467 lines
53 KiB
XML
467 lines
53 KiB
XML
<document id="C2366447FA4B493E81C438640AB10360" ID-CLB-Dataset="5162" ID-DOI="10.5281/zenodo.5160153" ID-GBIF-Dataset="821719e5-29b6-4a57-b04a-55633671eb80" ID-ISSN="1942-1354" ID-Zenodo-Dep="5160153" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628098133970" checkinUser="felipe" docAuthor="Woodruff, Robert E. & Warren E. Steiner, Jr." docDate="2011" docId="03C5BA39FF956E45FF1FCDF3C1A20050" docLanguage="en" docName="InsectaMundi.0157.1-6.pdf" docOrigin="Insecta Mundi 2011 (157)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Phyllophaga navassa Woodruff and Steiner 2011, new species" docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="FFFCC241FF976E43FFDFCB4CC3500437" masterDocTitle="new species of Phyllophaga Harris from the island of Navassa in the Caribbean (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="1" updateTime="1699045240715" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="E510B4DCEFAD7A5C8EFB52D8EF7BD94C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5CA3352780EB3B690ED3EFEEB8A1A6BD">
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<mods:title id="4DDD1DD6C371EE2E7FB3ABD0A6C114BC">new species of Phyllophaga Harris from the island of Navassa in the Caribbean (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:name id="AE3565B0AE1CB6289A1B2B9345112B0B" type="personal">
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<mods:roleTerm id="241B054D9D56018A454EED3A2E41E0F8">Author</mods:roleTerm>
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<mods:namePart id="E75D2BA738828938BDA860CCBA081E6F">Woodruff, Robert E.</mods:namePart>
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<mods:name id="C7B4E95F0B8A66ACA79968C7FA264565" type="personal">
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<mods:roleTerm id="7236A3C11A4CB300717B5F34114F6E2A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="10ADC3521AA30DC2BD35A033CBF3356C">Warren E. Steiner, Jr.</mods:namePart>
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<mods:typeOfResource id="7B0A6FFA6ED04CCC0299B6925EE3584D">text</mods:typeOfResource>
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<mods:title id="3F3DD8B33451DF0C50BB185F2C4D5BDA">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="D56B91F274C17A0CED635BFFA23D70C1">
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<mods:date id="16BA53CC548103FE2EE7D8694E668DCB">2011</mods:date>
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<mods:number id="961CF1F7462D213A1DC821C7F7A5FD39">2011-03-11</mods:number>
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<mods:detail id="E23634012221BCC53C21DB3CF01C8351" type="volume">
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<mods:number id="A3635C891EA1F337C7644D8625BBC355">2011</mods:number>
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<mods:detail id="6ECDF9F3F5B8DB092570FE2817AF28FE" type="issue">
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<mods:number id="1904DEE15A15776DB6BF685F42043B9A">157</mods:number>
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<mods:classification id="2C64390AAB4032CED182EE0026A7BDCB">journal article</mods:classification>
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<mods:identifier id="E21AA2D1ED13131349EE462C00BF9877" type="CLB-Dataset">5162</mods:identifier>
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<mods:identifier id="B1857DFFD45805C8F1AAACFC5EA56367" type="DOI">10.5281/zenodo.5160153</mods:identifier>
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<mods:identifier id="4B647AC1D5DD5EB067641D387305857F" type="GBIF-Dataset">821719e5-29b6-4a57-b04a-55633671eb80</mods:identifier>
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<mods:identifier id="E2672D25A35372A71B1CE5C23CB97E8E" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03C5BA39FF956E45FF1FCDF3C1A20050" ID-DOI="http://doi.org/10.5281/zenodo.5164040" ID-GBIF-Taxon="183919681" ID-Zenodo-Dep="5164040" LSID="urn:lsid:plazi:treatment:03C5BA39FF956E45FF1FCDF3C1A20050" httpUri="http://treatment.plazi.org/id/03C5BA39FF956E45FF1FCDF3C1A20050" lastPageId="6" lastPageNumber="5" pageId="2" pageNumber="1">
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<subSubSection id="C37658A4FF956E41FF1FCDF3C09602EF" box="[192,966,1727,1752]" pageId="2" pageNumber="1" type="nomenclature">
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<paragraph id="8BD30B2FFF956E41FF1FCDF3C09602EF" blockId="2.[192,966,1727,1752]" box="[192,966,1727,1752]" pageId="2" pageNumber="1">
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<heading id="D09BBC43FF956E41FF1FCDF3C09602EF" bold="true" box="[192,966,1727,1752]" fontSize="10" level="2" pageId="2" pageNumber="1" reason="3">
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<emphasis id="B918D73DFF956E41FF1FCDF3C09602EF" bold="true" box="[192,966,1727,1752]" pageId="2" pageNumber="1">
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<taxonomicName id="4C6C70ACFF956E41FF1FCDF3C04602EF" authority="Woodruff and Steiner" authorityName="Woodruff and Steiner" authorityYear="2011" box="[192,790,1727,1752]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="navassa" status="sp. nov.">
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<emphasis id="B918D73DFF956E41FF1FCDF3C2B602E0" bold="true" box="[192,486,1727,1752]" italics="true" pageId="2" pageNumber="1">Phyllophaga navassa</emphasis>
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Woodruff and Steiner
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22B6A46FF956E41FCFDCD8CC09602EF" box="[802,966,1728,1752]" pageId="2" pageNumber="1" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37658A4FF956E45FF1FCC4CC08E0030" lastPageId="6" lastPageNumber="5" pageId="2" pageNumber="1" type="materials_examined">
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<paragraph id="8BD30B2FFF956E40FF1FCC4CC214050C" blockId="2.[192,1440,1792,1880]" lastBlockId="3.[192,1440,195,315]" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1">
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<emphasis id="B918D73DFF956E41FF1FCC4CC2C2032F" bold="true" box="[192,402,1792,1816]" pageId="2" pageNumber="1">
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<typeStatus id="54D7B58DFF956E41FF1FCC4CC26F032F" box="[192,319,1792,1816]" pageId="2" pageNumber="1" type="holotype">Holotype</typeStatus>
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male.
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</emphasis>
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<collectingCountry id="F37B4BBFFF956E41FE44CC4DC10D032F" box="[411,605,1792,1816]" name="Navassa Island" pageId="2" pageNumber="1">Navassa Island</collectingCountry>
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[
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<collectingCountry id="F37B4BBFFF956E41FDB1CC4CC1F7032F" box="[622,679,1792,1816]" name="United States of America" pageId="2" pageNumber="1">U.S.</collectingCountry>
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], central forest area, 70M,
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<geoCoordinate id="EE586DE8FF956E41FC22CC4CC7D5032F" box="[1021,1157,1792,1816]" degrees="18" direction="north" minutes="24.08" orientation="latitude" pageId="2" pageNumber="1" precision="9" value="18.401333">18º24.08’N</geoCoordinate>
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,
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<geoCoordinate id="EE586DE8FF956E41FB4DCC4CC665032F" box="[1170,1333,1792,1816]" degrees="75" direction="west" minutes="00.694" orientation="longitude" pageId="2" pageNumber="1" precision="1" value="-75.011566">75º00.694' W</geoCoordinate>
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,
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<date id="FFD22DEFFF956E41FA9CCC4CC3AD030F" pageId="2" pageNumber="1" value="1998-07-28">28 July 1998</date>
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, collrs. W. E. Steiner, J. M. Swearingen,
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<emphasis id="B918D73DFF956E41FD05CC6DC043030F" box="[730,787,1824,1848]" italics="true" pageId="2" pageNumber="1">et al.</emphasis>
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/ At blacklight in gap in mixed forest (
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<emphasis id="B918D73DFF956E41FB0ECC6DC2F9036F" italics="true" pageId="2" pageNumber="1">
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<taxonomicName id="4C6C70ACFF956E41FB0ECC6DC645030F" box="[1233,1301,1825,1848]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="1" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
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,
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<taxonomicName id="4C6C70ACFF956E41FAC0CC6DC6CC030F" box="[1311,1436,1825,1848]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="1" phylum="Tracheophyta" rank="genus">Metopium</taxonomicName>
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,
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<taxonomicName id="4C6C70ACFF956E41FF1FCC0CC2F9036F" authority=", Thrinax" authorityName="Thrinax" box="[192,425,1856,1880]" class="Magnoliopsida" family="Polygonaceae" genus="Coccoloba" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="1" phylum="Tracheophyta" rank="genus">Coccoloba, Thrinax</taxonomicName>
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</emphasis>
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) on limestone. [
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<collectingCountry id="F37B4BBFFF956E41FDBACC0CC1CA036F" box="[613,666,1856,1880]" name="United States of America" pageId="2" pageNumber="1">U.S.</collectingCountry>
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National Museum of Natural History, Smithsonian Institution].
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<emphasis id="B918D73DFF946E40FF1FCB8FC28A04EC" bold="true" box="[192,474,195,219]" pageId="3" pageNumber="2">General description</emphasis>
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(
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<figureCitation id="135717AAFF946E40FE33CB8FC16804EC" box="[492,568,195,219]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 4</figureCitation>
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, habitus). Medium sized (L.
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<quantity id="4C94A6CAFF946E40FC78CB8FC0A704EC" box="[935,1015,195,219]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="2" unit="mm" value="15.0">15mm</quantity>
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, W.
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<quantity id="4C94A6CAFF946E40FBEDCB8FC72304EC" box="[1074,1139,195,219]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="2" unit="mm" value="8.0">8mm</quantity>
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), shiny, uniformly dark mahogany brown, convex, nearly glabrous dorsally, unremarkable for genus; genitalia simple, but distinctive, cylindrical, parameres fused ventrally at tip, aedeagus recessed, partly sclerotized, tip upturned (
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<figureCitation id="135717AAFF946E40FF16CA6FC265050C" box="[201,309,291,315]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 8-15</figureCitation>
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).
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF1FCA2FC1DC05EC" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF1FCA2FC258054C" bold="true" box="[192,264,355,379]" pageId="3" pageNumber="2">Head</emphasis>
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(
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<figureCitation id="135717AAFF946E40FEC9CA2FC226054C" box="[278,374,355,379]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 4, 5</figureCitation>
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). Clypeus barely emarginate, lateral angles smoothly rounded, medial indentation slight, margin carinate; clypeal punctures dense, with most coalescing, not setose. Head punctures similar to clypeus, with a few separated by a puncture diameter, and a typical impunctate band near base. Eyes large with setose canthus near center.
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF2FCAAFC271060C" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF2FCAAFC23605CC" bold="true" box="[240,358,483,507]" pageId="3" pageNumber="2">Antenna</emphasis>
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(Fig. 1,3). Nine segmented, 3 club segments nearly equal in length, antennomere 1 nearly equal to next 3 combined, 3rd and 4th more cylindrical and smaller than 2
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<superScript id="7C19A667FF946E40FB88C94DC7390638" attach="left" box="[1111,1129,513,527]" fontSize="6" pageId="3" pageNumber="2">nd</superScript>
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or 5
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<superScript id="7C19A667FF946E40FB7FC94DC7FF0638" attach="none" box="[1184,1199,513,527]" fontSize="6" pageId="3" pageNumber="2">th</superScript>
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; 6
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<superScript id="7C19A667FF946E40FB11C94DC78D0638" attach="left" box="[1230,1245,513,527]" fontSize="6" pageId="3" pageNumber="2">th</superScript>
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small and compressed.
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF2FC90FC75606AC" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF2FC90FC229066C" bold="true" box="[240,377,579,603]" pageId="3" pageNumber="2">Pronotum</emphasis>
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(Fig. 4,5). Convex, shiny, punctures sparse, shallow, non-setiferous; lateral margin slightly crenulate at origin of long, curved lateral golden setae; anterior margin complete, posterior margin barely indicated, fading medially, smooth with only few scattered punctures.
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF2FC9EFC70A068C" blockId="3.[192,1441,355,1563]" box="[240,1114,675,699]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF2FC9EFC2D4068C" bold="true" box="[240,388,675,699]" pageId="3" pageNumber="2">Scutellum.</emphasis>
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Triangular, slightly convex, punctate, posteriorly flattened.
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF2FC98FC178070C" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF2FC98FC21706EC" bold="true" box="[240,327,707,731]" pageId="3" pageNumber="2">Elytra</emphasis>
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(
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<figureCitation id="135717AAFF946E40FE8AC98FC2C906EC" box="[341,409,707,731]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 4</figureCitation>
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) Convex, flattened medially, somewhat rugulose horizontally, striae barely indicated, disc glabrous, lateral third and humerus with fine scattered setae; marginal golden setae longest below humerus, similar to pronotal setae. Sutural interval raised, broadest in posterior fifth; terminating in slight projection (not toothed).
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF2FC80FC12A07AC" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF2FC80FC227076C" bold="true" box="[240,375,835,859]" pageId="3" pageNumber="2">Pygidium</emphasis>
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(
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<figureCitation id="135717AAFF946E40FE5AC80FC29C076C" box="[389,460,835,859]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 6</figureCitation>
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). Broadly oval/triangular,evenly convex, shiny, glabrous, with few, shallow, nonsetigerous punctures; posterior lateral margin carinate (inaccurately termed “bead” in older literature) with elongate, golden setae medially.
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</paragraph>
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<paragraph id="8BD30B2FFF946E40FF2FC8EFC09007EC" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
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<emphasis id="B918D73DFF946E40FF2FC8EFC260078C" bold="true" box="[240,304,931,955]" pageId="3" pageNumber="2">Legs</emphasis>
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(
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<figureCitation id="135717AAFF946E40FEE1C8EFC2E7078C" box="[318,439,931,955]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 2-5, 7</figureCitation>
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). Relatively long, especially tarsi; all claws not cleft with sharp termination, middle tooth small, nearer base than tip (anterior right claw in
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<figureCitation id="135717AAFF946E40FCB5C88FC0E107EC" box="[874,945,963,987]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD30B2FFF946E40FF29C8AFC6CC000C" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
|
||
Anterior tibia tridentate (
|
||
<figureCitation id="135717AAFF946E40FDF0C8AFC12507CC" box="[559,629,995,1019]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 5</figureCitation>
|
||
), anterior tooth elongate, middle sharp, triangular, posterior tooth reduced; teeth spaced nearly equidistant; spur narrow, cylindrical, sharp, half length of 1
|
||
<superScript id="7C19A667FF946E40FAD4CF4DC6480038" attach="left" box="[1291,1304,1025,1039]" fontSize="6" pageId="3" pageNumber="2">st</superScript>
|
||
tarsal segment. Anterior tarsal segments cylindrical, nearly equal in diameter, segments 2-4 subequal in length.
|
||
</paragraph>
|
||
<paragraph id="8BD30B2FFF946E40FF2FCF0FC1DC00EC" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
|
||
Meso-and metatibia with longitudinal carina and incomplete transverse carina (
|
||
<figureCitation id="135717AAFF946E40FB6DCF0FC7A7006C" box="[1202,1271,1091,1115]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 7</figureCitation>
|
||
), teeth prominent with long, golden setae. Metatibial spurs (
|
||
<figureCitation id="135717AAFF946E40FCD5CF2FC000004C" box="[778,848,1123,1147]" captionStart="Figure 1-7" captionStartId="4.[192,277,1773,1795]" captionTargetBox="[192,1438,192,1754]" captionTargetId="figure-21@4.[192,1440,192,1754]" captionTargetPageId="4" captionText="Figure 1-7. Phyllophaga navassa, n.sp. 1) Right antenna. 2) Right metatibial apex. 3) Right protibial claw. 4) Habitus, adult male. 5) Left protibia, tarsus, clypeus (line = 5mm). 6) Pygidium (caudal view). 7) Right metatibia (line = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="3" pageNumber="2">Fig. 2</figureCitation>
|
||
) moveable, long (reaching near middle of tarsal segment 2), narrow, slightly curved, inner surface flattened, inner spur about 20% longer than outer spur; terminus not noticeably notched; fringe of spines 14-16. Metatarsal segments long, cylindrical, 2
|
||
<superScript id="7C19A667FF946E40FA51CFEDC6F00098" attach="none" box="[1422,1440,1185,1199]" fontSize="6" pageId="3" pageNumber="2">nd</superScript>
|
||
longer than 1
|
||
<superScript id="7C19A667FF946E40FEBBCF8DC22100F8" attach="none" box="[356,369,1217,1231]" fontSize="6" pageId="3" pageNumber="2">st</superScript>
|
||
; claws as in other legs.
|
||
</paragraph>
|
||
<paragraph id="8BD30B2FFF946E40FF2FCFAFC67201AC" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
|
||
<emphasis id="B918D73DFF946E40FF2FCFAFC2EE00CC" bold="true" box="[240,446,1251,1275]" pageId="3" pageNumber="2">Male Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="135717AAFF946E40FE10CFAFC11000CC" box="[463,576,1251,1275]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 8-15</figureCitation>
|
||
). Of a simplified form, and unremarkable aedeagus. Parameres symmetrical, lateral views (
|
||
<figureCitation id="135717AAFF946E40FE38CE4FC116012C" box="[487,582,1283,1307]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 8-9</figureCitation>
|
||
) nearly identical. Caudally (
|
||
<figureCitation id="135717AAFF946E40FC75CE4FC750012C" box="[938,1024,1283,1307]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 10</figureCitation>
|
||
) with large broadly oval opening, parameres fused ventrally; dorsally (
|
||
<figureCitation id="135717AAFF946E40FDAACE6FC196010C" box="[629,710,1315,1339]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 14</figureCitation>
|
||
) opening v-shaped posteriorly. Each paramere with smoothly rounded lateral carina (
|
||
<figureCitation id="135717AAFF946E40FE3BCE0FC130016C" box="[484,608,1347,1371]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 12-13</figureCitation>
|
||
) about half length; nearly truncate ventrally. Aedeagus cylindrical, partly sclerotized, longer than parameres when extended (
|
||
<figureCitation id="135717AAFF946E40FCA5CE2FC754014C" box="[890,1028,1379,1403]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 8, 9, 11</figureCitation>
|
||
). Cylinder twisted and open at tip, with distinctive upturned medial dorsal process, barely protruding in repose (
|
||
<figureCitation id="135717AAFF946E40FBB6CECFC64401AC" box="[1129,1300,1411,1435]" captionStart="Figure 8-15" captionStartId="5.[192,276,1438,1460]" captionTargetBox="[192,1440,192,1418]" captionTargetId="figure-215@5.[192,1440,192,1418]" captionTargetPageId="5" captionText="Figure 8-15. Phyllophaga navassa, n.sp., male genitalia. 8,9,11) Aedeagus extended. 10,12-15) Aedeagus in repose. 8) Left lateral. 9) Right lateral. 10) Caudal. 11) dorsal/caudal. 12) Right lateral. 13) Laterocaudal. 14) Dorsal. 15) Left lateral (note upturned aedeagal tip)." figureDoi="http://doi.org/10.5281/zenodo.5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="3" pageNumber="2">Fig. 12, 13, 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD30B2FFF946E40FF2FCEEFC2E4022C" blockId="3.[192,1441,355,1563]" pageId="3" pageNumber="2">
|
||
<emphasis id="B918D73DFF946E40FF2FCEEFC20D018C" bold="true" box="[240,349,1443,1467]" pageId="3" pageNumber="2">Female.</emphasis>
|
||
Unknown. Because all specimens are males and were collected at light and flight intercept traps, the unknown females may be flightless and subterranean (as are some Hispaniolan high altitude species, see
|
||
<bibRefCitation id="EFFD76DEFF946E40FE93CEAFC1E401CC" author="Woodruff, R. E. & M. W. Sanderson" box="[332,692,1507,1531]" pageId="3" pageNumber="2" pagination="1 - 154" refId="ref3484" refString="Woodruff, R. E., and M. W. Sanderson. 2004. A revision of the Phyllophaga of Hispaniola (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi 18 (1 - 4): 1 - 154; 727 fig." type="book chapter" year="2004">Woodruff and Sanderson 2004</bibRefCitation>
|
||
). On a small windswept island such adaptation would obviously have survival value.
|
||
</paragraph>
|
||
<paragraph id="8BD30B2FFF946E40FF1FCD0FC23402EC" blockId="3.[192,1441,1603,1883]" pageId="3" pageNumber="2">
|
||
<emphasis id="B918D73DFF946E40FF1FCD0FC2D6026C" bold="true" box="[192,390,1603,1627]" pageId="3" pageNumber="2">Relationships.</emphasis>
|
||
Because 48 species of
|
||
<taxonomicName id="4C6C70ACFF946E40FD50CD0FC079026C" authorityName="Harris" authorityYear="1826" box="[655,809,1603,1627]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918D73DFF946E40FD50CD0FC079026C" box="[655,809,1603,1627]" italics="true" pageId="3" pageNumber="2">Phyllophaga</emphasis>
|
||
</taxonomicName>
|
||
are known from the nearest land mass (Hispaniola), it would be significant to establish the relationships of this isolated species. The simplicity of both body habitus and male genitalia imply that it may be a relictual, primitive species. Its colonization of
|
||
<collectingCountry id="F37B4BBFFF946E40FAE7CDC8C25F028C" name="Navassa Island" pageId="3" pageNumber="2">Navassa Island</collectingCountry>
|
||
suggests an old, durable, survivor from ancestral stock. DNA investigations may elucidate its relationships.
|
||
</paragraph>
|
||
<paragraph id="8BD30B2FFF946E40FF2FCDA8C0F6036C" blockId="3.[192,1441,1603,1883]" pageId="3" pageNumber="2">
|
||
It does not appear closely related (morphologically) to any other Caribbean
|
||
<taxonomicName id="4C6C70ACFF946E40FBB4CDAFC65402CC" authorityName="Harris" authorityYear="1826" box="[1131,1284,1763,1787]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918D73DFF946E40FBB4CDAFC65402CC" box="[1131,1284,1763,1787]" italics="true" pageId="3" pageNumber="2">Phyllophaga</emphasis>
|
||
</taxonomicName>
|
||
known to us. It is most similar to
|
||
<taxonomicName id="4C6C70ACFF946E40FE6FCC48C1E7032C" authority="Blanchard" authorityName="Blanchard" box="[432,695,1795,1819]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="hogardi">
|
||
<emphasis id="B918D73DFF946E40FE6FCC48C17E032C" box="[432,558,1795,1819]" italics="true" pageId="3" pageNumber="2">P. hogardi</emphasis>
|
||
Blanchard
|
||
</taxonomicName>
|
||
, one of the most abundant, widespread (and economic) species on Hispaniola. It does not possess the distinctive spine-like projections on the elytra sutural tip, and it lacks the “brain-like” mass at the aedeagal tip of that species.
|
||
</paragraph>
|
||
<caption id="DF135BA7FF936E47FF1FCDA1C2070309" ID-DOI="http://doi.org/10.5281/zenodo.5160155" ID-Zenodo-Dep="5160155" httpUri="https://zenodo.org/record/5160155/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,277,1773,1795]" targetBox="[192,1438,192,1754]" targetPageId="4">
|
||
<paragraph id="8BD30B2FFF936E47FF1FCDA1C2070309" blockId="4.[192,1440,1773,1854]" pageId="4" pageNumber="3">
|
||
<emphasis id="B918D73DFF936E47FF1FCDA1C2170334" bold="true" box="[192,327,1773,1795]" pageId="4" pageNumber="3">Figure 1-7.</emphasis>
|
||
<taxonomicName id="4C6C70ACFF936E47FE91CDA2C16D0333" authority="Woodruff & Warren E. Steiner, 2011" authorityName="Woodruff & Warren E. Steiner" authorityYear="2011" box="[334,573,1774,1796]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="navassa" status="sp. nov.">
|
||
<emphasis id="B918D73DFF936E47FE91CDA2C16D0333" box="[334,573,1774,1796]" italics="true" pageId="4" pageNumber="3">Phyllophaga navassa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A22B6A46FF936E47FD94CDA3C1D10333" box="[587,641,1775,1796]" pageId="4" pageNumber="3" rank="species">n.sp.</taxonomicNameLabel>
|
||
<emphasis id="B918D73DFF936E47FD57CDA1C1CF0334" bold="true" box="[648,671,1773,1795]" pageId="4" pageNumber="3">1)</emphasis>
|
||
Right antenna.
|
||
<emphasis id="B918D73DFF936E47FC87CDA1C03F0334" bold="true" box="[856,879,1773,1795]" pageId="4" pageNumber="3">2)</emphasis>
|
||
Right metatibial apex.
|
||
<emphasis id="B918D73DFF936E47FBA4CDA1C7C20334" bold="true" box="[1147,1170,1773,1795]" pageId="4" pageNumber="3">3)</emphasis>
|
||
Right protibial claw.
|
||
<emphasis id="B918D73DFF936E47FA56CDA1C6CF0334" bold="true" box="[1417,1439,1773,1795]" pageId="4" pageNumber="3">4)</emphasis>
|
||
Habitus, adult male.
|
||
<emphasis id="B918D73DFF936E47FE74CC46C2910317" bold="true" box="[427,449,1802,1824]" pageId="4" pageNumber="3">5)</emphasis>
|
||
Left protibia, tarsus, clypeus (line = 5mm).
|
||
<emphasis id="B918D73DFF936E47FC7ACC46C0EB0317" bold="true" box="[933,955,1802,1824]" pageId="4" pageNumber="3">6)</emphasis>
|
||
Pygidium (caudal view).
|
||
<emphasis id="B918D73DFF936E47FB0CCC46C7B90317" bold="true" box="[1235,1257,1802,1824]" pageId="4" pageNumber="3">7)</emphasis>
|
||
Right metatibia (line = 5mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF135BA7FF926E46FF1FCED2C04401D9" ID-DOI="http://doi.org/10.5281/zenodo.5160167" ID-Zenodo-Dep="5160167" httpUri="https://zenodo.org/record/5160167/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,276,1438,1460]" targetBox="[192,1440,192,1418]" targetPageId="5">
|
||
<paragraph id="8BD30B2FFF926E46FF1FCED2C04401D9" blockId="5.[192,1440,1438,1518]" pageId="5" pageNumber="4">
|
||
<emphasis id="B918D73DFF926E46FF1FCED2C2060183" bold="true" box="[192,342,1438,1460]" pageId="5" pageNumber="4">Figure 8-15.</emphasis>
|
||
<taxonomicName id="4C6C70ACFF926E46FEBFCED3C1010183" authority="Woodruff & Warren E. Steiner, 2011" authorityName="Woodruff & Warren E. Steiner" authorityYear="2011" box="[352,593,1439,1460]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="navassa" status="sp. nov.">
|
||
<emphasis id="B918D73DFF926E46FEBFCED3C1010183" box="[352,593,1439,1460]" italics="true" pageId="5" pageNumber="4">Phyllophaga navassa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A22B6A46FF926E46FDBECED3C1C90183" box="[609,665,1439,1460]" pageId="5" pageNumber="4" rank="species">n.sp.</taxonomicNameLabel>
|
||
, male genitalia.
|
||
<emphasis id="B918D73DFF926E46FC84CED2C0FA0183" bold="true" box="[859,938,1438,1460]" pageId="5" pageNumber="4">8,9,11)</emphasis>
|
||
Aedeagus extended.
|
||
<emphasis id="B918D73DFF926E46FB41CED2C65A0183" bold="true" box="[1182,1290,1438,1460]" pageId="5" pageNumber="4">10,12-15)</emphasis>
|
||
Aedeagus in repose.
|
||
<emphasis id="B918D73DFF926E46FEC8CEF7C27D01E6" bold="true" box="[279,301,1467,1489]" pageId="5" pageNumber="4">8)</emphasis>
|
||
Left lateral.
|
||
<emphasis id="B918D73DFF926E46FE1BCEF7C28B01E6" bold="true" box="[452,475,1467,1489]" pageId="5" pageNumber="4">9)</emphasis>
|
||
Right lateral.
|
||
<emphasis id="B918D73DFF926E46FD5DCEF7C1F701E6" bold="true" box="[642,679,1467,1489]" pageId="5" pageNumber="4">10)</emphasis>
|
||
Caudal.
|
||
<emphasis id="B918D73DFF926E46FCD1CEF7C06201E6" bold="true" box="[782,818,1467,1489]" pageId="5" pageNumber="4">11)</emphasis>
|
||
dorsal/caudal.
|
||
<emphasis id="B918D73DFF926E46FC00CEF7C75401E6" bold="true" box="[991,1028,1467,1489]" pageId="5" pageNumber="4">12)</emphasis>
|
||
Right lateral.
|
||
<emphasis id="B918D73DFF926E46FB74CEF7C78001E6" bold="true" box="[1195,1232,1467,1489]" pageId="5" pageNumber="4">13)</emphasis>
|
||
Laterocaudal.
|
||
<emphasis id="B918D73DFF926E46FAA4CEF7C6F001E6" bold="true" box="[1403,1440,1467,1489]" pageId="5" pageNumber="4">14)</emphasis>
|
||
Dorsal.
|
||
<emphasis id="B918D73DFF926E46FEC7CE9BC26D01DA" bold="true" box="[280,317,1495,1517]" pageId="5" pageNumber="4">15)</emphasis>
|
||
Left lateral (note upturned aedeagal tip).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD30B2FFF926E45FF1FCD54C08E0030" blockId="5.[192,1440,1560,1904]" lastBlockId="6.[192,1441,879,1032]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
<emphasis id="B918D73DFF926E46FF1FCD54C2B10207" bold="true" box="[192,481,1560,1584]" pageId="5" pageNumber="4">Specimens examined</emphasis>
|
||
[57].
|
||
<typeStatus id="54D7B58DFF926E46FDFCCD54C1C20207" box="[547,658,1560,1584]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D6AC0A6FF926E46FD14CD54C0FF0207" box="[715,943,1560,1585]" pageId="5" pageNumber="4" type="male" typeStatus="paratype">56 paratype males</specimenCount>
|
||
. All from
|
||
<collectingCountry id="F37B4BBFFF926E46FBFDCD55C7B10207" box="[1058,1249,1560,1584]" name="Navassa Island" pageId="5" pageNumber="4">Navassa Island</collectingCountry>
|
||
, collected by W. E. Steiner, J. M. Swearingen,
|
||
<emphasis id="B918D73DFF926E46FD9ECD75C1290267" box="[577,633,1592,1616]" italics="true" pageId="5" pageNumber="4">et al</emphasis>
|
||
. (numbers of
|
||
<typeStatus id="54D7B58DFF926E46FCF4CD75C0F70266" box="[811,935,1593,1617]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
in brackets): Central forest area,
|
||
<specimenCount id="9D6AC0A6FF926E46FA84CD74C6CB0267" box="[1371,1435,1592,1616]" pageId="5" pageNumber="4" type="generic">
|
||
<quantity id="4C94A6CAFF926E46FA84CD74C6C80267" box="[1371,1432,1592,1616]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="4" unit="m" value="70.0">70m</quantity>
|
||
.
|
||
</specimenCount>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FF1FCD14C2160247" box="[192,326,1622,1648]" degrees="18" direction="north" minutes="24.08" orientation="latitude" pageId="5" pageNumber="4" precision="9" value="18.401333">
|
||
18
|
||
<superScript id="7C19A667FF926E46FF01CD1AC3B60253" attach="right" box="[222,230,1622,1636]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
24.08’N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FE8ECD14C28C0247" box="[337,476,1622,1648]" degrees="75" direction="west" minutes="00.69" orientation="longitude" pageId="5" pageNumber="4" precision="9" value="-75.0115">
|
||
75
|
||
<superScript id="7C19A667FF926E46FEB1CD1AC2260253" attach="none" box="[366,374,1622,1636]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
00.69’W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFD22DEFFF926E46FE39CD14C1D30247" box="[486,643,1624,1648]" pageId="5" pageNumber="4" value="1998-07-28">28 July 1998</date>
|
||
, at black light in gap of mixed forest (
|
||
<emphasis id="B918D73DFF926E46FB9ACD15C27402A7" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C6C70ACFF926E46FB9ACD15C7DB0247" box="[1093,1163,1625,1648]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FB4ACD15C6440247" box="[1173,1300,1625,1648]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Metopium</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FAC1CD14C27402A7" authority=", Thrinax" authorityName="Thrinax" class="Magnoliopsida" family="Polygonaceae" genus="Coccoloba" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Coccoloba, Thrinax</taxonomicName>
|
||
</emphasis>
|
||
) on limestone [3]; bluff of southwest rim,
|
||
<specimenCount id="9D6AC0A6FF926E46FCC7CD34C00802A7" box="[792,856,1656,1680]" pageId="5" pageNumber="4" type="generic">
|
||
<quantity id="4C94A6CAFF926E46FCC7CD34C00302A7" box="[792,851,1656,1680]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="4" unit="m" value="65.0">65m</quantity>
|
||
.
|
||
</specimenCount>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FCBDCD34C0B802A7" box="[866,1000,1654,1680]" degrees="18" direction="north" minutes="23.75" orientation="latitude" pageId="5" pageNumber="4" precision="9" value="18.395834">
|
||
18
|
||
<superScript id="7C19A667FF926E46FC5FCD3AC0D802B3" attach="none" box="[896,904,1654,1668]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
23.75’N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FC2DCD34C72D02A7" box="[1010,1149,1654,1680]" degrees="75" direction="west" minutes="00.94" orientation="longitude" pageId="5" pageNumber="4" precision="9" value="-75.01566">
|
||
75
|
||
<superScript id="7C19A667FF926E46FBCFCD3AC74802B3" attach="none" box="[1040,1048,1654,1668]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
00.94’W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFD22DEFFF926E46FB58CD34C61A02A7" box="[1159,1354,1656,1680]" pageId="5" pageNumber="4" value="1998-07-25" valueMax="1998-07-30" valueMin="1998-07-25">25-30 July 1998</date>
|
||
, flightintercept/yellow pans in Malaise trap, open mixed forest (
|
||
<emphasis id="B918D73DFF926E46FC5DCDD5C7890287" box="[898,1241,1688,1712]" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C6C70ACFF926E46FC5DCDD5C7520287" box="[898,1026,1689,1712]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Metopium</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FBD2CDD4C7DB0287" box="[1037,1163,1688,1712]" class="Magnoliopsida" family="Polygonaceae" genus="Coccoloba" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Coccoloba</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FB49CDD5C7890287" box="[1174,1241,1689,1712]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
</emphasis>
|
||
) at rim of upper terrace, limestone and red oolitic soil [1]; central forest area,
|
||
<specimenCount id="9D6AC0A6FF926E46FC6ACDF4C0A502E7" box="[949,1013,1720,1744]" pageId="5" pageNumber="4" type="generic">
|
||
<quantity id="4C94A6CAFF926E46FC6ACDF4C0A202E7" box="[949,1010,1720,1744]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="4" unit="m" value="70.0">70m</quantity>
|
||
.
|
||
</specimenCount>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FBDDCDF4C7D902E7" box="[1026,1161,1718,1744]" degrees="18" direction="north" minutes="23.99" orientation="latitude" pageId="5" pageNumber="4" precision="9" value="18.399834">
|
||
18
|
||
<superScript id="7C19A667FF926E46FBFFCDFAC77802F3" attach="none" box="[1056,1064,1718,1732]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
23.99’N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FB4ACDF4C67102E7" box="[1173,1313,1718,1744]" degrees="75" direction="west" minutes="00.67" orientation="longitude" pageId="5" pageNumber="4" precision="9" value="-75.01117">
|
||
75
|
||
<superScript id="7C19A667FF926E46FB6CCDFAC7EB02F3" attach="none" box="[1203,1211,1718,1732]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
00.67’W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFD22DEFFF926E46FAF2CDF4C23202C7" pageId="5" pageNumber="4" value="1998-07-26" valueMax="1998-08-04" valueMin="1998-07-26">26 July-4 August 1998</date>
|
||
, flight-intercept/yellow pans in Malaise trap in gap of mixed forest (
|
||
<emphasis id="B918D73DFF926E46FB18CD95C1040327" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C6C70ACFF926E46FB18CD95C65D02C7" box="[1223,1293,1753,1776]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FAC3CD95C6CC02C7" box="[1308,1436,1753,1776]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Metopium</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FF1FCDB4C26E0327" box="[192,318,1784,1808]" class="Magnoliopsida" family="Polygonaceae" genus="Coccoloba" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Coccoloba</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FE93CDB4C1040327" authority=", Thrinax" authorityName="Thrinax" box="[332,596,1784,1808]" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Sideroxylon, Thrinax</taxonomicName>
|
||
</emphasis>
|
||
) on limestone [3]; same except Malaise trap in gap of mixed forest (
|
||
<emphasis id="B918D73DFF926E46FF16CC55C06F0307" box="[201,831,1816,1840]" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C6C70ACFF926E46FF16CC55C25F0307" box="[201,271,1817,1840]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FEC2CC55C2CD0307" box="[285,413,1817,1840]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Metopium</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FE74CC54C1790307" box="[427,553,1816,1840]" class="Magnoliopsida" family="Polygonaceae" genus="Coccoloba" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Coccoloba</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FDE9CC54C06F0307" authority=", Thrinax" authorityName="Thrinax" box="[566,831,1816,1840]" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Sideroxylon, Thrinax</taxonomicName>
|
||
</emphasis>
|
||
) on limestone [1]; E. end of east savanna,
|
||
<specimenCount id="9D6AC0A6FF926E46FA84CC54C6CB0307" box="[1371,1435,1816,1840]" pageId="5" pageNumber="4" type="generic">
|
||
<quantity id="4C94A6CAFF926E46FA84CC54C6C70307" box="[1371,1431,1816,1840]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="4" unit="m" value="65.0">65m</quantity>
|
||
.
|
||
</specimenCount>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FF1FCC74C2180367" box="[192,328,1846,1872]" degrees="18" direction="north" minutes="23.75" orientation="latitude" pageId="5" pageNumber="4" precision="9" value="18.395834">
|
||
18
|
||
<superScript id="7C19A667FF926E46FF01CC7AC3B60373" attach="none" box="[222,230,1846,1860]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
23.75’N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FE8DCC74C28D0367" box="[338,477,1846,1872]" degrees="75" direction="west" minutes="00.52" orientation="longitude" pageId="5" pageNumber="4" precision="9" value="-75.00867">
|
||
75
|
||
<superScript id="7C19A667FF926E46FEB0CC7AC2270373" attach="none" box="[367,375,1846,1860]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
00.52’W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFD22DEFFF926E46FE37CC74C1C90367" box="[488,665,1848,1872]" pageId="5" pageNumber="4" value="1998-08-01">1 August 1998</date>
|
||
, at black light at edge of grass savanna and mixed forest (
|
||
<emphasis id="B918D73DFF926E46FA8ACC75C2EF0347" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C6C70ACFF926E46FA8ACC75C6CB0367" box="[1365,1435,1849,1872]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FF1FCC15C26D0347" box="[192,317,1881,1904]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Metopium</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF926E46FE98CC14C2EF0347" box="[327,447,1880,1904]" class="Magnoliopsida" family="Polygonaceae" genus="Coccoloba" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Coccoloba</taxonomicName>
|
||
</emphasis>
|
||
) on limestone and red oolitic soil [3]; near lighthouse,
|
||
<specimenCount id="9D6AC0A6FF926E46FBE2CC14C72B0347" box="[1085,1147,1880,1904]" pageId="5" pageNumber="4" type="generic">
|
||
<quantity id="4C94A6CAFF926E46FBE2CC14C7280347" box="[1085,1144,1880,1904]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="4" unit="m" value="80.0">80m</quantity>
|
||
.
|
||
</specimenCount>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FB5ACC14C65A0347" box="[1157,1290,1878,1904]" degrees="18" direction="north" minutes="23.82" orientation="latitude" pageId="5" pageNumber="4" precision="9" value="18.397">
|
||
18
|
||
<superScript id="7C19A667FF926E46FB7CCC1AC7FB0353" attach="right" box="[1187,1195,1878,1892]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
23.82’N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE586DE8FF926E46FACCCC14C6CB0347" box="[1299,1435,1878,1904]" degrees="75" direction="west" minutes="00.74" orientation="longitude" pageId="5" pageNumber="4" precision="9" value="-75.01234">
|
||
75
|
||
<superScript id="7C19A667FF926E46FAEFCC1AC6680353" attach="none" box="[1328,1336,1878,1892]" fontSize="6" pageId="5" pageNumber="4">o</superScript>
|
||
00.74’W
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFD22DEFFF916E45FF1FC83CC2FF07BF" box="[192,431,879,904]" pageId="6" pageNumber="5" value="1998-07-24" valueMax="1998-08-04" valueMin="1998-07-24">24 July-4 Aug. 1998</date>
|
||
, taken in Malaise Trap, edge of open weedy scrub and mixed forest (
|
||
<emphasis id="B918D73DFF916E45FB0EC83CC2730790" italics="true" pageId="6" pageNumber="5">
|
||
<taxonomicName id="4C6C70ACFF916E45FB0EC83CC64507B0" box="[1233,1301,880,903]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="5" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF916E45FAC0C83CC2730790" authority=", Thrinax" authorityName="Thrinax" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="5" phylum="Tracheophyta" rank="genus">Metopium, Thrinax</taxonomicName>
|
||
</emphasis>
|
||
) on limestone [2]; same except
|
||
<date id="FFD22DEFFF916E45FD4CC8C3C07D079F" box="[659,813,911,936]" pageId="6" pageNumber="5" value="1998-07-26">26 July 1998</date>
|
||
, at black light in open weedy scrub near mixed forest (
|
||
<emphasis id="B918D73DFF916E45FF16C8FCC15007F0" box="[201,512,943,967]" italics="true" pageId="6" pageNumber="5">
|
||
<taxonomicName id="4C6C70ACFF916E45FF16C8FCC25D07F0" box="[201,269,944,967]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="5" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6C70ACFF916E45FEC8C8FCC15007F0" authority=", Thrinax" authorityName="Thrinax" box="[279,512,943,967]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="5" phylum="Tracheophyta" rank="genus">Metopium, Thrinax</taxonomicName>
|
||
</emphasis>
|
||
) on limestone and red oolitic soil [1].
|
||
<typeStatus id="54D7B58DFF916E45FC6BC8FCC77F07FF" box="[948,1071,944,968]" pageId="6" pageNumber="5" type="paratype">Paratypes</typeStatus>
|
||
deposited in the
|
||
<collectingCountry id="F37B4BBFFF916E45FB29C8FCC66107FF" box="[1270,1329,944,968]" name="United States of America" pageId="6" pageNumber="5">U. S.</collectingCountry>
|
||
National Museum of Natural History,
|
||
<collectingRegion id="49A8C5CDFF916E45FDC0C883C1E707D0" box="[543,695,975,999]" country="United States of America" name="Washington" pageId="6" pageNumber="5">Washington</collectingRegion>
|
||
, D.C., the
|
||
<collectingRegion id="49A8C5CDFF916E45FCEDC883C08307DF" box="[818,979,975,1000]" country="United States of America" name="Florida" pageId="6" pageNumber="5">Florida State</collectingRegion>
|
||
Collection of Arthropods, Gainesville,
|
||
<collectingRegion id="49A8C5CDFF916E45FF1FC8A3C24D0030" box="[192,285,1007,1031]" country="United States of America" name="Florida" pageId="6" pageNumber="5">Florida</collectingRegion>
|
||
, and the
|
||
<collectingRegion id="49A8C5CDFF916E45FE56C8A3C2B00030" box="[393,480,1007,1031]" country="United States of America" name="Illinois" pageId="6" pageNumber="5">Illinois</collectingRegion>
|
||
Natural History Survey, Urbana,
|
||
<collectingRegion id="49A8C5CDFF916E45FC5FC8A3C08A0030" box="[896,986,1007,1031]" country="United States of America" name="Illinois" pageId="6" pageNumber="5">Illinois</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF135BA7FF916E45FF1FC852C03D0765" ID-DOI="http://doi.org/10.5281/zenodo.5160169" ID-Zenodo-Dep="5160169" httpUri="https://zenodo.org/record/5160169/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,276,798,820]" targetBox="[192,1440,193,779]" targetPageId="6">
|
||
<paragraph id="8BD30B2FFF916E45FF1FC852C03D0765" blockId="6.[192,1440,798,850]" pageId="6" pageNumber="5">
|
||
<emphasis id="B918D73DFF916E45FF1FC852C2320703" bold="true" box="[192,354,798,820]" pageId="6" pageNumber="5">Figure 16-17.</emphasis>
|
||
The Island of Navassa.
|
||
<emphasis id="B918D73DFF916E45FDABC852C1C90703" bold="true" box="[628,665,798,820]" pageId="6" pageNumber="5">16)</emphasis>
|
||
Outline map with major features (courtesy of Wikipedia).
|
||
<emphasis id="B918D73DFF916E45FAF1C852C6030703" bold="true" box="[1326,1363,798,820]" pageId="6" pageNumber="5">17)</emphasis>
|
||
Aerial view of the island from satellite imagery (courtesy of NASA).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37658A4FF916E45FF1FCF7CC1A20050" pageId="6" pageNumber="5" type="etymology">
|
||
<paragraph id="8BD30B2FFF916E45FF1FCF7CC1A20050" blockId="6.[192,1440,1071,1127]" pageId="6" pageNumber="5">
|
||
<emphasis id="B918D73DFF916E45FF1FCF7CC20B007F" bold="true" box="[192,347,1072,1096]" pageId="6" pageNumber="5">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="4C6C70ACFF916E45FDE2CF7CC1F00070" authorityName="Woodruff & Warren E. Steiner" authorityYear="2011" box="[573,672,1072,1095]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="navassa">
|
||
<emphasis id="B918D73DFF916E45FDE2CF7CC1F00070" box="[573,672,1072,1095]" italics="true" pageId="6" pageNumber="5">navassa</emphasis>
|
||
</taxonomicName>
|
||
is the same as this remote island, the only locality from which it is known. It is applied as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |