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<document id="DA4C61D237F2FC2E3C548C65999D6E04" ID-DOI="10.5281/zenodo.282460" ID-GBIF-Dataset="15d39ae0-4f0a-4ecc-83ab-2f99ae496d37" ID-ISSN="1175-5326" ID-Zenodo-Dep="282460" ID-ZooBank="9F22E784-360E-4DEB-A306-868F014D5F4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460536120703" checkinUser="plazi" docAuthor="Hamilton, Andrew" docDate="2012" docId="03C64273FFEE37578EFEFD3AFE1FFA16" docLanguage="en" docName="zt03497p059.pdf" docOrigin="Zootaxa 3497" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cephisus Stal" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFFF3A0BFFE837518E69FFB5FFDAFFA1" masterDocTitle="Revision of Neotropical aphrophorine spittlebugs, part 1: Ptyelini (Hemiptera, Cercopoidea)" masterLastPageNumber="59" masterPageNumber="41" pageNumber="47" updateTime="1698316418071" updateUser="plazi">
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<mods:title id="5320292DDF46C19E0DEA7E179F5233C1">Revision of Neotropical aphrophorine spittlebugs, part 1: Ptyelini (Hemiptera, Cercopoidea)</mods:title>
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<mods:date id="97CA4028E9B71BBC3B0A5944A51D81B6">2012</mods:date>
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<treatment id="03C64273FFEE37578EFEFD3AFE1FFA16" ID-DOI="http://doi.org/10.5281/zenodo.6173458" ID-GBIF-Taxon="119527692" ID-Zenodo-Dep="6173458" LSID="urn:lsid:plazi:treatment:03C64273FFEE37578EFEFD3AFE1FFA16" httpUri="http://treatment.plazi.org/id/03C64273FFEE37578EFEFD3AFE1FFA16" lastPageNumber="47" pageId="6" pageNumber="47">
<subSubSection id="C375A0EEFFEE37578EFEFD3AFEE7FD08" box="[151,317,655,681]" pageId="6" pageNumber="47" type="nomenclature">
<paragraph id="8BD0F365FFEE37578EFEFD3AFEE7FD08" blockId="6.[151,317,655,681]" box="[151,317,655,681]" pageId="6" pageNumber="47">
<heading id="D0984409FFEE37578EFEFD3AFEE7FD08" bold="true" box="[151,317,655,681]" fontSize="11" level="1" pageId="6" pageNumber="47" reason="1">
<taxonomicName id="4C6F88E6FFEE37578EFEFD3AFEE7FD08" ID-CoL="3KKC" ID-ENA="295234" authority="Stal" authorityName="Stal" box="[151,317,655,681]" class="Insecta" family="Aphrophoridae" genus="Cephisus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91B2F77FFEE37578EFEFD3AFEE7FD08" bold="true" box="[151,317,655,681]" pageId="6" pageNumber="47">
<emphasis id="B91B2F77FFEE37578EFEFD3AFEDCFD08" bold="true" box="[151,262,655,681]" italics="true" pageId="6" pageNumber="47">Cephisus</emphasis>
Stål
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375A0EEFFEE37578EFEFD62FE8FFCAA" pageId="6" pageNumber="47" type="reference_group">
<paragraph id="8BD0F365FFEE37578EFEFD62FE8FFCAA" blockId="6.[151,1436,726,779]" pageId="6" pageNumber="47">
<treatmentCitationGroup id="AB7FD44BFFEE37578EFEFD62FE8FFCAA" pageId="6" pageNumber="47">
<taxonomicName id="4C6F88E6FFEE37578EFEFD62FE5CFD4D" ID-CoL="3KKC" authority="Stal, 1866:67" authorityName="Stal" authorityPageNumber="67" authorityYear="1866" box="[151,390,726,749]" class="Insecta" family="Aphrophoridae" genus="Cephisus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91B2F77FFEE37578EFEFD62FF2EFD4C" box="[151,244,727,749]" italics="true" pageId="6" pageNumber="47">Cephisus</emphasis>
<treatmentCitation id="0ACED574FFEE37578E95FD63FE5CFD4D" author="Stal" box="[252,390,726,748]" page="67" pageId="6" pageNumber="47" year="1866">
<bibRefCitation id="EFFE8E94FFEE37578E95FD63FEB2FD4D" author="Stal" box="[252,360,726,748]" pageId="6" pageNumber="47" refString="Stal, C. (1866) Analecta Hemipterologica. Berliner Entomologische Zeitschrift, 10, 151 - 172, 281 - 394." type="journal article" year="1866">Stål, 1866</bibRefCitation>
:67
</treatmentCitation>
</taxonomicName>
. Validation by inclusion of species:
<treatmentCitation id="0ACED574FFEE37578C94FD63FC55FD4D" author="Stal" box="[765,911,726,748]" page="384" pageId="6" pageNumber="47" year="1866">
<bibRefCitation id="EFFE8E94FFEE37578C94FD63FCB8FD4D" author="Stal" box="[765,866,726,748]" pageId="6" pageNumber="47" refString="Stal, C. (1866) Analecta Hemipterologica. Berliner Entomologische Zeitschrift, 10, 151 - 172, 281 - 394." type="journal article" year="1866">Stål 1866</bibRefCitation>
:384
</treatmentCitation>
. Type-species by monotypy:
<taxonomicName id="4C6F88E6FFEE37578AD5FD62FE8BFCAA" ID-CoL="FKKV" authority="Walker, 1851" authorityName="Walker" authorityYear="1851" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="siccifolia">
<emphasis id="B91B2F77FFEE37578AD5FD62FA46FD4C" box="[1212,1436,727,749]" italics="true" pageId="6" pageNumber="47">Aphrophora siccifolia</emphasis>
<bibRefCitation id="EFFE8E94FFEE37578EAEFD40FE8BFCAA" author="Walker" box="[199,337,757,779]" pageId="6" pageNumber="47" refString="Walker, F. (1851) List of the specimens of homopterous insects in the collection of the British Museum, 3, 637 - 907." type="journal article" year="1851">Walker, 1851</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C375A0EEFFEE37578EFEFC82FC74FCEE" box="[151,942,823,848]" pageId="6" pageNumber="47" type="distribution">
<paragraph id="8BD0F365FFEE37578EFEFC82FC74FCEE" blockId="6.[151,1437,823,1463]" box="[151,942,823,848]" pageId="6" pageNumber="47">
<emphasis id="B91B2F77FFEE37578EFEFC82FEEAFCF1" bold="true" box="[151,304,823,848]" pageId="6" pageNumber="47">Distribution.</emphasis>
New world tropics north to southern Texas and Arizona.
</paragraph>
</subSubSection>
<subSubSection id="C375A0EEFFEE37578EAEFCEEFEB2FA86" pageId="6" pageNumber="47" type="diagnosis">
<paragraph id="8BD0F365FFEE37578EAEFCEEFEB2FA86" blockId="6.[151,1437,823,1463]" pageId="6" pageNumber="47">
<emphasis id="B91B2F77FFEE37578EAEFCEEFE9BFCD5" bold="true" box="[199,321,859,884]" pageId="6" pageNumber="47">Diagnosis.</emphasis>
The only aphrophorine spittlebug in the new world with a median carina on the face. Crown declivous; face more or less carinate on midline, at least on lower half of frons; head much narrower than pronotum; antennal ledges finely unicarinate; frons with low, rounded median ridge; clypellus tip narrow, extending between fore coxae; lora discoid; genae strongly constricted next to widest part of lora, broad below; rostrum extending to mesocoxae, or to mesotrochanters in
<taxonomicName id="4C6F88E6FFEE37578D3EFC58FB82FBA5" authority="Walker" authorityName="Walker" box="[855,1112,1004,1028]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="siccifolia">
<emphasis id="B91B2F77FFEE37578D3EFC58FC3FFBA5" box="[855,997,1004,1028]" italics="true" pageId="6" pageNumber="47">C. siccifolia</emphasis>
(Walker)
</taxonomicName>
. Pronotum with shallow to prominent median fossa near fore border. Tegmina strongly narrowed to obliquely truncate tips.
<collectingCountry id="F378B3F5FFEE37578AB7FBBAFACCFB86" box="[1246,1302,1039,1063]" name="India" pageId="6" pageNumber="47">Hind</collectingCountry>
wings with circumambient vein sinuate at end of each cell between Cu1 and vannal fold (
<figureCitation id="1354EFE0FFEE37578A72FB81FBB8FBED" box="[1051,1122,1076,1100]" captionStart="FIGURES 3 4" captionStartId="4.[151,264,1704,1727]" captionTargetBox="[175,1408,326,1677]" captionTargetId="figure@4.[169,1419,317,1697]" captionTargetPageId="4" captionText="FIGURES 3 4. Morphology of head (3) and wings (4) of Ptyelini. 3 A. face of Cephisus sanguisuga, holotype; 3 B. head and pronotum, ventrolateral aspect, of C. xanthocephala, holotype; 3 C. head and propleuron of unidentified Cercopinae; 4 A. tegmen of C. siccifolia, ventral aspect showing ventral flange (“ Sc ” of authors); 4 B. hindwing of same; 4 C. same, of Ptyelus sp.; 4 D. same, of Eulepyronia grossa Schmidt, holotype." httpUri="https://zenodo.org/record/282463/files/figure.png" pageId="6" pageNumber="47">Fig. 4</figureCitation>
B).
<collectingCountry id="F378B3F5FFEE37578AE4FB81FB1FFBED" box="[1165,1221,1076,1100]" name="India" pageId="6" pageNumber="47">Hind</collectingCountry>
tibiae each with a double pecten of 1415 black-tipped spines; basal tarsomeres each with 1 pecten row of 67 such spines; 2nd tarsomeres each with 79 tiny spines set in deeply V-notched border. Male anal tube surmounted by hoodlike articulated base (
<figureCitation id="1354EFE0FFEE37578F31FB17FE47FB1B" box="[344,413,1186,1210]" captionStart="FIGURES 5 15" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[221,1369,524,1773]" captionTargetId="figure@5.[217,1369,519,1863]" captionTargetPageId="5" captionText="FIGURES 5 15. Male genitalia of Ptyelini in lateral aspect (A), caudal aspect of theca (B) and widest aspect of style (C). 5. genital capsule of Ptyelus; 6. same, of Cephisus; 7. syntype male of C. sanguisuga; 8. C. diminutus, holotype; 9. C. laticeps, holotype; 10. C. xanthocephala; 11. C. magnificus, holotype; 12. C. variolosus; 13. C. siccifolia; 14. C. jacobii; 15. C. brevipennis, holotype." httpUri="https://zenodo.org/record/282464/files/figure.png" pageId="6" pageNumber="47">Fig. 6</figureCitation>
A); segment X very short; subgenital plates truncate; styles very large, with long setae on inner faces, apices with tiny process on caudal margin and large dorsal process hooked mesad (
<figureCitation id="1354EFE0FFEE37578AACFB72FAE2FB7E" box="[1221,1336,1223,1247]" captionStart="FIGURES 5 15" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[221,1369,524,1773]" captionTargetId="figure@5.[217,1369,519,1863]" captionTargetPageId="5" captionText="FIGURES 5 15. Male genitalia of Ptyelini in lateral aspect (A), caudal aspect of theca (B) and widest aspect of style (C). 5. genital capsule of Ptyelus; 6. same, of Cephisus; 7. syntype male of C. sanguisuga; 8. C. diminutus, holotype; 9. C. laticeps, holotype; 10. C. xanthocephala; 11. C. magnificus, holotype; 12. C. variolosus; 13. C. siccifolia; 14. C. jacobii; 15. C. brevipennis, holotype." httpUri="https://zenodo.org/record/282464/files/figure.png" pageId="6" pageNumber="47">Figs 715</figureCitation>
C); theca long, curved caudad, armed with 2 long spines, mesal pair retrorse, lateral pair divergent or curled around shaft (Figs (615)AB).
</paragraph>
</subSubSection>
<subSubSection id="C375A0EEFFEE37578EAEFA86FE1FFA16" pageId="6" pageNumber="47" type="discussion">
<paragraph id="8BD0F365FFEE37578EAEFA86FE1FFA16" blockId="6.[151,1437,823,1463]" pageId="6" pageNumber="47">
<emphasis id="B91B2F77FFEE37578EAEFA86FEE1FAEA" bold="true" box="[199,315,1331,1355]" pageId="6" pageNumber="47">Remarks.</emphasis>
Similar to the Old World genus
<taxonomicName id="4C6F88E6FFEE37578CAAFA86FCB4FAEA" box="[707,878,1331,1355]" class="Insecta" family="Aphrophoridae" genus="Cnemidanomia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91B2F77FFEE37578CAAFA86FCB4FAEA" box="[707,878,1331,1355]" italics="true" pageId="6" pageNumber="47">Cnemidanomia</emphasis>
</taxonomicName>
, but face medially carinate, body broader, with pronotum distinctly wider than head (
<figureCitation id="1354EFE0FFEE37578C20FAE2FD4AFACE" box="[585,656,1367,1391]" captionStart="FIGURE 2" captionStartId="3.[151,250,928,951]" captionTargetBox="[217,1369,193,906]" captionTargetId="figure@3.[217,1369,193,907]" captionTargetPageId="3" captionText="FIGURE 2. Head and pronotum, dorsolateral (perpendicular) aspect. A. C. laticeps, holotype; B. C. siccifolia; C. C. sanguisuga from Amazonian basin; D-F. C. variolosus complex (D. specimen near C. brevipennis from Guerrero; E. specimen # CNCHB 1905 11 from Chiapas; F. specimen # CNCHB 1906 11 from Veracruz)." httpUri="https://zenodo.org/record/282462/files/figure.png" pageId="6" pageNumber="47">Fig. 2</figureCitation>
), hump-backed, usually with long lateral margins of pronotum, and male anal tube with hoodlike articulated base (
<figureCitation id="1354EFE0FFEE37578CD1FACFFD24FA33" box="[696,766,1402,1426]" captionStart="FIGURES 5 15" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[221,1369,524,1773]" captionTargetId="figure@5.[217,1369,519,1863]" captionTargetPageId="5" captionText="FIGURES 5 15. Male genitalia of Ptyelini in lateral aspect (A), caudal aspect of theca (B) and widest aspect of style (C). 5. genital capsule of Ptyelus; 6. same, of Cephisus; 7. syntype male of C. sanguisuga; 8. C. diminutus, holotype; 9. C. laticeps, holotype; 10. C. xanthocephala; 11. C. magnificus, holotype; 12. C. variolosus; 13. C. siccifolia; 14. C. jacobii; 15. C. brevipennis, holotype." httpUri="https://zenodo.org/record/282464/files/figure.png" pageId="6" pageNumber="47">Fig. 6</figureCitation>
A) probably homologous with basal half of segment X in
<taxonomicName id="4C6F88E6FFEE37578EFEFA2AFE98FA16" box="[151,322,1439,1463]" class="Insecta" family="Aphrophoridae" genus="Cnemidanomia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91B2F77FFEE37578EFEFA2AFE98FA16" box="[151,322,1439,1463]" italics="true" pageId="6" pageNumber="47">Cnemidanomia</emphasis>
</taxonomicName>
(
<figureCitation id="1354EFE0FFEE37578F38FA2AFE7EFA16" box="[337,420,1439,1463]" captionStart="FIGURES 16 19" captionStartId="8.[151,264,1942,1965]" captionTargetBox="[313,1267,498,1917]" captionTargetId="figure@8.[313,1273,481,1921]" captionTargetPageId="8" captionText="FIGURES 16 19. Habitus of genera of Ptyelini (A), male genitalic capsule in lateral aspect (B) and male theca (C) in caudal aspect. 16. Mandesa; 17. Cnemidanomia; 18. Novaphrophara; 19. Hymettus." httpUri="https://zenodo.org/record/282465/files/figure.png" pageId="6" pageNumber="47">Fig. 17</figureCitation>
B).
</paragraph>
</subSubSection>
</treatment>
</document>