319 lines
38 KiB
XML
319 lines
38 KiB
XML
<document id="D2FB79E436D8C2EBE5E11F4F887867A8" ID-DOI="10.1051/acarologia/20101988" ID-ISSN="2107-7207" ID-Zenodo-Dep="5393450" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630318902626" checkinUser="marcus" docAuthor="Nakamura, Y. - N., Fukumori, S. & Fujikawa, T." docDate="2010" docId="03CA8799FFE6FFE7BEE2BD8FFEA3B87A" docLanguage="en" docName="Acarologia.2010.50.4.439-477.pdf" docOrigin="Acarologia 50 (4)" docSource="http://dx.doi.org/10.1051/acarologia/20101988" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Tectocepheus kumayaensis Nakamura & Fukumori & Fujikawa 2010, sp. nov." docType="treatment" docVersion="3" lastPageNumber="450" masterDocId="FFF3FFE1FFEFFFEABD7AB94EFFB7BD5E" masterDocTitle="Oribatid Fauna (Acari, Oribatida) From The Kumaya Cave Of Iheya Village In Central Ryukyu Arc, South Japan, With A Description Of Several New Species" masterLastPageNumber="477" masterPageNumber="439" pageNumber="447" updateTime="1699225274479" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:titleInfo id="514B4360A706A36E41FC67D5E4B628ED">
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<mods:title id="9387A5E6AE2B0A2E42078A0D46EB0639">Oribatid Fauna (Acari, Oribatida) From The Kumaya Cave Of Iheya Village In Central Ryukyu Arc, South Japan, With A Description Of Several New Species</mods:title>
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<mods:name id="013C3139E34A0791CA750524959C8641" type="personal">
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<mods:roleTerm id="C36A5BEDC5F5C1436C64AD7ADEBF9F8E">Author</mods:roleTerm>
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<mods:namePart id="53973EC45575EF16ABE0DEF5165E9B8C">Nakamura, Y. - N.</mods:namePart>
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<mods:roleTerm id="FAF48E0CF75C4E41FD4613BD3537FC42">Author</mods:roleTerm>
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<mods:namePart id="6449F0A51F5AD95B959F951A01497C2D">Fukumori, S.</mods:namePart>
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<mods:name id="AAD60AEA640BDF0188CA83CB8B2A4D96" type="personal">
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<mods:roleTerm id="D53991797CAE5C97653E8E8FB92A7A27">Author</mods:roleTerm>
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<mods:namePart id="B0B2D568F0A642F6462AEFA5074B7D20">Fujikawa, T.</mods:namePart>
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<mods:typeOfResource id="0FA2DE8413C95194BF1E0C2E21F96424">text</mods:typeOfResource>
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<mods:title id="A20984AD61645EDD74FF8FA9C29B902B">Acarologia</mods:title>
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</mods:titleInfo>
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<mods:part id="F01BF0F2BECDF210615B35F7C220054B">
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<mods:date id="9BA5F389CF75BDDD4D8C40C7B76A8243">2010</mods:date>
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<mods:number id="B2314133E83036F242D9DD1BB67F9669">2010-12-22</mods:number>
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<mods:detail id="EDAC1475EDCAA9D23FBED54385F7C755" type="volume">
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<mods:number id="F8646701ABB689F7E4B221E391972EF2">50</mods:number>
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<mods:detail id="1007EE31AB291334102E7ADB03BDC692" type="issue">
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<mods:number id="6732DB30168706E92B4A74CEBEBA3FCF">4</mods:number>
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<mods:start id="725F00D319553315416468B7F286D73E">439</mods:start>
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<mods:url id="9CC73B0D7A21113D78B269B8195655DD">http://dx.doi.org/10.1051/acarologia/20101988</mods:url>
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<mods:classification id="3A2418C1EB6B468F2840D98746DCF7F7">journal article</mods:classification>
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<mods:identifier id="8ADC2A3AA9BDF982D20AB4F1BAF9F131" type="DOI">10.1051/acarologia/20101988</mods:identifier>
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<mods:identifier id="D8C5ECDA171F42F27959B56BB963DE64" type="ISSN">2107-7207</mods:identifier>
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<treatment id="03CA8799FFE6FFE7BEE2BD8FFEA3B87A" LSID="urn:lsid:plazi:treatment:03CA8799FFE6FFE7BEE2BD8FFEA3B87A" httpUri="http://treatment.plazi.org/id/03CA8799FFE6FFE7BEE2BD8FFEA3B87A" lastPageId="13" lastPageNumber="450" pageId="9" pageNumber="447">
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<subSubSection id="C3796504FFE6FFE3BEE2BD8FFB0DB868" pageId="9" pageNumber="447" type="nomenclature">
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<paragraph id="8BDC368FFFE6FFE3BEE2BD8FFA81B987" blockId="9.[920,1334,1217,1241]" box="[920,1334,1217,1241]" pageId="9" pageNumber="447">
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<heading id="D09481E3FFE6FFE3BEE2BD8FFA81B987" bold="true" box="[920,1334,1217,1241]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="447" reason="2">
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<emphasis id="B917EA9DFFE6FFE3BEE2BD8FFA81B987" bold="true" box="[920,1334,1217,1241]" pageId="9" pageNumber="447">
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<taxonomicName id="4C634D0CFFE6FFE3BEE2BD8FFB64B987" ID-CoL="8TB3Y" authority="Nakamura & Fukumori & Fujikawa, 2010" authorityName="Nakamura & Fukumori & Fujikawa" authorityYear="2010" box="[920,1235,1217,1241]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="447" phylum="Arthropoda" rank="species" species="kumayaensis" status="sp. nov.">
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<emphasis id="B917EA9DFFE6FFE3BEE2BD8FFB64B987" bold="true" box="[920,1235,1217,1241]" italics="true" pageId="9" pageNumber="447">Tectocepheus kumayaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22457E6FFE6FFE3B9A0BD8FFA81B987" box="[1242,1334,1217,1241]" pageId="9" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDC368FFFE6FFE3BE0EBDB5FB0DB868" blockId="9.[884,1371,1274,1334]" pageId="9" pageNumber="447">
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[Japanese name: Kumaya-kuwagatadani] (
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<figureCitation id="13582A0AFFE6FFE3B967BC50FBDDB86B" box="[1053,1130,1309,1334]" captionStart="FIGURE" captionStartId="11.[327,339,1845,1864]" captionTargetBox="[283,1257,403,1778]" captionTargetId="figure-4@11.[272,1316,377,1813]" captionTargetPageId="11" captionText="FIGURE 6: Tectocepheus kumayaensis sp. nov. A, Dorsal view; B, Camerostome; C, Genito-anal region." figureDoi="http://doi.org/10.5281/zenodo.5393462" httpUri="https://zenodo.org/record/5393462/files/figure.png" pageId="9" pageNumber="447">Figs. 6</figureCitation>
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and
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<figureCitation id="13582A0AFFE6FFE3B9DEBC50FB05B868" box="[1188,1202,1310,1334]" captionStart="FIGURE" captionStartId="12.[159,171,1067,1086]" captionTargetBox="[169,738,485,1034]" captionTargetId="figure-32@12.[165,755,477,1035]" captionTargetPageId="12" captionText="FIGURE 7: Tectocepheus kumayaensis sp. nov. A, Solenidial region on tarsus and tibia of leg I; B, Left epimeral region." figureDoi="http://doi.org/10.5281/zenodo.5393464" httpUri="https://zenodo.org/record/5393464/files/figure.png" pageId="9" pageNumber="447">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796504FFE6FFE7BE1ABC2DFEA3B87A" lastPageId="13" lastPageNumber="451" pageId="9" pageNumber="447" type="diagnosis">
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<paragraph id="8BDC368FFFE6FFE3BE1ABC2DFAD4BBA3" blockId="9.[826,1429,1379,1949]" pageId="9" pageNumber="447">
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Diagnosis — Body length 264 µm; width 178 µm. Rostral anterior margin without incision, broadly truncate with two concavities. Setae
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<emphasis id="B917EA9DFFE6FFE3B86FBCE4FA9CB89F" box="[1301,1323,1450,1473]" italics="true" pageId="9" pageNumber="447">in</emphasis>
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trigonal pyramidal, roughened. Sensilli with globular head. Bothridia with deep incision, without swelling or projection. Depression and dorsosejugal scissure absent. Humeral region with small triangular projection. Ten pairs of notogastral setae. Genito-anal setal formula: 6-1-2-3. Lyrifissures
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<emphasis id="B917EA9DFFE6FFE3B9B7BF32FB5ABBCA" box="[1229,1261,1660,1684]" italics="true" pageId="9" pageNumber="447">iad</emphasis>
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located along, near anterior margin of anal aperture. Epimeral setal formula: 3-1-3-3. Trochantera III and IV bearing carina with sharply pointed apex. Monodactyl.
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</paragraph>
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<paragraph id="8BDC368FFFE6FFE3BE1ABE5CFB18BA13" blockId="9.[826,1429,1379,1949]" pageId="9" pageNumber="447">
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Material examined —
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<typeStatus id="54D8882DFFE6FFE3B9DEBE5CFAA4BA74" box="[1188,1299,1810,1834]" pageId="9" pageNumber="447" type="holotype">Holotype</typeStatus>
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(Female) (NSMT-Ac 13590) from point C.
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</paragraph>
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<paragraph id="8BDC368FFFE6FFE3BE1ABE2FFC5FBAC3" blockId="9.[826,1429,1379,1949]" pageId="9" pageNumber="447">Etymology — After the name of sampling point, Kumaya Cave.</paragraph>
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<paragraph id="8BDC368FFFE5FFE0BCCBBFA9FB34BBA4" blockId="10.[433,1155,1767,1786]" box="[433,1155,1767,1786]" pageId="10" pageNumber="448">
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PLATE 2:
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<taxonomicName id="4C634D0CFFE5FFE0BF76BFA9FC26BBA4" authority="(Oudemans, 1902)" authorityName="Solenidial" baseAuthorityName="Oudemans" baseAuthorityYear="1902" box="[524,913,1767,1786]" class="Arachnida" family="Oppiidae" genus="Oppiella" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="448" phylum="Arthropoda" rank="species" species="nova" subGenus="Oppiella">
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<emphasis id="B917EA9DFFE5FFE0BF76BFA9FD57BBA7" box="[524,736,1767,1786]" italics="true" pageId="10" pageNumber="448">Oppiella (Oppiella) nova</emphasis>
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(
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<bibRefCitation id="EFF24B7EFFE5FFE0BF97BFA9FC3CBBA4" author="Oudemans A. C." box="[749,907,1767,1786]" pageId="10" pageNumber="448" pagination="50 - 64" refId="ref15545" refString="Oudemans A. C. 1902 - New list of dutch Acari - Tidjschr. Ent., 45, Verslagen: 50 - 64." type="journal article" year="1902">Oudemans, 1902</bibRefCitation>
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)
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</taxonomicName>
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(photos by Nakamura Y.)
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</paragraph>
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<caption id="DF1C6607FFE4FFE1BC3DBE7BFB58BA16" ID-DOI="http://doi.org/10.5281/zenodo.5393462" ID-Zenodo-Dep="5393462" box="[327,1263,1845,1864]" httpUri="https://zenodo.org/record/5393462/files/figure.png" pageId="11" pageNumber="449" startId="11.[327,339,1845,1864]" targetBox="[283,1257,403,1778]" targetPageId="11">
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<paragraph id="8BDC368FFFE4FFE1BC3DBE7BFB58BA16" blockId="11.[327,1263,1845,1864]" box="[327,1263,1845,1864]" pageId="11" pageNumber="449">
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FIGURE 6:
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<taxonomicName id="4C634D0CFFE4FFE1BCD6BE7BFD39BA16" authority="Nakamura & Fukumori & Fujikawa, 2010" authorityName="Nakamura & Fukumori & Fujikawa" authorityYear="2010" box="[428,654,1845,1864]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="449" phylum="Arthropoda" rank="species" species="kumayaensis" status="sp. nov.">
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<emphasis id="B917EA9DFFE4FFE1BCD6BE7BFD39BA16" box="[428,654,1845,1864]" italics="true" pageId="11" pageNumber="449">Tectocepheus kumayaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22457E6FFE4FFE1BFE9BE7BFD6ABA16" box="[659,733,1845,1864]" pageId="11" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
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A, Dorsal view; B, Camerostome; C, Genito-anal region.
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</paragraph>
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</caption>
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<paragraph id="8BDC368FFFE3FFE6BDBFB879FDF4BCE6" blockId="12.[159,762,311,440]" pageId="12" pageNumber="450">Measurements and body appearance — Body length 264 µm; width 178 µm. Body colour light brown. Whole body surface covered cerotegument; cerotegument irregularly granulate.</paragraph>
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<caption id="DF1C6607FFE3FFE6BDE5BD65FD2AB907" ID-DOI="http://doi.org/10.5281/zenodo.5393464" ID-Zenodo-Dep="5393464" httpUri="https://zenodo.org/record/5393464/files/figure.png" pageId="12" pageNumber="450" startId="12.[159,171,1067,1086]" targetBox="[169,738,485,1034]" targetPageId="12">
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<paragraph id="8BDC368FFFE3FFE6BDE5BD65FD2AB907" blockId="12.[159,761,1066,1113]" pageId="12" pageNumber="450">
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FIGURE 7:
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<taxonomicName id="4C634D0CFFE3FFE6BC7CBD64FE5FB963" authority="Nakamura & Fukumori & Fujikawa, 2010" authorityName="Nakamura & Fukumori & Fujikawa" authorityYear="2010" box="[262,488,1066,1085]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="450" phylum="Arthropoda" rank="species" species="kumayaensis" status="sp. nov.">
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<emphasis id="B917EA9DFFE3FFE6BC7CBD64FE5FB963" box="[262,488,1066,1085]" italics="true" pageId="12" pageNumber="450">Tectocepheus kumayaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22457E6FFE3FFE6BC97BD65FD8FB960" box="[493,568,1067,1086]" pageId="12" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
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A, Solenidial region on tarsus and tibia of leg I; B, Left epimeral region.
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</paragraph>
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</caption>
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<paragraph id="8BDC368FFFE3FFE6BDBFBDD7FEABBBE1" blockId="12.[159,762,1177,1951]" pageId="12" pageNumber="450">
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Prodorsum — Rostral anterior margin without incision, broadly truncate with two concavities (
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<figureCitation id="13582A0AFFE3FFE6BFB7BDF2FF73B9A9" captionStart="FIGURE" captionStartId="11.[327,339,1845,1864]" captionTargetBox="[283,1257,403,1778]" captionTargetId="figure-4@11.[272,1316,377,1813]" captionTargetPageId="11" captionText="FIGURE 6: Tectocepheus kumayaensis sp. nov. A, Dorsal view; B, Camerostome; C, Genito-anal region." figureDoi="http://doi.org/10.5281/zenodo.5393462" httpUri="https://zenodo.org/record/5393462/files/figure.png" pageId="12" pageNumber="450">Fig. 6A</figureCitation>
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). Setae
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<emphasis id="B917EA9DFFE3FFE6BC64BDAEFE83B9A9" box="[286,308,1248,1271]" italics="true" pageId="12" pageNumber="450">ro</emphasis>
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(33 µm) and
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<emphasis id="B917EA9DFFE3FFE6BCAFBD91FE50B9A9" box="[469,487,1247,1271]" italics="true" pageId="12" pageNumber="450">le</emphasis>
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(30 µm) spiniform, extending for a short distance anterior of rostral anterior margin; setae
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<emphasis id="B917EA9DFFE3FFE6BCE8BC68FE1FB863" box="[402,424,1318,1341]" italics="true" pageId="12" pageNumber="450">ro</emphasis>
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barbed unilaterally; setae
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<emphasis id="B917EA9DFFE3FFE6BF92BC6BFD4DB863" box="[744,762,1317,1341]" italics="true" pageId="12" pageNumber="450">le</emphasis>
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roughened throughout length;
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<emphasis id="B917EA9DFFE3FFE6BF6BBC07FD90B83E" box="[529,551,1353,1376]" italics="true" pageId="12" pageNumber="450">ro</emphasis>
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nearly as long as
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<emphasis id="B917EA9DFFE3FFE6BDE5BC22FF06B8DA" box="[159,177,1388,1412]" italics="true" pageId="12" pageNumber="450">le</emphasis>
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. Lamellar cuspis without dent nor swelling, not extending to level of rostral anterior margin. Rostral and lamellar transverse ridges distinct. Setae
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<emphasis id="B917EA9DFFE3FFE6BFBEBCFCFD6DB897" box="[708,730,1458,1481]" italics="true" pageId="12" pageNumber="450">in</emphasis>
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(5 µm) small trigonal pyramidal, roughened throughout length. Sensilli (45 µm) composed of thin, roughened stem and globular head which bears dark coloured spines. Setae
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<emphasis id="B917EA9DFFE3FFE6BC9EBF71FE4BBB08" box="[484,508,1599,1622]" italics="true" pageId="12" pageNumber="450">ex</emphasis>
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(8 µm) smooth, short, longer than setae
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<emphasis id="B917EA9DFFE3FFE6BCFCBF2CFE2BBB27" box="[390,412,1634,1657]" italics="true" pageId="12" pageNumber="450">in</emphasis>
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. Bothridia opened anterolaterally, with deep incision, without swelling or projection.
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</paragraph>
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<paragraph id="8BDC368FFFE3FFE6BDBFBF98FCD6BCCA" blockId="12.[159,762,1177,1951]" lastBlockId="12.[826,1429,311,1948]" pageId="12" pageNumber="450">
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Notogaster — Depression and dorsosejugal scissure absent. Humeral region with small triangular projection. Ten pairs of notogastral setae short setiform, roughened throughout length. Lyrifissures
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<emphasis id="B917EA9DFFE3FFE6BD9DBE2CFF4DBA27" box="[231,250,1890,1913]" italics="true" pageId="12" pageNumber="450">ia</emphasis>
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aligned perpendicular to notogastral outline, antero-laterally to setae
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<emphasis id="B917EA9DFFE3FFE6BC82BECBFDB4BAC2" box="[504,515,1925,1948]" italics="true" pageId="12" pageNumber="450">c</emphasis>
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<subScript id="17E734CAFFE3FFE6BF79BEC0FDBBBAC1" attach="left" box="[515,524,1934,1951]" fontSize="7" pageId="12" pageNumber="450">2</subScript>
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;
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<emphasis id="B917EA9DFFE3FFE6BF67BECBFD8EBAC2" box="[541,569,1925,1948]" italics="true" pageId="12" pageNumber="450">im</emphasis>
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obliquely, laterally to
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<emphasis id="B917EA9DFFE3FFE6BEF2B879FC2ABC11" box="[904,925,311,335]" italics="true" pageId="12" pageNumber="450">lp</emphasis>
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;
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<emphasis id="B917EA9DFFE3FFE6BED0B879FC08BC10" box="[938,959,311,334]" italics="true" pageId="12" pageNumber="450">ip</emphasis>
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perpendicular to notogastral outline between
|
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<emphasis id="B917EA9DFFE3FFE6BEF0B815FC20BC2C" box="[906,919,347,370]" italics="true" pageId="12" pageNumber="450">p</emphasis>
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<subScript id="17E734CAFFE3FFE6BEEDB82DFC17BC2A" attach="left" box="[919,928,355,372]" fontSize="7" pageId="12" pageNumber="450">1</subScript>
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||
and
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<emphasis id="B917EA9DFFE3FFE6BEA5B815FC5BBC2C" box="[991,1004,347,370]" italics="true" pageId="12" pageNumber="450">p</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6BE96B82DFC42BC2A" attach="left" box="[1004,1013,355,372]" fontSize="7" pageId="12" pageNumber="450">2</subScript>
|
||
. Setae
|
||
<emphasis id="B917EA9DFFE3FFE6B92AB814FBEABC2C" box="[1104,1117,346,370]" italics="true" pageId="12" pageNumber="450">h</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6B924B82DFBD0BC2A" attach="left" box="[1118,1127,355,372]" fontSize="7" pageId="12" pageNumber="450">3</subScript>
|
||
inserted antero-lateral to
|
||
<emphasis id="B917EA9DFFE3FFE6BE40B833FCEDBCCB" box="[826,858,381,405]" italics="true" pageId="12" pageNumber="450">gla</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC368FFFE3FFE6BE1AB8E2FB9CBE8C" blockId="12.[826,1429,311,1948]" pageId="12" pageNumber="450">
|
||
Ventral region — Genital (36 µm) and anal (57 µm) apertures almost pentagonal in form; distance (22 µm) between them appreciably shorter than half length of anal aperture. Genito-anal setal formula: 6-1-2-3; all setae smooth setiform (
|
||
<figureCitation id="13582A0AFFE3FFE6B9A4BB76FAF7BF0E" box="[1246,1344,568,592]" captionStart="FIGURE" captionStartId="11.[327,339,1845,1864]" captionTargetBox="[283,1257,403,1778]" captionTargetId="figure-4@11.[272,1316,377,1813]" captionTargetPageId="11" captionText="FIGURE 6: Tectocepheus kumayaensis sp. nov. A, Dorsal view; B, Camerostome; C, Genito-anal region." figureDoi="http://doi.org/10.5281/zenodo.5393462" httpUri="https://zenodo.org/record/5393462/files/figure.png" pageId="12" pageNumber="450">Fig. 6C</figureCitation>
|
||
). Setae
|
||
<emphasis id="B917EA9DFFE3FFE6BE40BB12FCF0BF2D" box="[826,839,604,627]" italics="true" pageId="12" pageNumber="450">g</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6BE3DBB2BFCE7BF28" attach="left" box="[839,848,613,630]" fontSize="7" pageId="12" pageNumber="450">1</subScript>
|
||
and
|
||
<emphasis id="B917EA9DFFE3FFE6BEF1BB12FC2FBF2D" box="[907,920,604,627]" italics="true" pageId="12" pageNumber="450">g</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6BEE2BB2BFC16BF28" attach="left" box="[920,929,613,630]" fontSize="7" pageId="12" pageNumber="450">2</subScript>
|
||
inserted nearer anterior margine of plates. Setae
|
||
<emphasis id="B917EA9DFFE3FFE6BEFABB31FC2EBFC8" box="[896,921,639,662]" italics="true" pageId="12" pageNumber="450">ag</emphasis>
|
||
inserted latero-posteriorly to genital aperture. Setae
|
||
<emphasis id="B917EA9DFFE3FFE6BEB5BBEFFC5FBFE7" box="[975,1000,673,697]" italics="true" pageId="12" pageNumber="450">ad</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6BE92BBE5FC46BFE2" attach="left" box="[1000,1009,683,700]" fontSize="7" pageId="12" pageNumber="450">1</subScript>
|
||
and
|
||
<emphasis id="B917EA9DFFE3FFE6B94CBBEFFBF8BFE7" box="[1078,1103,673,697]" italics="true" pageId="12" pageNumber="450">ad</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6B935BBE5FBEFBFE2" attach="left" box="[1103,1112,683,700]" fontSize="7" pageId="12" pageNumber="450">2</subScript>
|
||
aligned in postanal position,
|
||
<emphasis id="B917EA9DFFE3FFE6BE00BB8AFC24BF82" box="[890,915,708,732]" italics="true" pageId="12" pageNumber="450">ad</emphasis>
|
||
<quantity id="4C9B9B6AFFE3FFE6BEE9BB80FC09BF82" box="[915,958,708,735]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="450" unit="in" value="3.0">
|
||
<subScript id="17E734CAFFE3FFE6BEE9BB80FC2BBF81" attach="left" box="[915,924,718,735]" fontSize="7" pageId="12" pageNumber="450">3</subScript>
|
||
in
|
||
</quantity>
|
||
adanal. Lyrifissures
|
||
<emphasis id="B917EA9DFFE3FFE6B9B9BB8AFB54BF82" box="[1219,1251,708,732]" italics="true" pageId="12" pageNumber="450">iad</emphasis>
|
||
located along, near anterior margin of anal aperture. Sternal ridge distinct at epimeres borders 2-4; bo. 1,2, sj distinct. Epimeral setal formula: 3-1-3-3; setae smooth, minute setiform. Diarthric subcapitulum bearing 3 pairs of setae:
|
||
<emphasis id="B917EA9DFFE3FFE6BE97BA3AFC4EBED5" box="[1005,1017,884,907]" italics="true" pageId="12" pageNumber="450">a</emphasis>
|
||
(13 µm),
|
||
<emphasis id="B917EA9DFFE3FFE6B915BA3AFB33BED5" box="[1135,1156,884,907]" italics="true" pageId="12" pageNumber="450">m</emphasis>
|
||
(2 µm) and
|
||
<emphasis id="B917EA9DFFE3FFE6B861BA3AFA9FBED2" box="[1307,1320,884,908]" italics="true" pageId="12" pageNumber="450">h</emphasis>
|
||
(11 µm); setae thin smooth setiform; relative lengths:
|
||
<emphasis id="B917EA9DFFE3FFE6B830BAD6FAE1BEF1" box="[1354,1366,920,943]" italics="true" pageId="12" pageNumber="450">a</emphasis>
|
||
≈ 6x
|
||
<emphasis id="B917EA9DFFE3FFE6BE40BAF4FCF0BE8C" box="[826,839,954,978]" italics="true" pageId="12" pageNumber="450">h</emphasis>
|
||
;
|
||
<emphasis id="B917EA9DFFE3FFE6BE2EBAF5FCDEBE8C" box="[852,873,955,978]" italics="true" pageId="12" pageNumber="450">m</emphasis>
|
||
≈ 4x
|
||
<emphasis id="B917EA9DFFE3FFE6BED7BAF4FC0DBE8C" box="[941,954,954,978]" italics="true" pageId="12" pageNumber="450">h</emphasis>
|
||
(
|
||
<figureCitation id="13582A0AFFE3FFE6BEB3BAF4FBA8BE8C" box="[969,1055,954,978]" captionStart="FIGURE" captionStartId="11.[327,339,1845,1864]" captionTargetBox="[283,1257,403,1778]" captionTargetId="figure-4@11.[272,1316,377,1813]" captionTargetPageId="11" captionText="FIGURE 6: Tectocepheus kumayaensis sp. nov. A, Dorsal view; B, Camerostome; C, Genito-anal region." figureDoi="http://doi.org/10.5281/zenodo.5393462" httpUri="https://zenodo.org/record/5393462/files/figure.png" pageId="12" pageNumber="450">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC368FFFE3FFE6BE1ABAA7FAEFB98D" blockId="12.[826,1429,311,1948]" pageId="12" pageNumber="450">
|
||
Legs — Monodactyl; claws with few dens. Legs not studied. Trochantera III and IV bearing carina with sharply pointed apex (
|
||
<figureCitation id="13582A0AFFE3FFE6B9CDBD61FAB8B919" box="[1207,1295,1071,1095]" captionStart="FIGURE" captionStartId="12.[159,171,1067,1086]" captionTargetBox="[169,738,485,1034]" captionTargetId="figure-32@12.[165,755,477,1035]" captionTargetPageId="12" captionText="FIGURE 7: Tectocepheus kumayaensis sp. nov. A, Solenidial region on tarsus and tibia of leg I; B, Left epimeral region." figureDoi="http://doi.org/10.5281/zenodo.5393464" httpUri="https://zenodo.org/record/5393464/files/figure.png" pageId="12" pageNumber="450">Fig. 7B</figureCitation>
|
||
). On tarsus I, famulus
|
||
<emphasis id="B917EA9DFFE3FFE6BEBBBD1CFC7AB934" box="[961,973,1106,1130]" italics="true" pageId="12" pageNumber="450">ε</emphasis>
|
||
trigonal pyramidal situated between solenidia bacilliform
|
||
<emphasis id="B917EA9DFFE3FFE6B94CBD3BFBF0B9D3" box="[1078,1095,1141,1165]" italics="true" pageId="12" pageNumber="450">ω</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6B932BD31FBE6B9CE" attach="left" box="[1096,1105,1151,1168]" fontSize="7" pageId="12" pageNumber="450">1</subScript>
|
||
and setiform
|
||
<emphasis id="B917EA9DFFE3FFE6B981BD3BFABBB9D3" box="[1275,1292,1141,1165]" italics="true" pageId="12" pageNumber="450">ω</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6B877BD31FAA1B9CE" attach="left" box="[1293,1302,1151,1168]" fontSize="7" pageId="12" pageNumber="450">2</subScript>
|
||
(
|
||
<figureCitation id="13582A0AFFE3FFE6B852BD3BFA3FB9D3" box="[1320,1416,1141,1165]" captionStart="FIGURE" captionStartId="12.[159,171,1067,1086]" captionTargetBox="[169,738,485,1034]" captionTargetId="figure-32@12.[165,755,477,1035]" captionTargetPageId="12" captionText="FIGURE 7: Tectocepheus kumayaensis sp. nov. A, Solenidial region on tarsus and tibia of leg I; B, Left epimeral region." figureDoi="http://doi.org/10.5281/zenodo.5393464" httpUri="https://zenodo.org/record/5393464/files/figure.png" pageId="12" pageNumber="450">Fig. 7A</figureCitation>
|
||
). Solenidion
|
||
<emphasis id="B917EA9DFFE3FFE6BEC4BDD6FC78B9EE" box="[958,975,1176,1200]" italics="true" pageId="12" pageNumber="450">’</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6BEB5BDECFC6FB9ED" attach="left" box="[975,984,1186,1203]" fontSize="7" pageId="12" pageNumber="450">1</subScript>
|
||
on tibia I originating from apophysis, about three times longer than the length of
|
||
<emphasis id="B917EA9DFFE3FFE6B84FBDF5FAF1B98D" box="[1333,1350,1211,1235]" italics="true" pageId="12" pageNumber="450">ω</emphasis>
|
||
<subScript id="17E734CAFFE3FFE6B83DBD8BFAE7B988" attach="left" box="[1351,1360,1221,1238]" fontSize="7" pageId="12" pageNumber="450">1</subScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC368FFFE3FFE7BE1ABDA4FEA3B87A" blockId="12.[826,1429,311,1948]" lastBlockId="13.[159,762,311,1317]" lastPageId="13" lastPageNumber="451" pageId="12" pageNumber="450">
|
||
Remarks — Rostrum with two concavities of the new species is similar in appearance to those of
|
||
<taxonomicName id="4C634D0CFFE3FFE6B813BC40FB23B816" authority="Berlese, 1913" authorityName="Berlese" authorityYear="1913" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="450" phylum="Arthropoda" rank="species" species="alatus">
|
||
<emphasis id="B917EA9DFFE3FFE6B813BC40FC59B816" italics="true" pageId="12" pageNumber="450">Tectocepheus alatus</emphasis>
|
||
<bibRefCitation id="EFF24B7EFFE3FFE6BE82BC7EFB23B816" author="Berlese A." box="[1016,1172,1328,1352]" pageId="12" pageNumber="450" pagination="77 - 111" refId="ref14246" refString="Berlese A. 1913 - - Acari nuovi. Manipoli VII-VIII - Redia, 9: 77 - 111, pls. 1 - 8." type="journal article" year="1913">Berlese, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D0CFFE3FFE6B9ADBC7FFC58B832" authority="Fujikawa, 2001" authorityName="Fujikawa" authorityYear="2001" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="450" phylum="Arthropoda" rank="species" species="shirakamiensis">
|
||
<emphasis id="B917EA9DFFE3FFE6B9ADBC7FFA22B816" box="[1239,1429,1328,1352]" italics="true" pageId="12" pageNumber="450">T. shirakamiensis</emphasis>
|
||
<bibRefCitation id="EFF24B7EFFE3FFE6BE40BC1DFC58B832" author="Fujikawa T." box="[826,1007,1363,1388]" pageId="12" pageNumber="450" pagination="23 - 30" refId="ref14602" refString="Fujikawa T. 2001 - A new and three known species of Tectocepheidae and Nodocepheidae from northeastern part of Nippon including the Shirakami-sanchi World Heritage Area (Acari: Oribatida) - Edaphologia, (67): 23 - 30." type="journal article" year="2001">Fujikawa, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, the new species differs from them in form of cuspis, dorsosejugal region, humeral region, and situation or direction of lyrifissures
|
||
<emphasis id="B917EA9DFFE3FFE6BEE4BCF3FC06B88A" box="[926,945,1469,1492]" italics="true" pageId="12" pageNumber="450">ia</emphasis>
|
||
and
|
||
<emphasis id="B917EA9DFFE3FFE6BE8FBCF3FBA2B88B" box="[1013,1045,1469,1493]" italics="true" pageId="12" pageNumber="450">iad</emphasis>
|
||
. The new species has a small body size, verrucose globular sensilli and subpallel lyrifissures
|
||
<emphasis id="B917EA9DFFE3FFE6BE93BF4DFBBEBB45" box="[1001,1033,1539,1563]" italics="true" pageId="12" pageNumber="450">iad</emphasis>
|
||
to anal aperture as is the case of
|
||
<taxonomicName id="4C634D0CFFE3FFE6BE40BF69FBF2BB60" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[826,1093,1574,1598]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="450" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B917EA9DFFE3FFE6BE40BF69FC2BBB60" box="[826,924,1575,1598]" italics="true" pageId="12" pageNumber="450">T. minor</emphasis>
|
||
<bibRefCitation id="EFF24B7EFFE3FFE6BED2BF68FBF2BB60" author="Berlese A." box="[936,1093,1574,1598]" pageId="12" pageNumber="450" pagination="235 - 252" refId="ref14201" refString="Berlese A. 1903 (1904) - - Spicilegia Zoologica. Acari Nuovi - Redia, 1: 235 - 252." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
[1904]
|
||
<emphasis id="B917EA9DFFE3FFE6B9D9BF69FB57BB60" box="[1187,1248,1575,1598]" italics="true" pageId="12" pageNumber="450">sensu</emphasis>
|
||
<bibRefCitation id="EFF24B7EFFE3FFE6B996BF68FA22BB60" author="Bernini, F." box="[1260,1429,1574,1598]" pageId="12" pageNumber="450" refId="ref14294" refString="Bernini, F. 1973 - Notulae Oribatologicae VII. Gli Oribatei (Acarida) dell'isolotto di Basiluzzo (Isole Eolie) - Lav. Soc. It. Biogeogr. N. S., III: 355 - 480, Tav. I - XVII." type="book" year="1973">Bernini (1973)</bibRefCitation>
|
||
and Japanese specimens collected by Fujikawa. At first she considered these Japanese specimens as
|
||
<taxonomicName id="4C634D0CFFE3FFE6B8FABF23FA99BBF9" authority="Knulle, 1954" authorityName="Knulle" authorityYear="1954" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="450" phylum="Arthropoda" rank="species" species="cuspidentatus">
|
||
<emphasis id="B917EA9DFFE3FFE6B8FABF23FC67BBF9" italics="true" pageId="12" pageNumber="450">T. cuspidentatus</emphasis>
|
||
<bibRefCitation id="EFF24B7EFFE3FFE6BEAFBFC1FBD1BBF9" author="Knulle W." box="[981,1126,1679,1703]" pageId="12" pageNumber="450" pagination="280 - 305" refId="ref15130" refString="Knulle W. 1954 - Die Arten der Gattung Tectocepheus Berlese (Acarina: Oribatei) - Zool. Anz., 152: 280 - 305." type="journal article" year="1954">Knülle, 1954</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFF24B7EFFE3FFE6B90EBFC1FA91BBF9" author="Fujikawa T." box="[1140,1318,1679,1703]" pageId="12" pageNumber="450" pagination="205 - 213" refId="ref14492" refString="Fujikawa T. 1988 - Two species belonging to the genus Tectocepheus from nature farm at Nayoro in Northern Japan (Acari; Oribatei) - Acarologia, 29 (2): 205 - 213." type="journal article" year="1988">Fujikawa, 1988</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, because (1) she could not find out similar characters like rostrum with two deep incisions between Japanese specimens and Berlese Collection (204/6, 217/27- 30 and 82/36-40) in addition slides in the Berlese Collection were hardly examinable for her, and (2) she could find that Japanese specimens beared a stricking resemblance in the rostrum shape, sensilli and lamellar cusps, and situation of lyrifissures
|
||
<emphasis id="B917EA9DFFE2FFE7BFA3B879FD4EBC11" box="[729,761,311,335]" italics="true" pageId="13" pageNumber="451">iad</emphasis>
|
||
with a specimen of
|
||
<taxonomicName id="4C634D0CFFE2FFE7BCE2B815FDF8BC2C" authorityName="Knulle" authorityYear="1954" box="[408,591,346,370]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="cuspidentatus">
|
||
<emphasis id="B917EA9DFFE2FFE7BCE2B815FDF8BC2C" box="[408,591,346,370]" italics="true" pageId="13" pageNumber="451">T. cuspidentatus</emphasis>
|
||
</taxonomicName>
|
||
in Zoological Museum Copenhagen. Latter the Japanese specimens were identified as
|
||
<taxonomicName id="4C634D0CFFE2FFE7BCB0B8EFFD4EBCE6" authority="(Fujikawa, 2001)" baseAuthorityName="Fujikawa" baseAuthorityYear="2001" box="[458,761,416,440]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B917EA9DFFE2FFE7BCB0B8EFFD9DBCE6" box="[458,554,417,440]" italics="true" pageId="13" pageNumber="451">T. minor</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF24B7EFFE2FFE7BF46B8EEFD45BCE6" author="Fujikawa T." box="[572,754,416,440]" pageId="13" pageNumber="451" pagination="23 - 30" refId="ref14602" refString="Fujikawa T. 2001 - A new and three known species of Tectocepheidae and Nodocepheidae from northeastern part of Nippon including the Shirakami-sanchi World Heritage Area (Acari: Oribatida) - Edaphologia, (67): 23 - 30." type="journal article" year="2001">Fujikawa, 2001</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
according to
|
||
<bibRefCitation id="EFF24B7EFFE2FFE7BC44B88DFDD0BC85" author="Nubel-Reidelbach E." box="[318,615,451,475]" pageId="13" pageNumber="451" pagination="3 - 94" refId="ref15511" refString="Nubel-Reidelbach E. 1994 - Taxonomie und Systematik der Gattung Tectocepheus Berlese, 1895 (Acari, Oribatei) - andrias, Staatliches Museum fur Naturkunde Karlsruhe, 12: 3 - 94." type="journal article" year="1994">Nübel-Reidelbach (1994)</bibRefCitation>
|
||
. After that, Mahunk and Mahunka-Papp (1995) and Laumann and al (2007) pointed out that
|
||
<taxonomicName id="4C634D0CFFE2FFE7BF7ABB44FDEABF7F" baseAuthorityName="Fujikawa" baseAuthorityYear="2001" box="[512,605,522,545]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B917EA9DFFE2FFE7BF7ABB44FDEABF7F" box="[512,605,522,545]" italics="true" pageId="13" pageNumber="451">T. minor</emphasis>
|
||
</taxonomicName>
|
||
has medially weakly protruding rostral anterial margin, cusps bearing inner and outer dens, globular sensilli, notogaster without depressions, bothridium ventrally expanded, trochanter IV bearing a dorsal spur and lyrifissures
|
||
<emphasis id="B917EA9DFFE2FFE7BC5CBBF6FEF1BF8E" box="[294,326,696,720]" italics="true" pageId="13" pageNumber="451">iad</emphasis>
|
||
situated subparallel to anal aperture.
|
||
<bibRefCitation id="EFF24B7EFFE2FFE7BDE5BB92FEE5BFAD" author="Fujikawa T." box="[159,338,731,756]" pageId="13" pageNumber="451" pagination="1 - 82" refId="ref14568" refString="Fujikawa T. 1995 - Comparison among populations of Tectocepheus velatus (Michael, 1880) from forests, grasslands and crop field - Edaphologia, (55): 1 - 82." type="journal article" year="1995">Fujikawa (1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF24B7EFFE2FFE7BC25BB92FE22BFAA" author="Fujikawa T." box="[351,405,732,756]" pageId="13" pageNumber="451" pagination="11 - 46" refId="ref14529" refString="Fujikawa T. 1999 - Individual variations of two reared oribatid species, Tectocepheus velatus (Michael, 1880) and Oppiella nova (Oudemans, 1902) - Edaphologia, (62): 11 - 46." type="journal article" year="1999">1999</bibRefCitation>
|
||
) could not observe individual variation in presence or absence of rostral incisions, form of rostral anterior margin, presence or absence of dens of cusps, and situation of
|
||
<emphasis id="B917EA9DFFE2FFE7BF4ABA0BFDE7BE03" box="[560,592,837,861]" italics="true" pageId="13" pageNumber="451">iad</emphasis>
|
||
subparallel to anal aperture. Now we consider that the abovementioned Japanese specimens should be reidentified as
|
||
<taxonomicName id="4C634D0CFFE2FFE7BD80BAE1FE18BE98" authorityName="Knulle" authorityYear="1954" box="[250,431,942,966]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="cuspidentatus">
|
||
<emphasis id="B917EA9DFFE2FFE7BD80BAE1FE18BE98" box="[250,431,942,966]" italics="true" pageId="13" pageNumber="451">T. cuspidentatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C634D0CFFE2FFE7BCB3BAE0FD28BE98" baseAuthorityName="Fujikawa" baseAuthorityYear="2001" box="[457,671,942,966]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B917EA9DFFE2FFE7BCB3BAE0FD28BE98" box="[457,671,942,966]" italics="true" pageId="13" pageNumber="451">Tectocepheus minor</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFF24B7EFFE2FFE7BFA8BAE0FE7FBEB7" author="Fujita, M. & Fujiyama, S." pageId="13" pageNumber="451" pagination="36 - 45" refId="ref14702" refString="Fujita, M., Fujiyama, S. 2001 - How can the minor species, Tectocepheus minor (Oribatida), dominate T. velatus in a no-tillage crop field? - Pedobiologia, 45: 36 - 45." type="journal article" year="2001">Fujita and Fujiyama (2001)</bibRefCitation>
|
||
should be also included in the species,
|
||
<taxonomicName id="4C634D0CFFE2FFE7BC36BABBFF51B971" authority=", Fujita and Fujiyama (2001)" authorityName=", Fujita and Fujiyama" authorityYear="2001" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="cuspidentatus">
|
||
<emphasis id="B917EA9DFFE2FFE7BC36BABBFE48B952" box="[332,511,1012,1036]" italics="true" pageId="13" pageNumber="451">T. cuspidentatus</emphasis>
|
||
,
|
||
<bibRefCitation id="EFF24B7EFFE2FFE7BF77BABAFF51B971" author="Fujita, M. & Fujiyama, S." pageId="13" pageNumber="451" pagination="36 - 45" refId="ref14702" refString="Fujita, M., Fujiyama, S. 2001 - How can the minor species, Tectocepheus minor (Oribatida), dominate T. velatus in a no-tillage crop field? - Pedobiologia, 45: 36 - 45." type="journal article" year="2001">Fujita and Fujiyama (2001)</bibRefCitation>
|
||
</taxonomicName>
|
||
suggested that, unlike
|
||
<taxonomicName id="4C634D0CFFE2FFE7BC86BD56FDD1B971" baseAuthorityName="Michael" baseAuthorityYear="1880" box="[508,614,1047,1071]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="velatus">
|
||
<emphasis id="B917EA9DFFE2FFE7BC86BD56FDD1B971" box="[508,614,1047,1071]" italics="true" pageId="13" pageNumber="451">T. velatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0CFFE2FFE7BF0FBD56FF61B90C" authorityName=", Fujita and Fujiyama" authorityYear="2001" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="451" phylum="Arthropoda" rank="species" species="cuspidentatus">
|
||
<emphasis id="B917EA9DFFE2FFE7BF0FBD56FF61B90C" italics="true" pageId="13" pageNumber="451">T. cuspidentatus</emphasis>
|
||
</taxonomicName>
|
||
could migrate in response to seasonal environmental changes. The new species differs from all the species of the genus in having rostral anterior margin with two concavities without incisions, cusps without dens and swelling, trigonal pyramidal interlamellar setae, and form of carina on trochanter III and IV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |