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<document id="442E2F4748732F1F6C55E3ED54FB7D63" ID-DOI="10.5281/zenodo.5319073" ID-ISSN="0374-1036" ID-Zenodo-Dep="5319073" ID-ZooBank="D78BA247-CF17-40E7-BB2D-399F1D778793D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630109735402" checkinUser="marcus" docAuthor="Kment, Petr" docDate="2015" docId="03C5FB49A96398157A834424FF10FE48" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.591-624.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Nene undulatum Kment 2015, sp. nov." docType="treatment" docVersion="4" lastPageNumber="619" masterDocId="FFFC8331A97698097B4D4038FFECFFFB" masterDocTitle="Two new genera of Madagascan Pentatominae (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="624" masterPageNumber="591" pageNumber="612" updateTime="1699154704771" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2536BD6BE73BF543E5FD352C369BBD74">Two new genera of Madagascan Pentatominae (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="BBCCF77BC2360036C24B8AC040EF7A66">Kment, Petr</mods:namePart>
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<mods:typeOfResource id="9AA26F3C6173C66DE2309144C264022D">text</mods:typeOfResource>
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<mods:title id="4FB626143EC6A6C2F62BBF00B1572077">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="1EC2B3DAE4F6A6AD8E121F05A194E542">
<mods:date id="D46A83EE3A8777E6B04274A64EB53EA0">2015</mods:date>
<mods:detail id="CAD378401652048E365A9518B6CA9DC3" type="pubDate">
<mods:number id="9C01E0C1DEDBF1931FB3376180E08B15">2015-12-31</mods:number>
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<mods:identifier id="2404BEBE256EA7EAB6B2950185BE4B0C" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03C5FB49A96398157A834424FF10FE48" LSID="urn:lsid:plazi:treatment:03C5FB49A96398157A834424FF10FE48" httpUri="http://treatment.plazi.org/id/03C5FB49A96398157A834424FF10FE48" lastPageId="28" lastPageNumber="619" pageId="21" pageNumber="612">
<subSubSection id="C37619D4A963981C7A834424FD08FBC8" box="[462,740,1052,1075]" pageId="21" pageNumber="612" type="nomenclature">
<paragraph id="8BD34A5FA963981C7A834424FD08FBC8" blockId="21.[462,740,1052,1075]" box="[462,740,1052,1075]" pageId="21" pageNumber="612">
<heading id="D09BFD33A963981C7A834424FD08FBC8" bold="true" box="[462,740,1052,1075]" centered="true" fontSize="8" level="2" pageId="21" pageNumber="612" reason="3">
<emphasis id="B918964DA963981C7A834424FD08FBC8" bold="true" box="[462,740,1052,1075]" pageId="21" pageNumber="612">
<taxonomicName id="4C6C31DCA963981C7A834424FD64FBC8" authority="Kment, 2015" authorityName="Kment" authorityYear="2015" box="[462,648,1052,1075]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="612" phylum="Arthropoda" rank="species" species="undulatum" status="sp. nov.">
<emphasis id="B918964DA963981C7A834424FD64FBC8" bold="true" box="[462,648,1052,1075]" italics="true" pageId="21" pageNumber="612">Nene undulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B2B36A963981C79C24424FD08FBC8" box="[655,740,1052,1075]" pageId="21" pageNumber="612" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37619D4A963981C7B3C4471FE5DF9D9" pageId="21" pageNumber="612" type="materials_examined">
<paragraph id="8BD34A5FA963981C7B3C4471FCC6FBA7" blockId="21.[113,1090,1097,1570]" box="[113,810,1097,1116]" pageId="21" pageNumber="612">
<emphasis id="B918964DA963981C7B3C4471FF06FBA7" bold="true" box="[113,234,1097,1116]" pageId="21" pageNumber="612">
<typeStatus id="54D7F4FDA963981C7B3C4471FF72FBA7" box="[113,158,1097,1116]" pageId="21" pageNumber="612">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F37B0ACFA963981C7BA24471FEBBFBA7" box="[239,343,1097,1116]" name="Madagascar" pageId="21" pageNumber="612">Madagascar</collectingCountry>
,
<collectingRegion id="49A884BDA963981C7A2D4471FDE3FBA7" box="[352,527,1097,1116]" country="Madagascar" name="Toamasina" pageId="21" pageNumber="612">Toamasina Province</collectingRegion>
, Rogez,
<geoCoordinate id="EE582C98A963981C79144471FD57FBA7" box="[601,699,1097,1116]" degrees="18" direction="south" minutes="48" orientation="latitude" pageId="21" pageNumber="612" precision="15" seconds="00" value="-18.8">18°4800″S</geoCoordinate>
<geoCoordinate id="EE582C98A963981C798D4471FCC9FBA7" box="[704,805,1097,1116]" degrees="48" direction="east" minutes="37" orientation="longitude" pageId="21" pageNumber="612" precision="15" seconds="06" value="48.618332">48°3706″E</geoCoordinate>
.
</paragraph>
<paragraph id="8BD34A5FA963981C7B3C445CFE5DF9D9" blockId="21.[113,1090,1097,1570]" pageId="21" pageNumber="612">
<emphasis id="B918964DA963981C7B3C445CFF1AFB8C" bold="true" box="[113,246,1124,1143]" pageId="21" pageNumber="612">Type material.</emphasis>
<materialsCitation id="3B044002A963981C7BB1445CFE1BFB69" collectionCode="NMPC" pageId="21" pageNumber="612" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D7F4FDA963981C7BB1445CFEA3FB8E" box="[252,335,1124,1143]" pageId="21" pageNumber="612" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED7DD29AA963981C7A3B445CFE5BFB8C" box="[374,439,1124,1143]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="612">NMPC</collectionCode>
), Rogez / Madagascar / Mus.Praha [p] // COLLECTIO / NATIONAL MU- SEUM / Praha, Czech Republic [p] // J [p] //
</materialsCitation>
<typeStatus id="54D7F4FDA963981C7AB04447FD90FB69" box="[509,636,1151,1170]" pageId="21" pageNumber="612" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="4C6C31DCA963981C79C04447FD28FB69" authorityName="Kment" authorityYear="2015" box="[653,708,1151,1170]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="617" phylum="Arthropoda" rank="genus">
<emphasis id="B918964DA963981C79C04447FD28FB69" box="[653,708,1151,1170]" italics="true" pageId="21" pageNumber="612">NENE</emphasis>
</taxonomicName>
/
<emphasis id="B918964DA963981C79984447FCBBFB69" box="[725,855,1151,1170]" italics="true" pageId="21" pageNumber="612">UNDULATUM</emphasis>
/ sp. nov. / det. P. KMENT 2015 [p, red label]. The
<typeStatus id="54D7F4FDA963981C7A0244A1FE76FB57" box="[335,410,1177,1196]" pageId="21" pageNumber="612" type="holotype">holotype</typeStatus>
is pinned through scutellum, the detached right antennal segment IV sticking to the rest of the antenna.
<materialsCitation id="3B044002A963981C7A15448CFD88FB19" collectionCode="NMPC" pageId="21" pageNumber="612" specimenCount="1" typeStatus="paratype">
<typeStatus id="54D7F4FDA963981C7A15448CFE5CFB3E" box="[344,432,1204,1223]" pageId="21" pageNumber="612" type="paratype">PARATYPES</typeStatus>
: 1 J, Rogez / Madagascar / Mus.Praha [p] // COLLECTIO / NATIONAL MUSEUM / Praha, Czech Republic [p] // J [p] (
<collectionCode id="ED7DD29AA963981C795044F7FDB2FB19" box="[541,606,1231,1250]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="612">NMPC</collectionCode>
)
</materialsCitation>
; male genitalia dissected: the dissected genital capsule, one paramere and the proctiger are glued on a separate piece of card, the phallus is stored in a plastic microvial in glycerol, all being attached to the same pin with the specimen;
<materialsCitation id="3B044002A963981C79D9453CFD48FA9C" collectionCode="NMPC, BMNH" country="Madagascar" county="Genus" determinerName="W. E. China" location="Rogez" municipality="Brit. Mus." pageId="21" pageNumber="612" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D6A81D6A963981C79D9453CFD59FAEC" box="[660,693,1278,1303]" pageId="21" pageNumber="612" type="female">1 ♀</specimenCount>
,
<location id="8EB31C84A963981C798E453CFD11FAEC" LSID="urn:lsid:plazi:treatment:03C5FB49A96398157A834424FF10FE48:8EB31C84A963981C798E453CFD11FAEC" box="[707,765,1284,1303]" country="Madagascar" county="Genus" municipality="Brit. Mus." name="Rogez" pageId="21" pageNumber="612">Rogez</location>
/
<location id="8EB31C84A963981C785D453CFC9BFAEC" LSID="urn:lsid:plazi:treatment:03C5FB49A96398157A834424FF10FE48:8EB31C84A963981C785D453CFC9BFAEC" box="[784,887,1284,1303]" country="Madagascar" county="Genus" municipality="Brit. Mus." name="Madagascar" pageId="21" pageNumber="612">Madagascar</location>
/
<location id="8EB31C84A963981C78C4453CFC08FAEC" LSID="urn:lsid:plazi:treatment:03C5FB49A96398157A834424FF10FE48:8EB31C84A963981C78C4453CFC08FAEC" box="[905,996,1284,1303]" country="Madagascar" county="Genus" municipality="Brit. Mus." name="Mus. Praha" pageId="21" pageNumber="612">Mus.Praha</location>
[p] // CO- LLECTIO / NATIONAL MUSEUM / Praha, Czech Republic [p] //
<specimenCount id="9D6A81D6A963981C79FC4520FD2DFACA" box="[689,705,1304,1329]" pageId="21" pageNumber="612" type="female"></specimenCount>
[p] (
<collectionCode id="ED7DD29AA963981C79BD4527FCDDFAC9" box="[752,817,1311,1330]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="612">NMPC</collectionCode>
). 1 J, [M]aroantsetra [hw, first letter illegible] // INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F37B0ACFA963981C79504501FD59FAB7" box="[541,693,1337,1356]" name="Madagascar" pageId="21" pageNumber="612">MADAGASCAR</collectingCountry>
[p] //
<collectingMunicipality id="6BB7D025A963981C79A14501FCA9FAB7" box="[748,837,1337,1356]" pageId="21" pageNumber="612">Brit. Mus.</collectingMunicipality>
/ 1965-338 [p] //
<collectingCounty id="62B232D3A963981C78924501FBFAFAB7" box="[991,1046,1337,1356]" pageId="21" pageNumber="612">Genus</collectingCounty>
near /
<taxonomicName id="4C6C31DCA963981C7B30456CFF20FA9C" authorityName="Horvath" authorityYear="1904" box="[125,204,1364,1383]" class="Insecta" family="Pentatomidae" genus="Triplatyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="620" phylum="Arthropoda" rank="genus">Triplatyx</taxonomicName>
[hw] /
<determinerName id="7E0FA69BA963981C7A41456CFE85FA9C" box="[268,361,1364,1383]" pageId="21" pageNumber="612">W.E.China</determinerName>
det. 196[p]4[hw] // J [p] (
<collectionCode id="ED7DD29AA963981C7914456CFD72FA9C" box="[601,670,1364,1383]" pageId="21" pageNumber="612">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B044002A963981C79E3456CFF2DFA67" collectionCode="MRAC" country="Madagascar" pageId="21" pageNumber="612" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
1 J
<specimenCount id="9D6A81D6A963981C7999456CFCE8FA9C" box="[724,772,1358,1383]" pageId="21" pageNumber="612" type="female">2 ♀♀</specimenCount>
, environs de / Tananarive [hw, pale blue label with black margins] // Madagascar [hw, pale blue label with black margins] // J or
<specimenCount id="9D6A81D6A963981C78D64550FC47FA7A" box="[923,939,1384,1409]" pageId="21" pageNumber="612" type="female"></specimenCount>
[p, respectively] (
<collectionCode id="ED7DD29AA963981C7B3A45B1FF57FA67" box="[119,187,1417,1436]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="21" pageNumber="612" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
; the male
<typeStatus id="54D7F4FDA963981C7A5345B1FE8BFA67" box="[286,359,1417,1436]" pageId="21" pageNumber="612" type="paratype">paratype</typeStatus>
has a detached but not dissected genital capsule glued on a separate piece of card attached to the same pin;
<materialsCitation id="3B044002A963981C7A1C459CFD66F9FC" collectionCode="MNHN" country="Madagascar" location="All" pageId="21" pageNumber="612" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
one of the female
<typeStatus id="54D7F4FDA963981C7ABD459CFDAEFA4C" box="[496,578,1444,1463]" pageId="21" pageNumber="612" type="paratype">paratypes</typeStatus>
has dissected abdomen glued to the specimen and internal genitalia stored in glycerol in a plastic microvial attached to the same pin.
<specimenCount id="9D6A81D6A963981C79A64587FCE7FA2A" box="[747,779,1464,1490]" pageId="21" pageNumber="612" type="female">1 ♀</specimenCount>
, environs de / Tananarive [hw, pale blue label with black margins] // Madagascar [hw, pale blue label with black margins] // MUSEUM PARIS [p] //
<specimenCount id="9D6A81D6A963981C7F4045EBFBF1FA17" box="[1037,1053,1491,1516]" pageId="21" pageNumber="612" type="female"></specimenCount>
[p] (
<collectionCode id="ED7DD29AA963981C7B3A45CCFF51F9FC" box="[119,189,1524,1543]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="612">MNHN</collectionCode>
).
<location id="8EB31C84A963981C7B8645CCFF0AF9FC" LSID="urn:lsid:plazi:treatment:03C5FB49A96398157A834424FF10FE48:8EB31C84A963981C7B8645CCFF0AF9FC" box="[203,230,1524,1543]" country="Madagascar" name="All" pageId="21" pageNumber="612">All</location>
<typeStatus id="54D7F4FDA963981C7BA145CCFED2F9FC" box="[236,318,1524,1543]" pageId="21" pageNumber="612" type="paratype">paratypes</typeStatus>
bear the following identification label
</materialsCitation>
:
<typeStatus id="54D7F4FDA963981C79D845CCFCFDF9FC" box="[661,785,1524,1543]" pageId="21" pageNumber="612" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="4C6C31DCA963981C786E45CCFCB6F9FC" authorityName="Kment" authorityYear="2015" box="[803,858,1524,1543]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="617" phylum="Arthropoda" rank="genus">
<emphasis id="B918964DA963981C786E45CCFCB6F9FC" box="[803,858,1524,1543]" italics="true" pageId="21" pageNumber="612">NENE</emphasis>
</taxonomicName>
/
<emphasis id="B918964DA963981C782645CCFC01F9FC" box="[875,1005,1524,1543]" italics="true" pageId="21" pageNumber="612">UNDULATUM</emphasis>
/ sp. nov. / det. P. KMENT 2015 [p, red label].
</paragraph>
</subSubSection>
<caption id="DF131AD7A960981F7BDA44ADFC3BFB03" ID-DOI="http://doi.org/10.5281/zenodo.5363141" ID-Zenodo-Dep="5363141" httpUri="https://zenodo.org/record/5363141/files/figure.png" pageId="22" pageNumber="613" startId="22.[151,188,1173,1192]" targetBox="[187,1088,228,1158]" targetPageId="22">
<paragraph id="8BD34A5FA960981F7BDA44ADFC3BFB03" blockId="22.[151,1127,1173,1272]" pageId="22" pageNumber="613">
Figs 4447.
<taxonomicName id="4C6C31DCA960981F7A4D44ADFE66FB53" authority="Kment, 2015" authorityName="Kment" authorityYear="2015" box="[256,394,1173,1192]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="613" phylum="Arthropoda" rank="species" species="elongatum" status="sp. nov.">
<emphasis id="B918964DA960981F7A4D44ADFE66FB53" box="[256,394,1173,1192]" italics="true" pageId="22" pageNumber="613">Nene elongatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B2B36A960981F7AC344ADFE3DFB53" box="[398,465,1173,1192]" pageId="22" pageNumber="613" rank="species">sp. nov.</taxonomicNameLabel>
4445 head (44 dorsal view, 45 lateral view). 46 humerus of pronotum, dorsal view. 18 meso- and metapleuron with external scent efferent system of the metathoracic scent gland (evaporatorium dotted).Abbreviations: at antenniferous tubercle, bc buccula, ev evaporatorium, lb1 labial segment I, lr labrum, mp mandibular plate, o ostiole, pe peritreme, sp spiracle. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C37619D4A96098157BDA450BFD72FEE8" lastPageId="28" lastPageNumber="619" pageId="22" pageNumber="613" type="description">
<paragraph id="8BD34A5FA96098107BDA450BFDC1FED4" blockId="22.[151,1127,1331,1578]" lastBlockId="25.[113,1090,216,431]" lastPageId="25" lastPageNumber="616" pageId="22" pageNumber="613">
<emphasis id="B918964DA960981F7BDA450BFDEDFAB1" bold="true" box="[151,513,1331,1354]" pageId="22" pageNumber="613">
Description of adult.
<emphasis id="B918964DA960981F7AC7450BFDEDFAB1" bold="true" box="[394,513,1331,1354]" italics="true" pageId="22" pageNumber="613">Coloration</emphasis>
</emphasis>
(
<figureCitation id="135756DAA960981F7942450BFD9EFAB1" box="[527,626,1331,1354]" captionStart="Figs 68" captionStartId="5.[113,150,1269,1288]" captionTargetBox="[113,1088,219,1242]" captionTargetId="figure-11@5.[113,1088,219,1242]" captionTargetPageId="5" captionText="Figs 68. Nene undulatum sp. nov., habitus of adult (J, paratype from Maroantsetra, body length 14.0 mm): 6 dorsal view, 7 ventral view, 8 lateral view." figureDoi="http://doi.org/10.5281/zenodo.5363125" httpUri="https://zenodo.org/record/5363125/files/figure.png" pageId="22" pageNumber="613">Figs 68</figureCitation>
). Entire body and legs brick red, dorsally with only few narrow black (on anterior margin of humeri, on inner margin of pronotal calli, in foveae in anterolateral angles of scutellum) and brown markings (anterolaterally on margins of connexival segments). Body ventrally and femora with numerous irregularly scattered black markings, most prominent on meso- and metapleuron; abdominal ventrite VII anteromedially with large black triangular spot pointed posteriad (
<figureCitation id="135756DAA960981F79E545EBFD0BFA11" box="[680,743,1491,1514]" captionStart="Figs 68" captionStartId="5.[113,150,1269,1288]" captionTargetBox="[113,1088,219,1242]" captionTargetId="figure-11@5.[113,1088,219,1242]" captionTargetPageId="5" captionText="Figs 68. Nene undulatum sp. nov., habitus of adult (J, paratype from Maroantsetra, body length 14.0 mm): 6 dorsal view, 7 ventral view, 8 lateral view." figureDoi="http://doi.org/10.5281/zenodo.5363125" httpUri="https://zenodo.org/record/5363125/files/figure.png" pageId="22" pageNumber="613">Fig. 7</figureCitation>
); femora with few larger black spots on dorsal surface, the most lateral spot projecting ventrally and forming narrow, undulated, more or less complete ring. Antennal segment I reddish with black spots medially or in entire basal half, remaining antennal segments black with reddish apices. Labium reddish, apical segment darkened. Spiracles of abdominal ventrites IIIVII black. Claws black apically. Membrane brown with dark brown veins.
</paragraph>
<caption id="DF131AD7A961981E7B3C4582FBC3F9CC" ID-DOI="http://doi.org/10.5281/zenodo.5363143" ID-Zenodo-Dep="5363143" httpUri="https://zenodo.org/record/5363143/files/figure.png" pageId="23" pageNumber="614" startId="23.[113,150,1466,1485]" targetBox="[151,1051,219,1438]" targetPageId="23">
<paragraph id="8BD34A5FA961981E7B3C4582FBC3F9CC" blockId="23.[113,1089,1466,1591]" pageId="23" pageNumber="614">
Figs 4855.
<taxonomicName id="4C6C31DCA961981E7B934582FE87FA36" authority="Kment, 2015" authorityName="Kment" authorityYear="2015" box="[222,363,1466,1485]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="614" phylum="Arthropoda" rank="species" species="elongatum" status="sp. nov.">
<emphasis id="B918964DA961981E7B934582FE87FA36" box="[222,363,1466,1485]" italics="true" pageId="23" pageNumber="614">Nene elongatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B2B36A961981E7A3F4582FE5AFA36" box="[370,438,1466,1485]" pageId="23" pageNumber="614" rank="species">sp. nov.</taxonomicNameLabel>
48 head, ventral view (magnification 37×). 49 meso- and metapleuron with external scent efferent system of the metathoracic scent gland (25×). 5051 ostiole and peritreme (50 60×, 51 230×). 5253 metathoracic spiracle (52 75×, 53 300×). 54 female external genitalia (30×). 55 detail of trichobothria and spiracle on ventrite VI (190×). Abbreviations: ev evaporatorium, o ostiole, pes peritremal surface, sp spiracle, tr trichobothrium. Scale bars: 0.1 mm 51, 53, 55; 0.5 mm 50, 52; 1 mm 48, 49, 54.
</paragraph>
</caption>
<caption id="DF131AD7A96E98117BDA45A7FCC2F9CC" ID-DOI="http://doi.org/10.5281/zenodo.5363145" ID-Zenodo-Dep="5363145" httpUri="https://zenodo.org/record/5363145/files/figure.png" pageId="24" pageNumber="615" startId="24.[151,188,1439,1458]" targetBox="[151,1126,219,1374]" targetPageId="24">
<paragraph id="8BD34A5FA96E98117BDA45A7FCC2F9CC" blockId="24.[151,1127,1439,1591]" pageId="24" pageNumber="615">
Figs 5662.
<taxonomicName id="4C6C31DCA96E98117A4F45A7FE62FA49" authority="Kment, 2015" authorityName="Kment" authorityYear="2015" box="[258,398,1439,1458]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="615" phylum="Arthropoda" rank="species" species="elongatum" status="sp. nov.">
<emphasis id="B918964DA96E98117A4F45A7FE62FA49" box="[258,398,1439,1458]" italics="true" pageId="24" pageNumber="615">Nene elongatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B2B36A96E98117AD945A7FE3BFA49" box="[404,471,1439,1458]" pageId="24" pageNumber="615" rank="species">sp. nov.</taxonomicNameLabel>
5659 genital capsule: 56 undissected, dorsal view (magnification 35×); 57 dissected, dorsal view (35×). 58 undissected, posterior view (42×); 59 dissected, lateral view (37×). 60 proctiger, dorsal view (100×). 61 paramere, extracted (85×). 62 paramere, in situ (95×). Abbreviations: dif dorsal rim infolding, dr dorsal ridge of ventral wall, lif lateral rim infolding, ml membranous lobe of lateral rim infolding, p paramere, pap posterior aperture, vif ventral rim infolding, vr ventral ridge of ventral wall, vwd ventral wall depression. Scale bars: 0.2 mm 6062; 0.5 mm 5659.
</paragraph>
</caption>
<paragraph id="8BD34A5FA96F98137BDD4100FEBBFED4" blockId="25.[113,1090,216,431]" lastBlockId="26.[151,1127,216,591]" lastPageId="26" lastPageNumber="617" pageId="25" pageNumber="616">
<emphasis id="B918964DA96F98107BDD4100FE4BFEB4" bold="true" box="[144,423,312,335]" italics="true" pageId="25" pageNumber="616">Integument and vestiture.</emphasis>
Body dorsally (
<figureCitation id="135756DAA96F981079034100FD60FEB4" box="[590,652,312,335]" captionStart="Figs 68" captionStartId="5.[113,150,1269,1288]" captionTargetBox="[113,1088,219,1242]" captionTargetId="figure-11@5.[113,1088,219,1242]" captionTargetPageId="5" captionText="Figs 68. Nene undulatum sp. nov., habitus of adult (J, paratype from Maroantsetra, body length 14.0 mm): 6 dorsal view, 7 ventral view, 8 lateral view." figureDoi="http://doi.org/10.5281/zenodo.5363125" httpUri="https://zenodo.org/record/5363125/files/figure.png" pageId="25" pageNumber="616">Fig. 6</figureCitation>
) with irregular, small but deep punctures, concolorous to black, leaving more or less wide and callose impunctate interspaces (most prominent on head and pronotum); endocorium posteriorly with punctures more shallow than elsewhere; punctures on connexivum shallow and mostly concolorous. Body ventrally (
<figureCitation id="135756DAA96C98137BD240E0FEF5FF14" box="[159,281,216,239]" captionStart="Figs 4855" captionStartId="23.[113,150,1466,1485]" captionTargetBox="[151,1051,219,1438]" captionTargetId="figure-11@23.[151,1052,219,1438]" captionTargetPageId="23" captionText="Figs 4855. Nene elongatum sp. nov. 48 head, ventral view (magnification 37×). 49 meso- and metapleuron with external scent efferent system of the metathoracic scent gland (25×). 5051 ostiole and peritreme (50 60×, 51 230×). 5253 metathoracic spiracle (52 75×, 53 300×). 54 female external genitalia (30×). 55 detail of trichobothria and spiracle on ventrite VI (190×). Abbreviations: ev evaporatorium, o ostiole, pes peritremal surface, sp spiracle, tr trichobothrium. Scale bars: 0.1 mm 51, 53, 55; 0.5 mm 50, 52; 1 mm 48, 49, 54." figureDoi="http://doi.org/10.5281/zenodo.5363143" httpUri="https://zenodo.org/record/5363143/files/figure.png" pageId="26" pageNumber="617">Figs 4849</figureCitation>
) with small (head, connexiva) to large (pleura, base of abdomen), concolorous to black, irregularly distributed punctures. Femora with few scattered large black punctures; tibiae impunctate.
</paragraph>
<caption id="DF131AD7A96F98107B3C45ECFE39F9CC" ID-DOI="http://doi.org/10.5281/zenodo.5363147" ID-Zenodo-Dep="5363147" httpUri="https://zenodo.org/record/5363147/files/figure.png" pageId="25" pageNumber="616" startId="25.[113,150,1492,1511]" targetBox="[123,1077,478,1477]" targetPageId="25">
<paragraph id="8BD34A5FA96F98107B3C45ECFE39F9CC" blockId="25.[113,1089,1492,1591]" pageId="25" pageNumber="616">
Figs 6368.
<taxonomicName id="4C6C31DCA96F98107B9145ECFE84FA1C" authority="Kment, 2015" authorityName="Kment" authorityYear="2015" box="[220,360,1492,1511]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="616" phylum="Arthropoda" rank="species" species="elongatum" status="sp. nov.">
<emphasis id="B918964DA96F98107B9145ECFE84FA1C" box="[220,360,1492,1511]" italics="true" pageId="25" pageNumber="616">Nene elongatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B2B36A96F98107A2045ECFE5CFA1C" box="[365,432,1492,1511]" pageId="25" pageNumber="616" rank="species">sp. nov.</taxonomicNameLabel>
63 paramere. 6468 phallus: 6466 uninflated (64 ventral view, 65 dorsal view, 66 lateral view); 6768 inflated (67 ventral view, 68 lateral view).Abbreviations: aed aedeagus (= vesica), bp basal plate, mpp median penial plates, phth phallotheca, res endophallic reservoir, vpp ventral process of phallotheca. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BD34A5FA96C98137BF84100FD8AFE54" blockId="26.[151,1127,216,591]" pageId="26" pageNumber="617">
Body glabrous, only tibiae (on inferior surface and apically), tarsi (ventrally), ventral rim infolding of genital capsule (
<figureCitation id="135756DAA96C98137A8E4160FDD2FE94" box="[451,574,344,367]" captionStart="Figs 5662" captionStartId="24.[151,188,1439,1458]" captionTargetBox="[151,1126,219,1374]" captionTargetId="figure-10@24.[151,1127,219,1375]" captionTargetPageId="24" captionText="Figs 5662. Nene elongatum sp. nov. 5659 genital capsule: 56 undissected, dorsal view (magnification 35×); 57 dissected, dorsal view (35×). 58 undissected, posterior view (42×); 59 dissected, lateral view (37×). 60 proctiger, dorsal view (100×). 61 paramere, extracted (85×). 62 paramere, in situ (95×). Abbreviations: dif dorsal rim infolding, dr dorsal ridge of ventral wall, lif lateral rim infolding, ml membranous lobe of lateral rim infolding, p paramere, pap posterior aperture, vif ventral rim infolding, vr ventral ridge of ventral wall, vwd ventral wall depression. Scale bars: 0.2 mm 6062; 0.5 mm 5659." figureDoi="http://doi.org/10.5281/zenodo.5363145" httpUri="https://zenodo.org/record/5363145/files/figure.png" pageId="26" pageNumber="617">Figs 5658</figureCitation>
), and external female genitalia (sparsely on valvifers VIII and IX,
<figureCitation id="135756DAA96C98137A524140FE87FE74" box="[287,363,376,399]" captionStart="Figs 4855" captionStartId="23.[113,150,1466,1485]" captionTargetBox="[151,1051,219,1438]" captionTargetId="figure-11@23.[151,1052,219,1438]" captionTargetPageId="23" captionText="Figs 4855. Nene elongatum sp. nov. 48 head, ventral view (magnification 37×). 49 meso- and metapleuron with external scent efferent system of the metathoracic scent gland (25×). 5051 ostiole and peritreme (50 60×, 51 230×). 5253 metathoracic spiracle (52 75×, 53 300×). 54 female external genitalia (30×). 55 detail of trichobothria and spiracle on ventrite VI (190×). Abbreviations: ev evaporatorium, o ostiole, pes peritremal surface, sp spiracle, tr trichobothrium. Scale bars: 0.1 mm 51, 53, 55; 0.5 mm 50, 52; 1 mm 48, 49, 54." figureDoi="http://doi.org/10.5281/zenodo.5363143" httpUri="https://zenodo.org/record/5363143/files/figure.png" pageId="26" pageNumber="617">Fig. 54</figureCitation>
) covered with short, semierect to erect pale setae. Antennal segments III and IV densely covered with microtrichiae.
</paragraph>
<paragraph id="8BD34A5FA96C98137BF84180FDBEFE34" blockId="26.[151,1127,216,591]" box="[181,594,440,463]" pageId="26" pageNumber="617">
<emphasis id="B918964DA96C98137BF84180FEC8FE34" bold="true" box="[181,292,440,463]" italics="true" pageId="26" pageNumber="617">Structure.</emphasis>
See the generic description.
</paragraph>
<paragraph id="8BD34A5FA96C98127BF841E0FBC2FEA8" blockId="26.[151,1127,216,591]" lastBlockId="27.[113,1089,220,627]" lastPageId="27" lastPageNumber="618" pageId="26" pageNumber="617">
<emphasis id="B918964DA96C98137BF841E0FEBAFE14" bold="true" box="[181,342,472,495]" italics="true" pageId="26" pageNumber="617">Measurements</emphasis>
(in mm). Males (n = 4, except antennal segments III (n = 3) and IV (n = 2); median (minimummaximum); measurements of
<typeStatus id="54D7F4FDA96C981379D541C0FD18FDF4" box="[664,756,504,527]" pageId="26" pageNumber="617" type="holotype">holotype</typeStatus>
presented in square brackets). Body length 13.85 (12.6214.31) [14.31]; head: length 3.36 (3.233.58) [3.43], width (including eyes) 3.92 (3.634.02) [4.02], interocular width 2.94 (2.702.99) [2.99]; lengths of antennal segments: I 0.72 (0.690.74) [0.74], IIa 0.98 (0.931.03) [0.98], IIb 1.06 (0.931.13) [1.13], III 1.32 (1.231.37) [1.32], IV 1.421.47 [1.42]; pronotum: length 3.56 (3.383.72) [3.68], width 10.98 (9.6111.76) [10.98]; scutellum: length 6.47 (6.476.86) [6.47], width 6.03 (5.496.27) [6.08]; abdomen: width (across segment III) 10.88 (9.9011.37) [10.88].
</paragraph>
<caption id="DF131AD7A96C98137BDA45A7FEA0F9CC" ID-DOI="http://doi.org/10.5281/zenodo.5319081" ID-Zenodo-Dep="5319081" httpUri="https://zenodo.org/record/5319081/files/figure.png" pageId="26" pageNumber="617" startId="26.[151,188,1439,1458]" targetBox="[153,1113,631,1412]" targetPageId="26">
<paragraph id="8BD34A5FA96C98137BDA45A7FEA0F9CC" blockId="26.[151,1127,1439,1591]" pageId="26" pageNumber="617">
Figs 6971.
<taxonomicName id="4C6C31DCA96C98137BB245A7FE65FA49" authority="Kment, 2015" authorityName="Kment" authorityYear="2015" box="[255,393,1439,1458]" class="Insecta" family="Pentatomidae" genus="Nene" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="617" phylum="Arthropoda" rank="species" species="elongatum" status="sp. nov.">
<emphasis id="B918964DA96C98137BB245A7FE65FA49" box="[255,393,1439,1458]" italics="true" pageId="26" pageNumber="617">Nene elongatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B2B36A96C98137AC045A7FE23FA49" box="[397,463,1439,1458]" pageId="26" pageNumber="617" rank="species">sp. nov.</taxonomicNameLabel>
6970 external female genitalia (69 intact, 70 dissected). 71 spermatheca. Triangulin not depicted. Abbreviations: ar apical receptacle of spermatheca, dd distal duct of spermatheca, df distal flange, dil spermathecal dilation, div distal invagination of spermathecal duct, ip intermediate part of spermatheca, lt8,9 laterotergite VIII and IX, pd proximal duct of spermatheca, pf proximal flange, sp spiracle, rs ring sclerite, so sclerite surrounding spermathecal opening, vf8,9 valvifer VIII and IX, X segment X. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BD34A5FA96D98127BDD4164FEE3FDC8" blockId="27.[113,1089,220,627]" pageId="27" pageNumber="618">Females (n = 4, except antennal segments III and IV (n = 3); median (minimummaximum)). Body length 13.77 (13.6915.08); head: length 3.72 (3.683.72), width (including eyes) 4.07 (4.024.17), interocular width 3.04 (2.993.19); lengths of antennal segments: I 0.78 (0.740.78), IIa 0.98 (0.881.03), IIb 1.01 (0.981.03), III 1.27 (1.271.37), IV 1.42 (1.371.47); pronotum: length 3.75 (3.683.97), width 11.42 (11.1811.57); scutellum: length 6.96 (6.867.06), width 6.47; abdomen: width (across segment III) 11.47 (11.2712.06).</paragraph>
<paragraph id="8BD34A5FA96D98157B3C4204FD72FEE8" blockId="27.[113,1089,220,627]" lastBlockId="28.[151,1127,220,435]" lastPageId="28" lastPageNumber="619" pageId="27" pageNumber="618">
<emphasis id="B918964DA96D98127B3C4204FE98FDA8" bold="true" box="[113,372,572,595]" pageId="27" pageNumber="618">Intraspecific variation.</emphasis>
The specimens examined appear uniform in structure (except slight differences in measurements) and differ mostly in details of the black markings on the body venter and the antennal segment I. Only in some specimens from the environs of
<collectingRegion id="49A884BDA96A9815789A40E4FB8BFF08" box="[983,1127,220,243]" country="Madagascar" name="Antananarivo" pageId="28" pageNumber="619">Antananarivo</collectingRegion>
the venter of abdomen is more or less bright red.
</paragraph>
</subSubSection>
<subSubSection id="C37619D4A96A98157BDA4124FE1AFE88" pageId="28" pageNumber="619" type="etymology">
<paragraph id="8BD34A5FA96A98157BDA4124FBC6FEA8" blockId="28.[151,1127,220,435]" pageId="28" pageNumber="619">
<emphasis id="B918964DA96A98157BDA4124FEF4FEC8" bold="true" box="[151,280,284,307]" pageId="28" pageNumber="619">Etymology.</emphasis>
The species name is the Latin adjective
<emphasis id="B918964DA96A981579934124FCABFEC8" box="[734,839,284,307]" italics="true" pageId="28" pageNumber="619">undulatus</emphasis>
(-
<emphasis id="B918964DA96A9815782F4124FC83FEC8" box="[866,879,284,307]" italics="true" pageId="28" pageNumber="619">a</emphasis>
, -
<emphasis id="B918964DA96A981578C44124FC46FEC8" box="[905,938,284,307]" italics="true" pageId="28" pageNumber="619">um</emphasis>
), meaning wavy, waved, undulate, referring to the characteristic undulated lateral margins of abdomen.
</paragraph>
<paragraph id="8BD34A5FA96A98157BDA4164FE1AFE88" blockId="28.[151,1127,220,435]" box="[151,502,348,371]" pageId="28" pageNumber="619">
<emphasis id="B918964DA96A98157BDA4164FE6CFE88" bold="true" box="[151,384,348,371]" pageId="28" pageNumber="619">Habitat and biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37619D4A96A98157BDA4144FF10FE48" pageId="28" pageNumber="619" type="distribution">
<paragraph id="8BD34A5FA96A98157BDA4144FF10FE48" blockId="28.[151,1127,220,435]" pageId="28" pageNumber="619">
<emphasis id="B918964DA96A98157BDA4144FEC6FE68" bold="true" box="[151,298,380,403]" pageId="28" pageNumber="619">Distribution.</emphasis>
Humid and subhumid forest areas of north-eastern and eastern
<collectingCountry id="F37B0ACFA96A981578AB4144FB8BFE68" box="[998,1127,380,403]" name="Madagascar" pageId="28" pageNumber="619">Madagascar</collectingCountry>
(Fig. 74).
</paragraph>
</subSubSection>
</treatment>
</document>