageNumber="325" unit="mm" value="601.0" valueMax="606.0" valueMin="596.0">596606 µm</quantity>
long and
<quantity id="4C9D9B4FFF8942343E7810389F926B7B" box="[869,1012,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="4.095" metricValueMax="4.16" metricValueMin="4.029999999999999" pageId="24" pageNumber="325" unit="mm" value="409.5" valueMax="416.0" valueMin="403.0">403416 µm</quantity>
wide; dorsal shield
<quantity id="4C9D9B4FFF89423439C8103899026B7B" box="[1237,1380,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="5.795" metricValueMax="5.87" metricValueMin="5.72" pageId="24" pageNumber="325" unit="mm" value="579.5" valueMax="587.0" valueMin="572.0">572587 µm</quantity>
long and
<quantity id="4C9D9B4FFF8942343DDB10549D336B26" box="[198,341,584,608]" metricMagnitude="-1" metricUnit="m" metricValue="3.08" metricValueMax="3.09" metricValueMin="3.07" pageId="24" pageNumber="325" unit="mm" value="308.0" valueMax="309.0" valueMin="307.0">307309 µm</quantity>
wide, reticulate on marginal and posterior surface (anteromedial region smooth); bearing 39 pairs of setae: 22 pairs (
<emphasis id="B911EAB8FF8942343C7610709DC36BC2" box="[363,421,620,644]" italics="true" pageId="24" pageNumber="325">j1j6</emphasis>
,
<emphasis id="B911EAB8FF8942343CA910709D926BC2" box="[436,500,620,644]" italics="true" pageId="24" pageNumber="325">z1z6</emphasis>
,
<emphasis id="B911EAB8FF8942343F1E10709E226BC2" box="[515,580,620,644]" italics="true" pageId="24" pageNumber="325">s1s6</emphasis>
,
<emphasis id="B911EAB8FF8942343F4E10709EF46BC5" box="[595,658,620,644]" italics="true" pageId="24" pageNumber="325">r2r5</emphasis>
) in podonotal area and 17 pairs (
<emphasis id="B911EAB8FF8942343912107098346BC5" box="[1039,1106,620,644]" italics="true" pageId="24" pageNumber="325">J1J5</emphasis>
,
<emphasis id="B911EAB8FF894234397C107098CD6BC5" box="[1121,1195,620,644]" italics="true" pageId="24" pageNumber="325">Z1Z5</emphasis>
,
<emphasis id="B911EAB8FF89423439A7107099796BC2" box="[1210,1311,620,644]" italics="true" pageId="24" pageNumber="325">Px2Px3</emphasis>
,
<emphasis id="B911EAB8FF8942343833107099126BC5" box="[1326,1396,620,644]" italics="true" pageId="24" pageNumber="325">S1S5</emphasis>
) in opisthonotal area and one unpaired accessory setae between
<emphasis id="B911EAB8FF8942343E57108C9FEA6BE1" box="[842,908,656,680]" italics="true" pageId="24" pageNumber="325">J1J5</emphasis>
; with at least 20 pairs of discernible pore-like structures (four pairs of gland pores and 16 pairs of lyrifissures). Dorsal shield setae apparently heterogeneous in length and thickness: thin and smooth setae on podonotum markedly shorter than thickened, elongate and apically pilose ones on opisthonotum; lengths of some setae:
<emphasis id="B911EAB8FF8942343FE410E09F686A52" box="[761,782,764,788]" italics="true" pageId="24" pageNumber="325">
j
<date id="FFDB106AFF8942343E1910E09F686A52" box="[772,782,764,788]" pageId="24" pageNumber="325" value="1927-01-30">1</date>
</emphasis>
<quantity id="4C9D9B4FFF8942343E0510E09FE96A53" box="[792,911,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="24" pageNumber="325" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">2730 µm</quantity>
,
<emphasis id="B911EAB8FF8942343E8010E09FB16A52" box="[925,983,764,788]" italics="true" pageId="24" pageNumber="325">j2j6</emphasis>
and
<emphasis id="B911EAB8FF894234390F10E098346A52" box="[1042,1106,764,788]" italics="true" pageId="24" pageNumber="325">
z1z
<date id="FFDB106AFF894234395910E098346A52" box="[1092,1106,764,788]" pageId="24" pageNumber="325" value="2017-06-20">6</date>
</emphasis>
<quantity id="4C9D9B4FFF894234394610E098B56A53" box="[1115,1235,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="24" pageNumber="325" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 µm</quantity>
,
<emphasis id="B911EAB8FF89423439FC10E099476A52" box="[1249,1313,764,788]" italics="true" pageId="24" pageNumber="325">s1s6</emphasis>
and
<emphasis id="B911EAB8FF894234384110E099FA6A55" box="[1372,1436,764,788]" italics="true" pageId="24" pageNumber="325">
r2r
<date id="FFDB106AFF894234389310E099FA6A55" box="[1422,1436,764,787]" pageId="24" pageNumber="325" value="1933-05-27">5</date>
</emphasis>
<quantity id="4C9D9B4FFF8942343D8A113C9D6A6A7E" box="[151,268,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.3" metricValueMin="2.7" pageId="24" pageNumber="325" unit="mm" value="30.0" valueMax="33.0" valueMin="27.0">2733 µm</quantity>
, longest posterior setae
<quantity id="4C9D9B4FFF8942343F04113C9E036A7E" box="[537,613,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="24" pageNumber="325" unit="mm" value="87.0">87 µm</quantity>
.
</paragraph>
<paragraph id="8BDA36AAFF8942343DDA115898976C86" blockId="24.[151,1437,151,1941]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343DDA11589DEE6A1A" box="[199,392,836,860]" italics="true" pageId="24" pageNumber="325">Ventral idiosoma</emphasis>
(
<figureCitation id="135E2A2FFF8942343C8411589D896A1B" box="[409,495,836,861]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 58</figureCitation>
). Tritosternum with short columniform base (
<quantity id="4C9D9B4FFF8942343EE61158982E6A1A" box="[1019,1096,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="325" unit="mm" value="17.0">17 µm</quantity>
) and pilose laciniae (
<quantity id="4C9D9B4FFF8942343822115899F66A1A" box="[1343,1424,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="24" pageNumber="325" unit="mm" value="54.0">54 µm</quantity>
). Sternal shield subrectangular,
<quantity id="4C9D9B4FFF8942343CF1117B9E086AC6" box="[492,622,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" metricValueMax="1.06" metricValueMin="0.99" pageId="24" pageNumber="325" unit="mm" value="102.5" valueMax="106.0" valueMin="99.0">99106 µm</quantity>
long and
<quantity id="4C9D9B4FFF8942343FFD11749F086AC6" box="[736,878,872,896]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.1099999999999999" metricValueMin="1.04" pageId="24" pageNumber="325" unit="mm" value="107.5" valueMax="111.0" valueMin="104.0">104111 µm</quantity>
wide (L/W=0.95), widest at level of coxae IIIII (
<quantity id="4C9D9B4FFF8942343D8211909D5C6AE2" box="[159,314,908,932]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="1.77" metricValueMin="1.73" pageId="24" pageNumber="325" unit="mm" value="175.0" valueMax="177.0" valueMin="173.0">173177 µm</quantity>
), convex anteriorly (except shallow medial concavity) and concave posteriorly (occasionally posterior margin unsclerotised and not indicated, as in
<figureCitation id="135E2A2FFF8942343E2511B39FF56A8E" box="[824,915,943,968]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 59</figureCitation>
); whole surface smooth, without sculptural ornamentation; setae with tips not reaching bases of following ones (
<emphasis id="B911EAB8FF8942343EB811C89FA06AAA" box="[933,966,980,1004]" italics="true" pageId="24" pageNumber="325">st1</emphasis>
<quantity id="4C9D9B4FFF8942343ED011C898226AAB" box="[973,1092,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="24" pageNumber="325" unit="mm" value="43.5" valueMax="45.0" valueMin="42.0">4245 µm</quantity>
,
<emphasis id="B911EAB8FF894234394C11C898176AAA" box="[1105,1137,980,1004]" italics="true" pageId="24" pageNumber="325">st2</emphasis>
<quantity id="4C9D9B4FFF894234396411C898896AAB" box="[1145,1263,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="24" pageNumber="325" unit="mm" value="38.5" valueMax="40.0" valueMin="37.0">3740 µm</quantity>
, and
<emphasis id="B911EAB8FF894234383011C899286AAA" box="[1325,1358,980,1004]" italics="true" pageId="24" pageNumber="325">st3</emphasis>
<quantity id="4C9D9B4FFF894234384B11C89CA76D56" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="24" pageNumber="325" unit="mm" value="39.5" valueMax="42.0" valueMin="37.0">3742 µm</quantity>
); lyrifissures
<date id="FFDB106AFF8942343C4111E49DBD6D56" box="[348,475,1015,1040]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343C4111E49D186D56" box="[348,382,1016,1040]" italics="true" pageId="24" pageNumber="325">iv1</emphasis>
and
<emphasis id="B911EAB8FF8942343CA511E49DBD6D56" box="[440,475,1016,1040]" italics="true" pageId="24" pageNumber="325">iv2</emphasis>
</date>
slit-like. Metasternal setae (
<emphasis id="B911EAB8FF8942343E3D11E49F276D56" box="[800,833,1016,1040]" italics="true" pageId="24" pageNumber="325">st4</emphasis>
<quantity id="4C9D9B4FFF8942343E5411E49FD96D56" box="[841,959,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="24" pageNumber="325" unit="mm" value="34.5" valueMax="37.0" valueMin="32.0">3237 µm</quantity>
) and associated lyrifissures
<emphasis id="B911EAB8FF894234381E11E499436D56" box="[1283,1317,1016,1040]" italics="true" pageId="24" pageNumber="325">
<date id="FFDB106AFF894234381E11E499436D56" box="[1283,1317,1016,1040]" pageId="24" pageNumber="325">iv3</date>
</emphasis>
placed on soft integument; endopodal platelets IIIIV subtriangular, narrow, anteriorly free or connected to posterolateral margins of sternal shield. Epigynal shield tongue-shaped, elongate,
<quantity id="4C9D9B4FFF8942343EA91623982E6D1E" box="[948,1096,1087,1112]" metricMagnitude="-1" metricUnit="m" metricValue="2.785" metricValueMax="2.85" metricValueMin="2.7199999999999998" pageId="24" pageNumber="325" unit="mm" value="278.5" valueMax="285.0" valueMin="272.0">272285 µm</quantity>
long and
<quantity id="4C9D9B4FFF89423439DB1623993C6D1E" box="[1222,1370,1087,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.595" metricValueMax="1.63" metricValueMin="1.56" pageId="24" pageNumber="325" unit="mm" value="159.5" valueMax="163.0" valueMin="156.0">156163 µm</quantity>
wide (
<quantity id="4C9D9B4FFF8942343D8216789D526D3B" box="[159,308,1124,1149]" metricMagnitude="-1" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.26" metricValueMin="1.21" pageId="24" pageNumber="325" unit="mm" value="123.5" valueMax="126.0" valueMin="121.0">121126 µm</quantity>
at level of
<emphasis id="B911EAB8FF8942343CD916789D826D3D" box="[452,484,1124,1147]" italics="true" pageId="24" pageNumber="325">st5</emphasis>
); anterior hyaline margin irregularly curved, reaching level between
<emphasis id="B911EAB8FF894234380E167899526D3A" box="[1299,1332,1124,1148]" italics="true" pageId="24" pageNumber="325">st1</emphasis>
and
<emphasis id="B911EAB8FF8942343869167899F26D3A" box="[1396,1428,1124,1148]" italics="true" pageId="24" pageNumber="325">st2</emphasis>
; posterior portion with rounded margin and lineate ornamentation pattern forming usually 89 elongate cells; genital setae (
<emphasis id="B911EAB8FF8942343C2B16B09D316D85" box="[310,343,1196,1219]" italics="true" pageId="24" pageNumber="325">st5</emphasis>
<quantity id="4C9D9B4FFF8942343C7D16B09DCA6D82" box="[352,428,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="325" unit="mm" value="40.0">40 µm</quantity>
) placed on mediolateral margins and associated lyrifissures
<emphasis id="B911EAB8FF894234397D16B098E46D85" box="[1120,1154,1196,1219]" italics="true" pageId="24" pageNumber="325">
<date id="FFDB106AFF894234397D16B098E46D85" box="[1120,1154,1196,1219]" pageId="24" pageNumber="325">iv5</date>
</emphasis>
on soft integument; soft integument between
<emphasis id="B911EAB8FF8942343C6216CC9DC76DA1" box="[383,417,1232,1255]" italics="true" pageId="24" pageNumber="325">
<date id="FFDB106AFF8942343C6216CC9DC76DA1" box="[383,417,1232,1255]" pageId="24" pageNumber="325">iv5</date>
</emphasis>
and
<emphasis id="B911EAB8FF8942343CCB16CC9E666DAE" box="[470,512,1232,1256]" italics="true" pageId="24" pageNumber="325">Zv1</emphasis>
each with a pair of minute and narrow adgenital sclerites. Metapodal platelets long and narrow,
<quantity id="4C9D9B4FFF8942343C3716E89DC76C4B" box="[298,417,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="24" pageNumber="325" unit="mm" value="51.0" valueMax="52.0" valueMin="50.0">5052 µm</quantity>
long. Anal shield elongate, triangle-shaped,
<quantity id="4C9D9B4FFF8942343EB316E898436C4B" box="[942,1061,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="8.9" metricValueMin="8.6" pageId="24" pageNumber="325" unit="mm" value="87.5" valueMax="89.0" valueMin="86.0">8689 µm</quantity>
long and
<quantity id="4C9D9B4FFF89423439BD16E899716C4B" box="[1184,1303,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" metricValueMax="7.1" metricValueMin="6.2" pageId="24" pageNumber="325" unit="mm" value="66.5" valueMax="71.0" valueMin="62.0">6271 µm</quantity>
wide, with anterolateral angles obtusely pointed, anterior margin deeply concave and weak lineate-reticulate ornamentation on anterior portion; postanal seta
<quantity id="4C9D9B4FFF8942343CEF17209E006C13" box="[498,614,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="24" pageNumber="325" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">4045 µm</quantity>
long, thickened and longer than adanal setae (
<quantity id="4C9D9B4FFF8942343967172098956C12" box="[1146,1267,1340,1364]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="24" pageNumber="325" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">3032 µm</quantity>
); cribrum with wide band of spicules. Lateral and opisthogastric soft integument with 17 pairs of setae; setae different in length, each with base inserted in small and rounded scutal structure; setae
<emphasis id="B911EAB8FF8942343E9417989FC76CDA" box="[905,929,1412,1436]" italics="true" pageId="24" pageNumber="325">r6</emphasis>
,
<emphasis id="B911EAB8FF8942343EB217989FA86CDA" box="[943,974,1412,1436]" italics="true" pageId="24" pageNumber="325">R1</emphasis>
, and
<emphasis id="B911EAB8FF89423439161798984C6CDA" box="[1035,1066,1412,1436]" italics="true" pageId="24" pageNumber="325">R2</emphasis>
shorter and smooth, other longer and sparsely pilose distally. Poststigmatic section of peritrematal shield narrow, tapered posteriorly.
</paragraph>
<paragraph id="8BDA36AAFF8942343DDA17D09F816FFB" blockId="24.[151,1437,151,1941]" pageId="24" pageNumber="325">
<taxonomicName id="4C654D29FF8942343DDA17D09D306CA2" authorityName="Krefft" authorityYear="1871" box="[199,342,1484,1508]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="24" pageNumber="325" phylum="Nematoda" rank="genus">
<emphasis id="B911EAB8FF8942343DDA17D09D306CA2" box="[199,342,1484,1508]" italics="true" pageId="24" pageNumber="325">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="135E2A2FFF8942343C7817D09D8E6CA3" box="[357,488,1484,1509]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Figs 6062</figureCitation>
). Epistome subtriangular, with narrower base and rounded anterior margin (
<figureCitation id="135E2A2FFF894234382617D099F66CA3" box="[1339,1424,1484,1509]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 60</figureCitation>
). Corniculi weakly sclerotised, short and pointed, horn-like; internal malae short, divergent, finely serrate laterobasally, and with tips not reaching beyond corniculi; labrum long and blade-like (
<figureCitation id="135E2A2FFF8942343981140898926F6B" box="[1180,1268,1556,1581]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 61</figureCitation>
). Deutosternal furrow on ventral subcapitulum with almost parallel lateral margins, with six transverse rows of denticles (each with 812 denticles). Lengths of hypostomal and palpcoxal setae:
<emphasis id="B911EAB8FF8942343E6514409FF26F32" box="[888,916,1628,1652]" italics="true" pageId="24" pageNumber="325">
h
<date id="FFDB106AFF8942343E9514409FF26F32" box="[904,916,1628,1652]" pageId="24" pageNumber="325" value="1922-01-25">1</date>
</emphasis>
<quantity id="4C9D9B4FFF8942343E87144098696F33" box="[922,1039,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="24" pageNumber="325" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">2225 µm</quantity>
,
<emphasis id="B911EAB8FF8942343907144098516F32" box="[1050,1079,1628,1652]" italics="true" pageId="24" pageNumber="325">h2</emphasis>
<quantity id="4C9D9B4FFF8942343920144098E16F33" box="[1085,1159,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="mm" value="20.0">20 µm</quantity>
,
<emphasis id="B911EAB8FF894234398E144098C96F32" box="[1171,1199,1628,1652]" italics="true" pageId="24" pageNumber="325">h3</emphasis>
<quantity id="4C9D9B4FFF89423439A81440994C6F33" box="[1205,1322,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="24" pageNumber="325" unit="mm" value="33.5" valueMax="35.0" valueMin="32.0">3235 µm</quantity>
,
<emphasis id="B911EAB8FF8942343828144199296F32" box="[1333,1359,1629,1652]" italics="true" pageId="24" pageNumber="325">pc</emphasis>
<quantity id="4C9D9B4FFF894234384B14409CA66FDF" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="24" pageNumber="325" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">2730 µm</quantity>
. Palp setation as in
<figureCitation id="135E2A2FFF8942343CB514639D9B6FDE" box="[424,509,1663,1688]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 61</figureCitation>
. Shape and dentition of chelicera as in
<figureCitation id="135E2A2FFF8942343ED81463987F6FDE" box="[965,1049,1663,1688]" captionStart="FIGURES 5762" captionStartId="25.[151,264,1940,1963]" captionTargetBox="[190,1408,203,1923]" captionTargetId="figure@25.[182,1412,193,1931]" captionTargetPageId="25" captionText="FIGURES 5762. Holostaspis oophila, female. 57. Dorsal idiosoma; 58. Ventral idiosoma; 59. Sternal and epigynal shield (a specimen with unsclerotised posterior margin of sternal shield); 60. Epistome; 61. Ventral gnathosoma with palp; 62. Chelicera." httpUri="https://zenodo.org/record/2653390/files/figure.png" pageId="24" pageNumber="325">Fig. 62</figureCitation>
; lengths of cheliceral articles: the second segment
<quantity id="4C9D9B4FFF8942343C4D14B89DA36FFB" box="[336,453,1700,1725]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="6.7" metricValueMin="6.3" pageId="24" pageNumber="325" unit="mm" value="65.0" valueMax="67.0" valueMin="63.0">6367 µm</quantity>
, fixed digit
<quantity id="4C9D9B4FFF8942343F5114B89EF36FFB" box="[588,661,1700,1725]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="325" unit="mm" value="15.0">15 µm</quantity>
and movable digit
<quantity id="4C9D9B4FFF8942343E7014B89F856FFB" box="[877,995,1700,1725]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="24" pageNumber="325" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 µm</quantity>
.
</paragraph>
<paragraph id="8BDA36AAFF8942343DDA14D5985E6E6E" blockId="24.[151,1437,151,1941]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343DDA14D59C9A6FA6" box="[199,252,1737,1760]" italics="true" pageId="24" pageNumber="325">Legs</emphasis>
(
<figureCitation id="135E2A2FFF8942343C1714DB9DEB6FA6" box="[266,397,1735,1760]" captionStart="FIGURES 6366" captionStartId="26.[151,264,1589,1612]" captionTargetBox="[187,1390,254,1473]" captionTargetId="figure@26.[151,1436,193,1586]" captionTargetPageId="26" captionText="FIGURES 6366. Holostaspis oophila, female. 63. Leg I; 64. Leg II; 65. Leg III; 66. Leg IV." httpUri="https://zenodo.org/record/2653392/files/figure.png" pageId="24" pageNumber="325">Figs 6366</figureCitation>
). Tarsi IIV each with a pair of claws and pulvillus. Lengths of legs: leg I
<quantity id="4C9D9B4FFF89423439D814DB99336FA7" box="[1221,1365,1735,1761]" metricMagnitude="-1" metricUnit="m" metricValue="3.675" metricValueMax="3.7399999999999998" metricValueMin="3.61" pageId="24" pageNumber="325" unit="mm" value="367.5" valueMax="374.0" valueMin="361.0">361374 µm</quantity>
, leg II
<quantity id="4C9D9B4FFF8942343D8A14F09D4D6E43" box="[151,299,1772,1797]" metricMagnitude="-1" metricUnit="m" metricValue="2.66" metricValueMax="2.7199999999999998" metricValueMin="2.6" pageId="24" pageNumber="325" unit="mm" value="266.0" valueMax="272.0" valueMin="260.0">260272 µm</quantity>
, leg III
<quantity id="4C9D9B4FFF8942343C9514F09E7D6E43" box="[392,539,1772,1797]" metricMagnitude="-1" metricUnit="m" metricValue="2.785" metricValueMax="2.82" metricValueMin="2.75" pageId="24" pageNumber="325" unit="mm" value="278.5" valueMax="282.0" valueMin="275.0">275282 µm</quantity>
, leg IV
<quantity id="4C9D9B4FFF8942343F6714F09F686E43" box="[634,782,1772,1797]" metricMagnitude="-1" metricUnit="m" metricValue="3.3649999999999998" metricValueMax="3.3899999999999997" metricValueMin="3.34" pageId="24" pageNumber="325" unit="mm" value="336.5" valueMax="339.0" valueMin="334.0">334339 µm</quantity>
. Setae
<emphasis id="B911EAB8FF8942343E7D14F19F1C6E42" box="[864,890,1773,1796]" italics="true" pageId="24" pageNumber="325">av</emphasis>
and
<emphasis id="B911EAB8FF8942343EA814F19FB66E42" box="[949,976,1773,1796]" italics="true" pageId="24" pageNumber="325">pv</emphasis>
on genu II, tibia II and tibia IV slightly thickened; seta
<emphasis id="B911EAB8FF8942343C58150C9D076E6E" box="[325,353,1808,1832]" italics="true" pageId="24" pageNumber="325">ad</emphasis>
on trochanter IV,
<emphasis id="B911EAB8FF8942343F36150C9E336E6E" box="[555,597,1808,1832]" italics="true" pageId="24" pageNumber="325">ad1</emphasis>
on femur IV and
<emphasis id="B911EAB8FF8942343E00150D9F5E6E6E" box="[797,824,1809,1832]" italics="true" pageId="24" pageNumber="325">av</emphasis>
on genu IV thickened.
</paragraph>
<paragraph id="8BDA36AAFF8942343DDA15299F006E0B" blockId="24.[151,1437,151,1941]" box="[199,870,1844,1869]" pageId="24" pageNumber="325">MALE. Briefly described and illustrated by Berlese (1904b).</paragraph>
<paragraph id="8BDA36AAFF8942343DDA154B9E836ED3" blockId="24.[151,1437,151,1941]" pageId="24" pageNumber="325">
<emphasis id="B911EAB8FF8942343DDA154B9D776E36" bold="true" box="[199,273,1879,1904]" pageId="24" pageNumber="325">Notes.</emphasis>
<bibRefCitation id="EFF44B5BFF8942343C0A154B9DAF6E36" author="Vitzthum, H." box="[279,457,1879,1904]" pageId="24" pageNumber="325" pagination="1 - 86" refId="ref25895" refString="Vitzthum, H. (1925) Acarologische Beobachtungen. 8. Reihe. Archiv fur Naturgeschichte, Abteilung A, 10, 1 - 86." type="journal article" year="1925">Vitzthum (1925</bibRefCitation>
,
<bibRefCitation id="EFF44B5BFF8942343CC815449E686E36" author="Vitzthum, H." box="[469,526,1880,1904]" pageId="24" pageNumber="325" pagination="1 - 112" refId="ref25922" refString="Vitzthum, H. (1929) Ordnung: Milben, Acari. Die Tierwelt Mitteleuropas. Band 3, Lief. 3, Abteilung 7, 1 - 112." type="journal article" year="1929">1929</bibRefCitation>
) considered that
<taxonomicName id="4C654D29FF8942343FD015459F226E36" box="[717,836,1880,1904]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="oophila">
<emphasis id="B911EAB8FF8942343FD015459E876E36" box="[717,737,1881,1904]" italics="true" pageId="24" pageNumber="325">H</emphasis>
.
<emphasis id="B911EAB8FF8942343FF315449F226E36" box="[750,836,1880,1904]" italics="true" pageId="24" pageNumber="325">oophila</emphasis>
</taxonomicName>
was a synonym of
<taxonomicName id="4C654D29FF8942343906154598C66E36" authorityName="Kolenati" authorityYear="1858" box="[1051,1184,1880,1904]" class="Arachnida" family="Laelapidae" genus="Holostaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="isotricha">
<emphasis id="B911EAB8FF8942343906154598496E36" box="[1051,1071,1881,1904]" italics="true" pageId="24" pageNumber="325">H</emphasis>
.
<emphasis id="B911EAB8FF8942343921154498C66E36" box="[1084,1184,1880,1904]" italics="true" pageId="24" pageNumber="325">isotricha</emphasis>
</taxonomicName>
, but
<bibRefCitation id="EFF44B5BFF89423439CA154499FA6E36" author="Bregetova, N. G." box="[1239,1436,1880,1904]" pageId="24" pageNumber="325" pagination="486 - 556" refId="ref21303" refString="Bregetova, N. G. (1977) Family Laelaptidae Berlese, 1882. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil - Inhabiting Mites. Mesostigmata. Nauka, Leningrad, pp. 486 - 556. [in Russian]" type="book chapter" year="1977">Bregetova (1977)</bibRefCitation>
and subsequent authors treated both as valid species.
</paragraph>
</subSubSection>
</treatment>
</document>