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<document id="0F7C58ECF61A68021F849EF51E089C01" ID-DOI="10.11646/zootaxa.3700.3.7" ID-GBIF-Dataset="371f30c1-ff9d-4fde-9429-fc9d284caf03" ID-ISSN="1175-5326" ID-Zenodo-Dep="249376" ID-ZooBank="E3FA2E4B-DDF4-4FDB-8CF1-FFBA215D40CA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460663374311" checkinUser="plazi" docAuthor="Massa, Bruno" docDate="2013" docId="03C187FAFFA0B347FF64D56458BFFDBA" docLanguage="en" docName="zt03700p475.pdf" docOrigin="Zootaxa 3700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmethis cisti Fabricius 1787" docType="treatment" docVersion="13" lastPageNumber="438" masterDocId="FFF8FF82FFA2B344FFF3D33D5D23FFDA" masterDocTitle="Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species" masterLastPageNumber="475" masterPageNumber="435" pageNumber="437" updateTime="1698347717602" updateUser="plazi">
<mods:mods id="813E46B4881C0D7BBE0703ED5B1CC235" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="77875E4849C2179D97995092858E924D">Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species</mods:title>
</mods:titleInfo>
<mods:name id="533531BA4B6723C3C4BD133F0D0F5F02" type="personal">
<mods:role id="836F70E31C142252A6F609E138263E75">
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</mods:role>
<mods:namePart id="45C7FBCF4D4C6C9A44BC5DBA256BAA9D">Massa, Bruno</mods:namePart>
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<mods:title id="1D0F0F768CDA2DE4809BB3C30D4730CF">Zootaxa</mods:title>
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<mods:part id="0F34BE8F4E1D59D21C352700427FF2EA">
<mods:date id="FC17264EB9DCAA28F3A674C4BC6C83B7">2013</mods:date>
<mods:detail id="CC6EEB31549C9C2089496762950F5915" type="volume">
<mods:number id="B1982307459A441C3D24182F897597EB">3700</mods:number>
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<mods:detail id="C97294D9342AB849E4897322338D620D" type="issue">
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<mods:classification id="FF6C90C11FA36A6A7906E39773B3046A">journal article</mods:classification>
<mods:identifier id="A9795C62D0C5BA3F62B6A9116B57FE87" type="DOI">10.11646/zootaxa.3700.3.7</mods:identifier>
<mods:identifier id="3306BABE5E0D895B0520D362353C15E3" type="GBIF-Dataset">371f30c1-ff9d-4fde-9429-fc9d284caf03</mods:identifier>
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<treatment id="03C187FAFFA0B347FF64D56458BFFDBA" ID-DOI="http://doi.org/10.5281/zenodo.5697448" ID-GBIF-Taxon="119574877" ID-Zenodo-Dep="5697448" LSID="urn:lsid:plazi:treatment:03C187FAFFA0B347FF64D56458BFFDBA" httpUri="http://treatment.plazi.org/id/03C187FAFFA0B347FF64D56458BFFDBA" lastPageId="3" lastPageNumber="438" pageId="2" pageNumber="437">
<subSubSection id="C3726567FFA0B346FF64D5645F28F9A9" box="[151,523,1625,1651]" pageId="2" pageNumber="437" type="nomenclature">
<paragraph id="8BD736ECFFA0B346FF64D5645F28F9A9" blockId="2.[151,523,1625,1651]" box="[151,523,1625,1651]" pageId="2" pageNumber="437">
<heading id="D09F8180FFA0B346FF64D5645F28F9A9" bold="true" box="[151,523,1625,1651]" fontSize="11" level="1" pageId="2" pageNumber="437" reason="1">
<taxonomicName id="4C684D6FFFA0B346FF64D5645F28F9A9" ID-CoL="7CF3Q" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[151,523,1625,1651]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
<emphasis id="B91CEAFEFFA0B346FF64D5645F28F9A9" bold="true" box="[151,523,1625,1651]" pageId="2" pageNumber="437">
<emphasis id="B91CEAFEFFA0B346FF64D5645C12F9A9" bold="true" box="[151,305,1625,1651]" italics="true" pageId="2" pageNumber="437">Tmethis cisti</emphasis>
(Fabricius, 1787)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3726567FFA0B347FF64D5A258BFFDBA" lastPageId="3" lastPageNumber="438" pageId="2" pageNumber="437" type="materials_examined">
<paragraph id="8BD736ECFFA0B346FF64D5A25F2EF802" blockId="2.[151,1436,1695,2008]" pageId="2" pageNumber="437">
<emphasis id="B91CEAFEFFA0B346FF64D5A25C59F962" bold="true" box="[151,378,1695,1720]" pageId="2" pageNumber="437">Material examined</emphasis>
.
<collectingCountry id="F37F767CFFA0B346FE79D59D5CD0F96D" box="[394,499,1696,1719]" name="Morocco" pageId="2" pageNumber="437">Morocco</collectingCountry>
:
<collectingCountry id="F37F767CFFA0B346FE0CD5A25F70F962" box="[511,595,1695,1720]" name="Spain" pageId="2" pageNumber="437">Melilla</collectingCountry>
,
<specimenCount id="9D6EFD65FFA0B346FDACD59D5FA7F962" box="[607,644,1696,1720]" pageId="2" pageNumber="437" type="generic">3♂</specimenCount>
, 8♀; El Guerruau,
<specimenCount id="9D6EFD65FFA0B346FCACD59D5EA7F962" box="[863,900,1696,1720]" pageId="2" pageNumber="437" type="generic">3♂</specimenCount>
, 8♀; Mulay Rechid (Ulad Settut),
<specimenCount id="9D6EFD65FFA0B346FAE3D59D5816F962" box="[1296,1333,1696,1720]" pageId="2" pageNumber="437" type="generic">3♂</specimenCount>
; Toco El Arba (
<collectingCountry id="F37F767CFFA0B346FF2ED5F95C10F907" box="[221,307,1732,1757]" name="Spain" pageId="2" pageNumber="437">Melilla</collectingCountry>
),
<specimenCount id="9D6EFD65FFA0B346FEB5D5F95C48F906" box="[326,363,1732,1756]" pageId="2" pageNumber="437" type="generic">3♂</specimenCount>
, 8♀; Hasi Berkan,
<specimenCount id="9D6EFD65FFA0B346FDCCD5F95F46F906" box="[575,613,1732,1756]" pageId="2" pageNumber="437" type="generic">3♂</specimenCount>
, 8♀ (MNCN);
<collectingCountry id="F37F767CFFA0B346FCFDD5F95E45F907" box="[782,870,1732,1757]" name="Algeria" pageId="2" pageNumber="437">Algeria</collectingCountry>
: Ain M’Lila (Costantine) 18.
<date id="FFD6102CFFA0B346FB54D5F959D5F906" box="[1191,1270,1732,1756]" pageId="2" pageNumber="437" value="2001-05-01">
V.01,
<specimenCount id="9D6EFD65FFA0B346FB18D5F959D5F906" box="[1259,1270,1732,1756]" pageId="2" pageNumber="437" type="generic">1</specimenCount>
</date>
♂, 1♀; Dumas 20.
<date id="FFD6102CFFA0B346FF4AD5D55C2CF8DA" box="[185,271,1768,1792]" pageId="2" pageNumber="437" value="1949-05-01">V.49, 1</date>
♀;
<collectingCountry id="F37F767CFFA0B346FECBD5DA5CACF8DA" box="[312,399,1767,1792]" name="Tunisia" pageId="2" pageNumber="437">Tunisia</collectingCountry>
: Tamerza 26.
<date id="FFD6102CFFA0B346FDDED5DA5F83F8DA" box="[557,672,1767,1792]" pageId="2" pageNumber="437" value="1873-05-06">
V.1873,
<specimenCount id="9D6EFD65FFA0B346FD62D5DA5F83F8DA" box="[657,672,1767,1792]" pageId="2" pageNumber="437" type="generic">6</specimenCount>
</date>
♂, 9♀; Ludien 23.
<date id="FFD6102CFFA0B346FC88D5D55EDBF8DA" box="[891,1016,1768,1792]" pageId="2" pageNumber="437" value="1873-04-01">IV.1873, 1</date>
♀; Hammam el Lif 2.
<date id="FFD6102CFFA0B346FB0ED5DA58A9F8DA" box="[1277,1418,1767,1792]" pageId="2" pageNumber="437" value="1875-07-05">VII.1875, 5</date>
♀ (MSNG);
<quantity id="4C909B09FFA0B346FEF8D4315C76F8FF" box="[267,341,1804,1829]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="437" unit="km" value="15.0">15 km</quantity>
N El Jem 15.
<date id="FFD6102CFFA0B346FE05D4315F6FF8FE" box="[502,588,1804,1829]" pageId="2" pageNumber="437" value="1991-05-02">V.91, 2</date>
♂♂;
<collectingCountry id="F37F767CFFA0B346FD75D4315FF1F8FF" box="[646,722,1804,1829]" name="Tunisia" pageId="2" pageNumber="437">Djerba</collectingCountry>
17.
<date id="FFD6102CFFA0B346FD08D4315E79F8FE" box="[763,858,1804,1828]" pageId="2" pageNumber="437" value="1992-04-03">
IV.92,
<specimenCount id="9D6EFD65FFA0B346FCBFD4315E79F8FE" box="[844,858,1804,1828]" pageId="2" pageNumber="437" type="generic">3</specimenCount>
</date>
♂, 2♀; Sousse
<date id="FFD6102CFFA0B346FBFFD4315947F8FE" box="[1036,1124,1804,1828]" pageId="2" pageNumber="437" value="1979-05-30">30.V.79</date>
, ♂; Gafsa 2.
<date id="FFD6102CFFA0B346FB04D431587BF8FE" box="[1271,1368,1804,1828]" pageId="2" pageNumber="437" value="1979-06-01">VI.79, 1</date>
♀; La Skhira 1.
<date id="FFD6102CFFA0B346FF05D40D5C79F892" box="[246,346,1840,1864]" pageId="2" pageNumber="437" value="1979-06-01">VI.79, 1</date>
♀;
<quantity id="4C909B09FFA0B346FE8CD4125CE4F892" box="[383,455,1839,1864]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="437" unit="km" value="15.0">15 km</quantity>
N Gabes 16.
<date id="FFD6102CFFA0B346FDAAD4125F89F892" box="[601,682,1839,1864]" pageId="2" pageNumber="437" value="1991-05-01">
V.91,
<specimenCount id="9D6EFD65FFA0B346FD6DD40D5F89F892" box="[670,682,1840,1864]" pageId="2" pageNumber="437" type="generic">1</specimenCount>
</date>
♂; Mabeth 16.
<date id="FFD6102CFFA0B346FCBCD4125E82F892" box="[847,929,1839,1864]" pageId="2" pageNumber="437" value="1991-05-01">V.91, 1</date>
♀; Biserte, 1♀;
<quantity id="4C909B09FFA0B346FBA4D40D59BCF892" box="[1111,1183,1839,1864]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="437" unit="km" value="30.0">30 km</quantity>
S Gabes 16.
<date id="FFD6102CFFA0B346FADFD41258A2F892" box="[1324,1409,1839,1864]" pageId="2" pageNumber="437" value="1991-05-02">
V.91,
<specimenCount id="9D6EFD65FFA0B346FA82D40D58A2F892" box="[1393,1409,1840,1864]" pageId="2" pageNumber="437" type="generic">2</specimenCount>
</date>
♂; Korba 3.
<date id="FFD6102CFFA0B346FF04D4695C4BF8B6" box="[247,360,1876,1900]" pageId="2" pageNumber="437" value="1993-06-11">
VI.93,
<specimenCount id="9D6EFD65FFA0B346FEBFD4695C4BF8B6" box="[332,360,1876,1900]" pageId="2" pageNumber="437" type="generic">11</specimenCount>
</date>
♂, 5♀; Kerkenna Is. 31.
<date id="FFD6102CFFA0B346FD8AD4695FEDF8B6" box="[633,718,1876,1900]" pageId="2" pageNumber="437" value="1979-05-08">
V.79,
<specimenCount id="9D6EFD65FFA0B346FD33D4695FEDF8B6" box="[704,718,1876,1900]" pageId="2" pageNumber="437" type="generic">8</specimenCount>
</date>
♂, 6♀; Soliman 8.
<date id="FFD6102CFFA0B346FC6CD4695928F8B6" box="[927,1035,1876,1901]" pageId="2" pageNumber="437" value="1976-07-03">
VII.76,
<specimenCount id="9D6EFD65FFA0B346FC0ED4695928F8B6" box="[1021,1035,1876,1900]" pageId="2" pageNumber="437" type="generic">3</specimenCount>
</date>
♂; Sidi Mansoun 28.
<date id="FFD6102CFFA0B346FB09D4695879F8B6" box="[1274,1370,1876,1901]" pageId="2" pageNumber="437" value="1976-06-01">VI.76, 1</date>
♀; 16 Km N Gabés 5.
<date id="FFD6102CFFA0B346FEB3D44A5C89F84A" box="[320,426,1911,1936]" pageId="2" pageNumber="437" value="1976-07-03">
VII.76,
<specimenCount id="9D6EFD65FFA0B346FE6FD4455C89F84A" box="[412,426,1912,1936]" pageId="2" pageNumber="437" type="generic">3</specimenCount>
</date>
♂, 2♀; Sousse 7.
<date id="FFD6102CFFA0B346FD97D44A5FEEF84A" box="[612,717,1911,1936]" pageId="2" pageNumber="437" value="1976-07-01">VII.76, 1</date>
♀; 30 Km S Ben Gardhane 3.
<date id="FFD6102CFFA0B346FBE1D44A59A3F84A" box="[1042,1152,1911,1936]" pageId="2" pageNumber="437" value="1976-07-02">VII.76, 2</date>
♀ (MSNT);
<collectingCountry id="F37F767CFFA0B346FAF5D44A586FF84A" box="[1286,1356,1911,1936]" name="Libya" pageId="2" pageNumber="437">Libya</collectingCountry>
: Leptis Magna 18.
<date id="FFD6102CFFA0B346FEFFD4A15C4FF86E" box="[268,364,1948,1973]" pageId="2" pageNumber="437" value="2005-04-02">
IV.05,
<specimenCount id="9D6EFD65FFA0B346FEAFD4A15C4FF86E" box="[348,364,1948,1972]" pageId="2" pageNumber="437" type="generic">2</specimenCount>
</date>
♂; Tagiura 14.
<date id="FFD6102CFFA0B346FDFFD4A15F49F86E" box="[524,618,1948,1972]" pageId="2" pageNumber="437" value="1998-04-01">IV.98, 1</date>
♀ (BMCP); U. Caàm, Mizda, Tinaiat ben Galan,
<specimenCount id="9D6EFD65FFA0B346FB6FD4A159E2F86E" box="[1180,1217,1948,1972]" pageId="2" pageNumber="437" type="generic">4♂</specimenCount>
, 5♀ (MSNM); Ain Zara,
<specimenCount id="9D6EFD65FFA0B346FF2BD4FD5DDFF802" box="[216,252,1984,2008]" pageId="2" pageNumber="437" type="generic">1♂</specimenCount>
, 1♀; Homs,
<specimenCount id="9D6EFD65FFA0B346FE7FD4FD5C8DF802" box="[396,430,1984,2008]" pageId="2" pageNumber="437" type="generic">2♂</specimenCount>
(MZR).
</paragraph>
<paragraph id="8BD736ECFFA1B347FF34D3A45E4AFDE6" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="438">
<taxonomicName id="4C684D6FFFA1B347FF34D3A45C30FF6A" box="[199,275,153,176]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="438" phylum="Arthropoda" rank="species" species="cisti">
<emphasis id="B91CEAFEFFA1B347FF34D3A45C30FF6A" box="[199,275,153,176]" italics="true" pageId="3" pageNumber="438">T. cisti</emphasis>
</taxonomicName>
covers North Africa, from
<collectingCountry id="F37F767CFFA1B347FDB8D3A55F93FF75" box="[587,688,152,175]" name="Morocco" pageId="3" pageNumber="438">Morocco</collectingCountry>
to
<collectingCountry id="F37F767CFFA1B347FD24D3AA5E3BFF6A" box="[727,792,151,176]" name="Libya" pageId="3" pageNumber="438">Libya</collectingCountry>
(Tripolitania and Cyrenaica). Uvarov (1943) established that the
<typeStatus id="54D3884EFFA1B347FF00D3805C00FF0F" box="[243,291,189,213]" pageId="3" pageNumber="438">type</typeStatus>
locality was Northern
<collectingCountry id="F37F767CFFA1B347FDD5D3815F5EFF0F" box="[550,637,188,213]" name="Tunisia" pageId="3" pageNumber="438">Tunisia</collectingCountry>
. It is well characterized by a high pronotum keel, with deep transverse sulcus (mainly in males) (
<figureCitation id="13532A69FFA1B347FE4CD3E25F6AFF22" box="[447,585,223,248]" captionStart="FIGURES 32 – 39" captionStartId="28.[151,264,1715,1737]" captionTargetBox="[300,1286,184,1690]" captionTargetId="figure@28.[300,1287,183,1693]" captionTargetPageId="28" captionText="FIGURES 32 – 39. Lateral view of pronotum of some taxa described in the genus Tmethis, here revised." httpUri="https://zenodo.org/record/249379/files/figure.png" pageId="3" pageNumber="438">Figs. 32–36</figureCitation>
), male epiphallus with apical lobes large and broadly separated (
<figureCitation id="13532A69FFA1B347FADFD3E258A8FF22" box="[1324,1419,223,248]" captionStart="FIGURES 54 – 59" captionStartId="30.[151,264,1827,1849]" captionTargetBox="[290,1301,181,1799]" captionTargetId="figure@30.[286,1301,171,1806]" captionTargetPageId="30" captionText="FIGURES 54 – 59. Some of the taxa described in the genus Tmethis, view from above to show the pattern of wing colour pattern and the ratio between the length of wings and the body. Figs. 58 a and 58 b show the pattern of pronotum and wings of male and female of T. harterti n. status. Figs. 54 a and 55 a show the apical lobes, large and broadly separated, of epiphallus of T. cisti, while Figs. 59 b and 59 c show the relatively small and narrowly separated lobes of epiphallus of T. pulchripennis (after Uvarov 1943). Figs 54 b and 59 a show the differences in the spurs of hind tibiae, long and slender, with the first tarsal joint long, and its second pulvillus longer than the first or third in T. cisti; shorter and stouter, with first tarsal joint short, its second pulvillus not longer than the first or the third in T. pulchripennis (after Uvarov 1943)." httpUri="https://zenodo.org/record/249381/files/figure.png" pageId="3" pageNumber="438">Figs. 54</figureCitation>
a, 55a), spurs of hind tibiae long and slender (
<figureCitation id="13532A69FFA1B347FD8DD2395FF1FEC7" box="[638,722,260,285]" captionStart="FIGURES 54 – 59" captionStartId="30.[151,264,1827,1849]" captionTargetBox="[290,1301,181,1799]" captionTargetId="figure@30.[286,1301,171,1806]" captionTargetPageId="30" captionText="FIGURES 54 – 59. Some of the taxa described in the genus Tmethis, view from above to show the pattern of wing colour pattern and the ratio between the length of wings and the body. Figs. 58 a and 58 b show the pattern of pronotum and wings of male and female of T. harterti n. status. Figs. 54 a and 55 a show the apical lobes, large and broadly separated, of epiphallus of T. cisti, while Figs. 59 b and 59 c show the relatively small and narrowly separated lobes of epiphallus of T. pulchripennis (after Uvarov 1943). Figs 54 b and 59 a show the differences in the spurs of hind tibiae, long and slender, with the first tarsal joint long, and its second pulvillus longer than the first or third in T. cisti; shorter and stouter, with first tarsal joint short, its second pulvillus not longer than the first or the third in T. pulchripennis (after Uvarov 1943)." httpUri="https://zenodo.org/record/249381/files/figure.png" pageId="3" pageNumber="438">Fig. 54</figureCitation>
b). Tegmina are long and generally exceed the hind knees (
<figureCitation id="13532A69FFA1B347FA90D2395DFCFE9A" captionStart="FIGURES 40 – 53" captionStartId="29.[151,264,2001,2023]" captionTargetBox="[220,1366,182,1957]" captionTargetId="figure@29.[220,1366,165,1979]" captionTargetPageId="29" captionText="FIGURES 40 – 53. Lateral view of males and females of some taxa described in the genus Tmethis, here revised." httpUri="https://zenodo.org/record/249380/files/figure.png" pageId="3" pageNumber="438">Figs. 42–43</figureCitation>
,
<figureCitation id="13532A69FFA1B347FF1ED21A5C2FFE9A" box="[237,268,295,320]" captionStart="FIGURES 54 – 59" captionStartId="30.[151,264,1827,1849]" captionTargetBox="[290,1301,181,1799]" captionTargetId="figure@30.[286,1301,171,1806]" captionTargetPageId="30" captionText="FIGURES 54 – 59. Some of the taxa described in the genus Tmethis, view from above to show the pattern of wing colour pattern and the ratio between the length of wings and the body. Figs. 58 a and 58 b show the pattern of pronotum and wings of male and female of T. harterti n. status. Figs. 54 a and 55 a show the apical lobes, large and broadly separated, of epiphallus of T. cisti, while Figs. 59 b and 59 c show the relatively small and narrowly separated lobes of epiphallus of T. pulchripennis (after Uvarov 1943). Figs 54 b and 59 a show the differences in the spurs of hind tibiae, long and slender, with the first tarsal joint long, and its second pulvillus longer than the first or third in T. cisti; shorter and stouter, with first tarsal joint short, its second pulvillus not longer than the first or the third in T. pulchripennis (after Uvarov 1943)." httpUri="https://zenodo.org/record/249381/files/figure.png" pageId="3" pageNumber="438">55</figureCitation>
), hind wings are red inside with a central brown spot, which reaches the base of the red area, but this colour is very variable; for example in
<collectingCountry id="F37F767CFFA1B347FDA8D2715F8DFEBF" box="[603,686,332,357]" name="Tunisia" pageId="3" pageNumber="438">Tunisia</collectingCountry>
yellow (
<collectingCountry id="F37F767CFFA1B347FCE7D2715E41FEBF" box="[788,866,332,357]" name="Tunisia" pageId="3" pageNumber="438">Djerba</collectingCountry>
Is., Sousse), purple (Gafsa, Mareth, Gabes), blue (Skhira, Kerkenna Is.) and red tibiae (Korba, Biserte) have been found; in
<collectingCountry id="F37F767CFFA1B347FBF5D2525964FE52" box="[1030,1095,367,392]" name="Libya" pageId="3" pageNumber="438">Libya</collectingCountry>
bluish with red apex (Leptis Magna, East of Sirte), blue (East of Sirte, Agedabia) and vinaceous tibiae with red apex (Tagiura) have been detected; in
<collectingCountry id="F37F767CFFA1B347FEDFD28A5CA3FE0A" box="[300,384,439,464]" name="Algeria" pageId="3" pageNumber="438">Algeria</collectingCountry>
red (El Kantara, Tebessa, Constantine) and yellow tibiae (Dumas) have been recorded. Concerning the length of tegmina, specimens with abbreviated wings exist in different localities (e.g., Kerkenna Is., Korba); generally, when tegmina are shorter than abdomen, hind wings are shorter than tegmina and show a simplified colour pattern (less coloured, with faint brown spots).
</paragraph>
<paragraph id="8BD736ECFFA1B347FF34D17558BFFDBA" blockId="3.[151,1437,151,608]" box="[199,1436,583,608]" pageId="3" pageNumber="438">
In the following paragraphs I will examine the different populations listed as subspecies by Eades
<emphasis id="B91CEAFEFFA1B347FAFFD1745867FDBA" box="[1292,1348,583,608]" italics="true" pageId="3" pageNumber="438">et al.</emphasis>
(2013).
</paragraph>
</subSubSection>
</treatment>
</document>
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