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<document id="3CED36B496DAC738FE9D1F10EF767C7B" ID-CLB-Dataset="27671" ID-DOI="10.11646/zootaxa.4545.4.7" ID-GBIF-Dataset="6e699de7-3402-497b-ac7d-cf031d53b703" ID-ISSN="1175-5326" ID-Zenodo-Dep="2618341" ID-ZooBank="BBD8A9FD-BC9B-42D2-AD7D-CEB71ADF17DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553929527618" checkinUser="plazi" docAuthor="Ratsoavina, Fanomezana Mihaja, Raselimanana, Achille P., Scherz, Mark D., Rakotoarison, Andolalao, Razafindraibe, Jary H., Glaw, Frank &amp; Vences, Miguel" docDate="2019" docId="03C187C2FFAA9541FF22FF72FE784707" docLanguage="en" docName="zootaxa.4545.2.7.pdf" docOrigin="Zootaxa 4545 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Uroplatus finaritra Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences 2019, sp. nov." docType="treatment" docUuid="E6A1407B-7C9E-46F6-8FEC-D120A111A344" docUuidSource="ZooBank" docVersion="10" lastPageNumber="572" masterDocId="FFF8FFBAFFA99548FFB5FFE5FC02443C" masterDocTitle="Finaritra! A splendid new leaf-tailed gecko (Uroplatus) species from Marojejy National Park in north-eastern Madagascar" masterLastPageNumber="577" masterPageNumber="563" pageNumber="566" updateTime="1698715674977" updateUser="ExternalLinkService">
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<mods:title id="5E01AC6F1B179FB095FCE5E88DB7DB6C">Finaritra! A splendid new leaf-tailed gecko (Uroplatus) species from Marojejy National Park in north-eastern Madagascar</mods:title>
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<taxonomicName id="4C684D57FFAA954BFF22FF72FD7C448D" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[151,382,151,177]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAA954BFF22FF72FD7C448D" bold="true" box="[151,382,151,177]" italics="true" pageId="3" pageNumber="566">Uroplatus finaritra</emphasis>
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<figureCitation id="13532A51FFAA954BFF22FF5FFD0344EF" box="[151,257,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="5.[151,250,1917,1939]" captionStartId-1="6.[151,250,1623,1645]" captionStartId-2="7.[151,250,1744,1766]" captionStartId-3="11.[151,250,1965,1987]" captionStartId-4="12.[151,250,1534,1556]" captionTargetBox-0="[228,1370,195,1901]" captionTargetBox-1="[154,1433,195,1601]" captionTargetBox-2="[151,1407,409,1723]" captionTargetBox-3="[276,1315,205,1941]" captionTargetBox-4="[150,1436,483,1512]" captionTargetId-0="figure@5.[228,1370,195,1901]" captionTargetId-1="figure@6.[151,1436,193,1602]" captionTargetId-2="figure@7.[151,1436,409,1723]" captionTargetId-3="figure@11.[268,1322,193,1949]" captionTargetId-4="figure@12.[150,1436,483,1512]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 1. (a) Bayesian inference phylogenetic tree of the Uroplatus ebenaui group based on a 2326 bp alignment of four concatenated mitochondrial gene fragments (ND4, 12S rRNA, 16S rRNA, COX1). Only posterior probability values of 0.94 or greater are displayed at nodes. An outgroup (U. alluaudi) was used for rooting but omitted from the tree for graphical reasons. (b) Haplotype network independently calculated from sequences of the nuclear gene c-mos." captionText-1="FIGURE 2. Holotype of Uroplatus finaritra sp. nov. (UADBA-R 70489, APR 12522) in dorsal view (left) and a paratype (UADBA-R 70491, APR 12589) in ventral view (right) view. Scale bar indicates 10 mm." captionText-2="FIGURE 3. Variation among specimens of Uroplatus finaritra sp. nov. from Marojejy National Park. (a) photo probably showing paratype UADBA-R 70494 (APR 12692), (b) paratype UADBA-R 70491 (APR 12589), (c) paratype UADBA-R 70490 (APR 12590), (d) holotype UADBA-R 70489 (APR 12522), (e) paratype UADBA-R 70501 (MSZC 0253), (f) paratype ZSM 458/2016 (MSZC 0271). All specimens shown except (c) are females; note the differences in tail notching between this specimen and the others, and the presence of hemipenial bulges." captionText-3="FIGURE 4. Subadult female specimens of Uroplatus finaritra sp. nov. (ab) ZSM 458/2016 (MSZC 0271) and (cd) UADBA-R 70501 (MSZC 0253) in (a, d) dorsolateral and (bc) dorsal view." captionText-4="FIGURE 5. Oral mucosa colouration differences between (ac) Uroplatus finaritra sp. nov. from Marojejy and (df) U. phantasticus from (d) Fierenana, (e) Ranomafana, and (f) Kianjavato." httpUri-0="https://zenodo.org/record/2618343/files/figure.png" httpUri-1="https://zenodo.org/record/2618345/files/figure.png" httpUri-2="https://zenodo.org/record/2618347/files/figure.png" httpUri-3="https://zenodo.org/record/2618349/files/figure.png" httpUri-4="https://zenodo.org/record/2618351/files/figure.png" pageId="3" pageNumber="566">Figs. 15</figureCitation>
, Table 1
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<taxonomicName id="4C684D57FFAA954BFF22FEE7FCFF4524" box="[151,253,258,280]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B91CEAC6FFAA954BFF22FEE7FCFF4524" box="[151,253,258,280]" italics="true" pageId="3" pageNumber="566">Uroplatus</emphasis>
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sp. G—Raxworthy
<emphasis id="B91CEAC6FFAA954BFE7DFEE6FDFE4524" box="[456,508,258,280]" italics="true" pageId="3" pageNumber="566">et al.</emphasis>
(2008)
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<taxonomicName id="4C684D57FFAA954BFF22FEC4FCFF450B" box="[151,253,289,311]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B91CEAC6FFAA954BFF22FEC4FCFF450B" box="[151,253,289,311]" italics="true" pageId="3" pageNumber="566">Uroplatus</emphasis>
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sp. Ca9—
<bibRefCitation id="EFF94B25FFAA954BFED2FEC4FE59450B" author="Ratsoavina, F. M. &amp; Raminosoa, N. R. &amp; Louis, E. E. Jr. &amp; Raselimanana, A. P. &amp; Glaw, F. &amp; Vences, M." box="[359,603,289,311]" pageId="3" pageNumber="566" pagination="115 - 148" refId="ref8519" refString="Ratsoavina, F. M., Raminosoa, N. R., Louis, E. E. Jr., Raselimanana, A. P., Glaw, F. &amp; Vences, M. (2013) An overview of Madagascar's leaf tailed geckos (genus Uroplatus): species boundaries, candidate species and review of geographical distribution based on molecular data. Salamandra, 49, 115 - 148." type="journal article" year="2013">
Ratsoavina
<emphasis id="B91CEAC6FFAA954BFE6AFEC7FE10450B" box="[479,530,289,311]" italics="true" pageId="3" pageNumber="566">et al.</emphasis>
(2013)
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.
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(field number
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12522), adult female, collected on the eastern slope of
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by a tributary of the
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,
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,
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, northern
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, at ca.
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,
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(
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a.s.l.
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) on the night of
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<collectingDate id="EF92E9FCFFAA954BFD6AFE0BFF77463B" box="[735,885,494,519]" pageId="3" pageNumber="566" value="2016-05-15">15 May 2016</collectingDate>
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by
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<typeStatus id="54D38876FFAA954BFF72FDF6FD394616" box="[199,315,531,554]" pageId="3" pageNumber="566" type="paratype">Paratypes</typeStatus>
.
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<specimenCode id="DBCE9EAFFFAA954BFEFDFDF6FE144617" box="[328,534,531,555]" collectionCode="UADBA-R" pageId="3" pageNumber="566">UADBA-R 70493</specimenCode>
(field number
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12591), adult male
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,
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<specimenCode id="DBCE9EAFFFAA954BFC53FDF6F8B64617" box="[998,1204,531,555]" collectionCode="UADBA-R" pageId="3" pageNumber="566">UADBA-R 70490</specimenCode>
(
<collectionCode id="ED79AE11FFAA954BFB71FDF6F8FE4617" box="[1220,1276,531,555]" pageId="3" pageNumber="566">APR</collectionCode>
12590), adult male, and
</materialsCitation>
<materialsCitation id="3B003C89FFAA954BFEA7FDD2FF98464F" ID-GBIF-Occurrence="2147525715" collectingDate="2016-05-18" collectionCode="APR" collectorName="A. P. Raselimanana" country="Madagascar" county="Sava Region" elevation="780" latitude="-14.4559" location="Marojejy" longLatPrecision="7" longitude="49.7749" municipality="Marojejy National Park" pageId="3" pageNumber="566" specimenCode="UADBA-R 70491" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBCE9EAFFFAA954BFEA7FDD2FDE04673" box="[274,482,567,591]" collectionCode="UADBA-R" pageId="3" pageNumber="566">UADBA-R 70491</specimenCode>
(
<collectionCode id="ED79AE11FFAA954BFE41FDD2FE2E4673" box="[500,556,567,591]" pageId="3" pageNumber="566">APR</collectionCode>
12589), adult female, three specimens collected in the same locality as the holotype (
<elevation id="0045D1E7FFAA954BFEBDFDBFFDC1464F" box="[264,451,602,627]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="8.1" metricValueMin="7.5" pageId="3" pageNumber="566" unit="m" value="780.0" valueMax="810.0" valueMin="750.0">
<quantity id="4C909B31FFAA954BFEBDFDBFFD8B464E" box="[264,393,602,627]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="8.1" metricValueMin="7.5" pageId="3" pageNumber="566" unit="m" value="780.0" valueMax="810.0" valueMin="750.0">750810 m</quantity>
a.s.l.
</elevation>
) on
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<collectingDate id="EF92E9FCFFAA954BFE4CFDBEFE92464F" box="[505,656,602,627]" pageId="3" pageNumber="566" value="2016-05-18">18 May 2016</collectingDate>
</date>
by
<collectorName id="269D5302FFAA954BFD0EFDBEFF98464F" box="[699,922,602,627]" pageId="3" pageNumber="566">A. P. Raselimanana</collectorName>
</materialsCitation>
;
<materialsCitation id="3B003C89FFAA954BFC10FDBEFCC246AB" ID-GBIF-Occurrence="2147525716" collectionCode="APR" country="Madagascar" pageId="3" pageNumber="566" specimenCode="UADBA-R 70492" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBCE9EAFFFAA954BFC10FDBEF870464F" box="[933,1138,603,627]" collectionCode="UADBA-R" pageId="3" pageNumber="566">UADBA-R 70492</specimenCode>
(
<collectionCode id="ED79AE11FFAA954BFB34FDBEF8BB464F" box="[1153,1209,603,627]" pageId="3" pageNumber="566">APR</collectionCode>
12691), adult male, and
</materialsCitation>
<materialsCitation id="3B003C89FFAA954BFF70FD9AF8BE46E3" ID-GBIF-Occurrence="2147525806" collectingDate="2016-05-28" collectionCode="APR" collectorName="A. P. Raselimanana" country="Madagascar" county="Sava Region" elevation="845" latitude="-14.5074" location="Marojejy" longLatPrecision="7" longitude="49.6153" municipality="Marojejy National Park" pageId="3" pageNumber="566" specimenCode="UADBA-R 70494" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBCE9EAFFFAA954BFF70FD9AFD9046AB" box="[197,402,639,663]" collectionCode="UADBA-R" pageId="3" pageNumber="566">UADBA-R 70494</specimenCode>
(
<collectionCode id="ED79AE11FFAA954BFE2AFD9AFDDA46AB" box="[415,472,639,663]" pageId="3" pageNumber="566">APR</collectionCode>
12692), juvenile female, two specimens collected on the western slope of
<location id="8EB7600FFFAA954BFAA6FD9BF97846AB" LSID="urn:lsid:plazi:treatment:03C187C2FFAA9541FF22FF72FE784707:8EB7600FFFAA954BFAA6FD9BF97846AB" box="[1299,1402,638,663]" country="Madagascar" county="Sava Region" latitude="-14.5074" longLatPrecision="7" longitude="49.6153" municipality="Marojejy National Park" name="Marojejy" pageId="3" pageNumber="566">Marojejy</location>
by
<location id="8EB7600FFFAA954BFF22FD47FD3A4687" LSID="urn:lsid:plazi:treatment:03C187C2FFAA9541FF22FF72FE784707:8EB7600FFFAA954BFF22FD47FD3A4687" box="[151,312,674,699]" country="Madagascar" county="Sava Region" latitude="-14.5074" longLatPrecision="7" longitude="49.6153" municipality="Marojejy National Park" name="Tsiasisa River" pageId="3" pageNumber="566">Tsiasisa River</location>
,
<quantity id="4C909B31FFAA954BFEF1FD46FD964687" box="[324,404,674,699]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="566" unit="km" value="2.2">2.2 km</quantity>
east of
<location id="8EB7600FFFAA954BFE45FD47FE994687" LSID="urn:lsid:plazi:treatment:03C187C2FFAA9541FF22FF72FE784707:8EB7600FFFAA954BFE45FD47FE994687" box="[496,667,674,699]" country="Madagascar" county="Sava Region" latitude="-14.5074" longLatPrecision="7" longitude="49.6153" municipality="Marojejy National Park" name="Antanimbaribe" pageId="3" pageNumber="566">Antanimbaribe</location>
,
<collectingMunicipality id="6BB3ACAEFFAA954BFD1DFD47FFB44687" box="[680,950,674,699]" pageId="3" pageNumber="566">Marojejy National Park</collectingMunicipality>
,
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, northern
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, at ca.
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,
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(
<elevation id="0045D1E7FFAA954BFE1CFD23FE3646E3" box="[425,564,710,735]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="3" pageNumber="566" unit="m" value="845.0">
<quantity id="4C909B31FFAA954BFE1CFD23FDF446E2" box="[425,502,710,735]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="3" pageNumber="566" unit="m" value="845.0">845 m</quantity>
a.s.l.
</elevation>
) on the night of
<date id="FFD61014FFAA954BFCB3FD22FFA646E3" box="[774,932,710,735]" pageId="3" pageNumber="566" value="2016-05-28">
<collectingDate id="EF92E9FCFFAA954BFCB3FD22FFA646E3" box="[774,932,710,735]" pageId="3" pageNumber="566" value="2016-05-28">28 May 2016</collectingDate>
</date>
by
<collectorName id="269D5302FFAA954BFC63FD22F8BE46E3" box="[982,1212,710,735]" pageId="3" pageNumber="566">A. P. Raselimanana</collectorName>
</materialsCitation>
;
<materialsCitation id="3B003C89FFAA954BFB7EFD22FDF8473F" ID-GBIF-Occurrence="2147525717" collectionCode="MSZC" country="Madagascar" pageId="3" pageNumber="566" specimenCode="UADBA-R 70501, MSZC 0253" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBCE9EAFFFAA954BFB7EFD22F99E46E3" box="[1227,1436,710,735]" collectionCode="UADBA-R" pageId="3" pageNumber="566">UADBA-R 70501</specimenCode>
(
<specimenCode id="DBCE9EAFFFAA954BFF2BFD0EFD32473F" box="[158,304,746,771]" collectionCode="MSZC" pageId="3" pageNumber="566">MSZC 0253</specimenCode>
), subadult female
</materialsCitation>
,
<materialsCitation id="3B003C89FFAA954BFDB0FD0EFEDE4752" ID-GBIF-Occurrence="2147525684" collectingDate="2016-11-21" collectingDateMax="2016-11-23" collectingDateMin="2016-11-21" collectionCode="ZSM, MSZC" collectorName="M. D. Scherz &amp; M. Bletz &amp; A. Rakotoarison &amp; J. Razafindraibe &amp; A. Razafimanantsoa &amp; M. Vences" country="Madagascar" county="Sava Region" elevation="450" latitude="-14.438" location="Camp" longLatPrecision="76" longitude="49.776" municipality="Marojejy National Park" pageId="3" pageNumber="566" specimenCode="ZSM 458, MSZC 0271" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBCE9EAFFFAA954BFDB0FD0EFE72473F" box="[517,624,746,771]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="566" type="Museum">ZSM 458</specimenCode>
/2016 (
<specimenCode id="DBCE9EAFFFAA954BFD75FD0EFF53473F" box="[704,849,747,771]" collectionCode="MSZC" pageId="3" pageNumber="566">MSZC 0271</specimenCode>
), subadult female, two specimens collected near the path below
<location id="8EB7600FFFAA954BFE90FCEAFD6A471B" LSID="urn:lsid:plazi:treatment:03C187C2FFAA9541FF22FF72FE784707:8EB7600FFFAA954BFE90FCEAFD6A471B" box="[293,360,783,807]" country="Madagascar" county="Sava Region" latitude="-14.438" longLatPrecision="76" longitude="49.776" municipality="Marojejy National Park" name="Camp" pageId="3" pageNumber="566">Camp</location>
1 (
<location id="8EB7600FFFAA954BFE22FCEAFE4F471B" LSID="urn:lsid:plazi:treatment:03C187C2FFAA9541FF22FF72FE784707:8EB7600FFFAA954BFE22FCEAFE4F471B" box="[407,589,782,807]" country="Madagascar" county="Sava Region" latitude="-14.438" longLatPrecision="76" longitude="49.776" municipality="Marojejy National Park" name="Camp Mantella" pageId="3" pageNumber="566">Camp Mantella</location>
) of
<collectingMunicipality id="6BB3ACAEFFAA954BFD33FCEBFF9F471B" box="[646,925,782,807]" pageId="3" pageNumber="566">Marojejy National Park</collectingMunicipality>
,
<collectingCounty id="62B64E58FFAA954BFC1BFCEAF847471B" box="[942,1093,782,807]" pageId="3" pageNumber="566">Sava Region</collectingCounty>
, northern
<collectingCountry id="F37F7644FFAA954BFB74FCEBF948471B" box="[1217,1354,782,807]" name="Madagascar" pageId="3" pageNumber="566">Madagascar</collectingCountry>
, at ca.
<geoCoordinate id="EE5C5013FFAA954BFF22FCD6FD034777" box="[151,257,819,843]" direction="south" orientation="latitude" pageId="3" pageNumber="566" precision="55" value="-14.438">14.438°S</geoCoordinate>
,
<geoCoordinate id="EE5C5013FFAA954BFEA4FCD7FD784777" box="[273,378,818,843]" direction="east" orientation="longitude" pageId="3" pageNumber="566" precision="55" value="49.776">49.776°E</geoCoordinate>
(
<elevation id="0045D1E7FFAA954BFE38FCD7FE1A4777" box="[397,536,818,843]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="566" unit="m" value="450.0">
<quantity id="4C909B31FFAA954BFE38FCD7FDD84776" box="[397,474,818,843]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="566" unit="m" value="450.0">450 m</quantity>
a.s.l.
</elevation>
) on
<date id="FFD61014FFAA954BFDE0FCD6FF624777" box="[597,864,818,843]" pageId="3" pageNumber="566" value="2016-11-21" valueMax="2016-11-23" valueMin="2016-11-21">
<collectingDate id="EF92E9FCFFAA954BFDE0FCD6FF624777" box="[597,864,818,843]" pageId="3" pageNumber="566" value="2016-11-21" valueMax="2016-11-23" valueMin="2016-11-21">2123 November 2016</collectingDate>
</date>
by
<collectorName id="269D5302FFAA954BFC26FCD6F8364777" box="[915,1076,818,843]" pageId="3" pageNumber="566">M. D. Scherz</collectorName>
,
<collectorName id="269D5302FFAA954BFBF6FCD6F8AF4777" box="[1091,1197,818,843]" pageId="3" pageNumber="566">M. Bletz</collectorName>
,
<collectorName id="269D5302FFAA954BFB09FCD6F9784777" box="[1212,1402,818,843]" pageId="3" pageNumber="566">A. Rakotoarison</collectorName>
,
<collectorName id="269D5302FFAA954BFA3FFCD6FD354753" pageId="3" pageNumber="566">J. Razafindraibe</collectorName>
,
<collectorName id="269D5302FFAA954BFEF7FCB2FE2A4753" box="[322,552,854,879]" pageId="3" pageNumber="566">A. Razafimanantsoa</collectorName>
, and
<collectorName id="269D5302FFAA954BFDD7FCB2FEDE4752" box="[610,732,855,878]" pageId="3" pageNumber="566">M. Vences</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C372655FFFAA954BFF72FC9FFF3A47E7" pageId="3" pageNumber="566" type="etymology">
<paragraph id="8BD736D4FFAA954BFF72FC9FFF3A47E7" blockId="3.[151,1437,422,1923]" pageId="3" pageNumber="566">
<emphasis id="B91CEAC6FFAA954BFF72FC9FFD4547AF" bold="true" box="[199,327,890,915]" italics="true" pageId="3" pageNumber="566">Etymology.</emphasis>
Finaritra is a Malagasy word for special greetings but also means healthy and happy. We refer to our delight in describing this splendid and exceptionally large species from a clade of generally small-sized leaftailed geckos. The name is an invariable noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C372655FFFAA954CFF72FC03FAAD477A" lastPageId="4" lastPageNumber="567" pageId="3" pageNumber="566" type="diagnosis">
<paragraph id="8BD736D4FFAA954BFF72FC03FF7C417F" blockId="3.[151,1437,422,1923]" pageId="3" pageNumber="566">
<emphasis id="B91CEAC6FFAA954BFF72FC03FE1F47C3" box="[199,541,998,1023]" italics="true" pageId="3" pageNumber="566">
<emphasis id="B91CEAC6FFAA954BFF72FC03FD4347C2" bold="true" box="[199,321,998,1022]" italics="true" pageId="3" pageNumber="566">Diagnosis.</emphasis>
<taxonomicName id="4C684D57FFAA954BFEF2FC03FE1F47C3" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[327,541,998,1023]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">Uroplatus finaritra</taxonomicName>
</emphasis>
<emphasis id="B91CEAC6FFAA954BFD91FC02FE7F47C3" bold="true" box="[548,637,999,1023]" pageId="3" pageNumber="566">
<taxonomicNameLabel id="A22F57BDFFAA954BFD91FC02FE7F47C3" box="[548,637,999,1023]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is assigned to the
<taxonomicName id="4C684D57FFAA954BFCFDFC03F81A47C2" baseAuthorityName="Boettger" baseAuthorityYear="1879" box="[840,1048,998,1022]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="ebenaui">
<emphasis id="B91CEAC6FFAA954BFCFDFC03F81A47C2" box="[840,1048,998,1022]" italics="true" pageId="3" pageNumber="566">Uroplatus ebenaui</emphasis>
</taxonomicName>
group based on its relatively small size and its triangular head with supraocular spines, and overall leaf-mimicking aspect. It is characterised by (1) a dark red oral mucosa, (2) a comparatively large body size (
<quantity id="4C909B31FFAA954BFCD5FBCBF808407A" box="[864,1034,1070,1095]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="9.53" metricValueMin="8.25" pageId="3" pageNumber="566" unit="mm" value="88.9" valueMax="95.3" valueMin="82.5">82.595.3 mm</quantity>
SVL in adults), (3) 78 lamellae (exceptionally 6) under the third finger and toe, and (4) a long (length 5365% of SVL), wide (width 1618% of SVL), leaf-shaped tail. It differs from all members of the
<taxonomicName id="4C684D57FFAA954BFC8EFB92FFD640B2" baseAuthorityName="Schneider" baseAuthorityYear="1797" box="[827,980,1142,1166]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B91CEAC6FFAA954BFC8EFB92FFD640B2" box="[827,980,1142,1166]" italics="true" pageId="3" pageNumber="566">U. fimbriatus</emphasis>
</taxonomicName>
group (
<taxonomicName id="4C684D57FFAA954BFB86FB92F8CE40B2" baseAuthorityName="Schneider" baseAuthorityYear="1797" box="[1075,1228,1142,1166]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B91CEAC6FFAA954BFB86FB92F8CE40B2" box="[1075,1228,1142,1166]" italics="true" pageId="3" pageNumber="566">U. fimbriatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D57FFAA954BFB68FB92F96D40B2" box="[1245,1391,1143,1166]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B91CEAC6FFAA954BFB68FB92F96D40B2" box="[1245,1391,1143,1166]" italics="true" pageId="3" pageNumber="566">U. giganteus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D57FFAA954BFA34FB92FCEA408F" authorityName="Böhme &amp; Ibisch" authorityYear="1990" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="henkeli">
<emphasis id="B91CEAC6FFAA954BFA34FB92FCEA408F" italics="true" pageId="3" pageNumber="566">U. henkeli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D57FFAA954BFF42FB7EFD6C408F" box="[247,366,1179,1203]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="sikorae">
<emphasis id="B91CEAC6FFAA954BFF42FB7EFD6C408F" box="[247,366,1179,1203]" italics="true" pageId="3" pageNumber="566">U. sikorae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C684D57FFAA954BFE18FB7EFE20408E" authorityName="Böhme &amp; Ibisch" authorityYear="1990" box="[429,546,1179,1202]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="sameiti">
<emphasis id="B91CEAC6FFAA954BFE18FB7EFE20408E" box="[429,546,1179,1202]" italics="true" pageId="3" pageNumber="566">U. sameiti</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C684D57FFAA954BFDD1FB7EFEE3408F" baseAuthorityName="Duméril &amp; Bibron" baseAuthorityYear="1836" box="[612,737,1179,1203]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91CEAC6FFAA954BFDD1FB7EFEE3408F" box="[612,737,1179,1203]" italics="true" pageId="3" pageNumber="566">U. lineatus</emphasis>
</taxonomicName>
by its smaller size (SVL
<quantity id="4C909B31FFAA954BFBB0FB7FF8A8408E" box="[1029,1194,1178,1203]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="9.53" metricValueMin="8.25" pageId="3" pageNumber="566" unit="mm" value="88.9" valueMax="95.3" valueMin="82.5">82.595.3 mm</quantity>
vs.
<quantity id="4C909B31FFAA954BFB6FFB7FF966408F" box="[1242,1380,1178,1203]" metricMagnitude="-1" metricUnit="m" metricValue="1.425" metricValueMax="2.0" metricValueMin="0.85" pageId="3" pageNumber="566" unit="mm" value="142.5" valueMax="200.0" valueMin="85.0">85200 mm</quantity>
) and the lack of lateral dermal fringes on any part of the body (vs. presence except in
<taxonomicName id="4C684D57FFAA954BFBF8FB5AF8C840EA" baseAuthorityName="Duméril &amp; Bibron" baseAuthorityYear="1836" box="[1101,1226,1214,1238]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91CEAC6FFAA954BFBF8FB5AF8C840EA" box="[1101,1226,1214,1238]" italics="true" pageId="3" pageNumber="566">U. lineatus</emphasis>
</taxonomicName>
). Also,
<taxonomicName id="4C684D57FFAA954BFA9EFB5BFCF440C7" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAA954BFA9EFB5BFCF440C7" italics="true" pageId="3" pageNumber="566">Uroplatus finaritra</emphasis>
</taxonomicName>
<emphasis id="B91CEAC6FFAA954BFF4BFB06FD5B40C7" bold="true" box="[254,345,1251,1275]" pageId="3" pageNumber="566">
<taxonomicNameLabel id="A22F57BDFFAA954BFF4BFB06FD5B40C7" box="[254,345,1251,1275]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
diverges from the species of the
<taxonomicName id="4C684D57FFAA954BFD62FB06FF6D40C7" baseAuthorityName="Schneider" baseAuthorityYear="1797" box="[727,879,1251,1275]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B91CEAC6FFAA954BFD62FB06FF6D40C7" box="[727,879,1251,1275]" italics="true" pageId="3" pageNumber="566">U. fimbriatus</emphasis>
</taxonomicName>
group by their lack of lateral body compression. The prominent triangular head, supraocular spines, and rather smooth skin distinguish
<taxonomicName id="4C684D57FFAA954BFBD1FAE3F9394123" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[1124,1339,1286,1311]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAA954BFBD1FAE3F9394123" box="[1124,1339,1286,1311]" italics="true" pageId="3" pageNumber="566">Uroplatus finaritra</emphasis>
</taxonomicName>
<emphasis id="B91CEAC6FFAA954BFAF7FAE2F99E4123" bold="true" box="[1346,1436,1287,1311]" pageId="3" pageNumber="566">
<taxonomicNameLabel id="A22F57BDFFAA954BFAF7FAE2F99E4123" box="[1346,1436,1287,1311]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<emphasis id="B91CEAC6FFAA954BFF61FACEFE85417F" box="[212,647,1323,1347]" italics="true" pageId="3" pageNumber="566">
<taxonomicName id="4C684D57FFAA954BFF61FACEFD54417F" authorityName="Mocquard" authorityYear="1894" box="[212,342,1323,1347]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="alluaudi">U. alluaudi</taxonomicName>
,
<taxonomicName id="4C684D57FFAA954BFED7FACEFDF0417F" authorityName="Mocquard" authorityYear="1908" box="[354,498,1323,1347]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="guentheri">U. guentheri</taxonomicName>
,
<taxonomicName id="4C684D57FFAA954BFE48FACEFE85417F" authorityName="Nussbaum &amp; Raxworthy" authorityYear="1994" box="[509,647,1323,1347]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="malahelo">U. malahelo</taxonomicName>
</emphasis>
and
<emphasis id="B91CEAC6FFAA954BFD0BFACEFF7C417F" box="[702,894,1323,1347]" italics="true" pageId="3" pageNumber="566">
<taxonomicName id="4C684D57FFAA954BFD0BFACEFF7B417F" box="[702,889,1323,1347]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="pietschmanni">U. pietschmanni</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD736D4FFAA954BFF72FAAAFD3841CB" blockId="3.[151,1437,422,1923]" pageId="3" pageNumber="566">
Amongst the
<taxonomicName id="4C684D57FFAA954BFEE8FAAAFDDA415A" baseAuthorityName="Boettger" baseAuthorityYear="1879" box="[349,472,1358,1382]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="ebenaui">
<emphasis id="B91CEAC6FFAA954BFEE8FAAAFDDA415A" box="[349,472,1358,1382]" italics="true" pageId="3" pageNumber="566">U. ebenaui</emphasis>
</taxonomicName>
group, the rather long tail (TAL/SVL 0.530.68) differentiates
<taxonomicName id="4C684D57FFAA954BFB28FAABF976415B" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[1181,1396,1358,1383]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAA954BFB28FAABF976415B" box="[1181,1396,1358,1383]" italics="true" pageId="3" pageNumber="566">Uroplatus finaritra</emphasis>
</taxonomicName>
<emphasis id="B91CEAC6FFAA954BFACFFAAAFCCA41B7" bold="true" pageId="3" pageNumber="566">
<taxonomicNameLabel id="A22F57BDFFAA954BFACFFAAAFCCA41B7" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C684D57FFAA954BFEA5FA96FD8F41B7" baseAuthorityName="Boettger" baseAuthorityYear="1879" box="[272,397,1395,1419]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="ebenaui">
<emphasis id="B91CEAC6FFAA954BFEA5FA96FD8F41B7" box="[272,397,1395,1419]" italics="true" pageId="3" pageNumber="566">U. ebenaui</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D57FFAA954BFE28FA96FDF441B7" box="[413,502,1395,1419]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fiera">
<emphasis id="B91CEAC6FFAA954BFE28FA96FDF441B7" box="[413,502,1395,1419]" italics="true" pageId="3" pageNumber="566">U. fiera</emphasis>
</taxonomicName>
,
<emphasis id="B91CEAC6FFAA954BFDB0FA96FE9441B7" box="[517,662,1395,1419]" italics="true" pageId="3" pageNumber="566">
<taxonomicName id="4C684D57FFAA954BFDB0FA96FE9041B7" authorityName="Ratsoavina, Gehring, Scherz, Vieites, Glaw &amp; Vences" authorityYear="2017" box="[517,658,1395,1419]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fotsivava">U. fotsivava</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C684D57FFAA954BFD65FA96FF6441B7" box="[720,870,1395,1419]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="kelirambo">
<emphasis id="B91CEAC6FFAA954BFD65FA96FF6441B7" box="[720,870,1395,1419]" italics="true" pageId="3" pageNumber="566">U. kelirambo</emphasis>
</taxonomicName>
(TAL/SVL 0.220.44), and furthermore it differs from
<taxonomicName id="4C684D57FFAA954BFF63FA72FD324193" box="[214,304,1431,1455]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fiera">
<emphasis id="B91CEAC6FFAA954BFF63FA72FD324193" box="[214,304,1431,1455]" italics="true" pageId="3" pageNumber="566">U. fiera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D57FFAA954BFEDFFA72FDF14193" authorityName="Ratsoavina, Gehring, Scherz, Vieites, Glaw &amp; Vences" authorityYear="2017" box="[362,499,1431,1455]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="fotsivava">
<emphasis id="B91CEAC6FFAA954BFEDFFA72FDF14193" box="[362,499,1431,1455]" italics="true" pageId="3" pageNumber="566">U. fotsivava</emphasis>
</taxonomicName>
by a pigmented oral mucosa (vs. unpigmented). The new species differs from
<taxonomicName id="4C684D57FFAA954BFA34FA72FD0341EF" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="finiavana">
<emphasis id="B91CEAC6FFAA954BFA34FA72FD0341EF" italics="true" pageId="3" pageNumber="566">U. finiavana</emphasis>
</taxonomicName>
by a wider tail (TAW/SVL 0.160.18 vs. 0.050.14), and by its pigmented oral mucosa (vs. unpigmented).
</paragraph>
<paragraph id="8BD736D4FFAA954BFF72F9E6FFE342F3" blockId="3.[151,1437,422,1923]" pageId="3" pageNumber="566">
By its large tail and pigmented oral mucosa, the new species is most similar to
<taxonomicName id="4C684D57FFAA954BFB81F9E6F8B34227" authorityName="Nussbaum &amp; Raxworthy" authorityYear="1995" box="[1076,1201,1539,1563]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="malama">
<emphasis id="B91CEAC6FFAA954BFB81F9E6F8B34227" box="[1076,1201,1539,1563]" italics="true" pageId="3" pageNumber="566">U. malama</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D57FFAA954BFB50F9E6F9964227" box="[1253,1428,1539,1563]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="phantasticus">
<emphasis id="B91CEAC6FFAA954BFB50F9E6F9964227" box="[1253,1428,1539,1563]" italics="true" pageId="3" pageNumber="566">U. phantasticus</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="4C684D57FFAA954BFE8AF9C2FDED4202" box="[319,495,1574,1598]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="phantasticus">
<emphasis id="B91CEAC6FFAA954BFE8AF9C2FDED4202" box="[319,495,1574,1598]" italics="true" pageId="3" pageNumber="566">U. phantasticus</emphasis>
</taxonomicName>
by larger size (SVL
<quantity id="4C909B31FFAA954BFD6AF9C3FF5B4202" box="[735,857,1574,1599]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" metricValueMax="9.5" metricValueMin="8.3" pageId="3" pageNumber="566" unit="mm" value="89.0" valueMax="95.0" valueMin="83.0">8395 mm</quantity>
vs.
<quantity id="4C909B31FFAA954BFC3CF9C3F8054203" box="[905,1031,1574,1599]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" metricValueMax="7.6" metricValueMin="5.2" pageId="3" pageNumber="566" unit="mm" value="64.0" valueMax="76.0" valueMin="52.0">5276 mm</quantity>
), a generally shorter tail (TAL/SVL 0.530.65 vs. 0.620.76), a typically higher number of lamellae under the third finger and toe (68 vs. 57), and the absence of black pigment of the oral mucosa (oral mucosa dark red vs. black). It differs from
<taxonomicName id="4C684D57FFAA954BFB53F98AF96642BA" authorityName="Nussbaum &amp; Raxworthy" authorityYear="1995" box="[1254,1380,1646,1670]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="malama">
<emphasis id="B91CEAC6FFAA954BFB53F98AF96642BA" box="[1254,1380,1646,1670]" italics="true" pageId="3" pageNumber="566">U. malama</emphasis>
</taxonomicName>
by a spinier integument in some adult males, by the absence of black pigment of the oral mucosa (oral mucosa dark red vs. black), and probably by a narrower tail (TAW/SVL 0.160.18 vs. 0.24).
</paragraph>
<paragraph id="8BD736D4FFAA954BFF72F93EFECB43BF" blockId="3.[151,1437,422,1923]" pageId="3" pageNumber="566">
In the mitochondrial phylogeny, the new species occupies an isolated position within the
<taxonomicName id="4C684D57FFAA954BFB78F93EF94942CF" baseAuthorityName="Boettger" baseAuthorityYear="1879" box="[1229,1355,1755,1779]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="ebenaui">
<emphasis id="B91CEAC6FFAA954BFB78F93EF94942CF" box="[1229,1355,1755,1779]" italics="true" pageId="3" pageNumber="566">U. ebenaui</emphasis>
</taxonomicName>
group, without affinities to the morphologically similar
<taxonomicName id="4C684D57FFAA954BFD03F91AFF67432A" box="[694,869,1790,1814]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="phantasticus">
<emphasis id="B91CEAC6FFAA954BFD03F91AFF67432A" box="[694,869,1790,1814]" italics="true" pageId="3" pageNumber="566">U. phantasticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D57FFAA954BFC2EF91AF815432A" authorityName="Nussbaum &amp; Raxworthy" authorityYear="1995" box="[923,1047,1790,1814]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="malama">
<emphasis id="B91CEAC6FFAA954BFC2EF91AF815432A" box="[923,1047,1790,1814]" italics="true" pageId="3" pageNumber="566">U. malama</emphasis>
</taxonomicName>
(which are not each other's closest relatives either). The nuclear gene c-mos reveals haplotype sharing of the new species with
<taxonomicName id="4C684D57FFAA954BFB10F8C6F9384307" box="[1189,1338,1827,1851]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="566" phylum="Chordata" rank="species" species="kelirambo">
<emphasis id="B91CEAC6FFAA954BFB10F8C6F9384307" box="[1189,1338,1827,1851]" italics="true" pageId="3" pageNumber="566">U. kelirambo</emphasis>
</taxonomicName>
and two undescribed candidate species (
<emphasis id="B91CEAC6FFAA954BFDADF8A2FE314362" box="[536,563,1863,1886]" italics="true" pageId="3" pageNumber="566">U.</emphasis>
sp. Ca3 and Ca4) occurring in the same region of
<collectingCountry id="F37F7644FFAA954BFB6FF8A3F9614363" box="[1242,1379,1862,1887]" name="Madagascar" pageId="3" pageNumber="566">Madagascar</collectingCountry>
, but morphologically very different by their short tails.
</paragraph>
<paragraph id="8BD736D4FFAD954CFF26FCE9FAAD477A" blockId="4.[147,1920,780,838]" pageId="4" pageNumber="567">
<emphasis id="B91CEAC6FFAD954CFF26FCE9FCFC4719" bold="true" box="[147,254,780,805]" pageId="4" pageNumber="567">TABLE 1</emphasis>
. Morphological measurements and scale counts of
<taxonomicName id="4C684D57FFAD954CFCACFCE9FFDD471A" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[793,991,780,806]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="567" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAD954CFCACFCE9FFDD471A" box="[793,991,780,806]" italics="true" pageId="4" pageNumber="567">Uroplatus finaritra</emphasis>
</taxonomicName>
<emphasis id="B91CEAC6FFAD954CFC52FCE9F83C4719" bold="true" box="[999,1086,780,805]" pageId="4" pageNumber="567">
<taxonomicNameLabel id="A22F57BDFFAD954CFC52FCE9F83C4719" box="[999,1086,780,805]" pageId="4" pageNumber="567" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
For abbreviations used, refer to Methods; additional abbreviations: M, male; F, female; sa, subadult; j, juvenile; HT,
<typeStatus id="54D38876FFAD954CFDBEFCC8FE64477A" box="[523,614,813,838]" pageId="4" pageNumber="567" type="holotype">holotype</typeStatus>
; PT,
<typeStatus id="54D38876FFAD954CFD29FCC8FEF4477A" box="[668,758,813,838]" pageId="4" pageNumber="567" type="paratype">paratype</typeStatus>
; na, not applicable (due to missing original tail); nm, not measured. All measurements in mm.
</paragraph>
</subSubSection>
<paragraph id="8BD736D4FFAD954CFF26FC97FB2C4166" pageId="4" pageNumber="567">
<table id="F968C474FFAD6AB7FF26FC97FB774166" box="[147,1909,882,1370]" gridcols="22" gridrows="9" pageId="4" pageNumber="567">
<tr id="35583496FFAD6AB7FF26FC97FB7747B7" box="[147,1909,882,907]" gridrow="0" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FC97FD4E47B7" box="[147,332,882,907]" gridcol="0" gridrow="0" pageId="4" pageNumber="567">Museum Number</th>
<th id="76895DEAFFAD6AB7FEDDFC97FDF247B7" box="[360,496,882,907]" gridcol="1" gridrow="0" pageId="4" pageNumber="567">Field number</th>
<th id="76895DEAFFAD6AB7FDB2FC97FE6447B7" box="[519,614,882,907]" gridcol="2" gridrow="0" pageId="4" pageNumber="567">Sex</th>
<th id="76895DEAFFAD6AB7FDCDFC97FEAA47B7" box="[632,680,882,907]" gridcol="3" gridrow="0" pageId="4" pageNumber="567">SVL</th>
<th id="76895DEAFFAD6AB7FD03FC97FEE147B7" box="[694,739,882,907]" gridcol="4" gridrow="0" pageId="4" pageNumber="567">TaL</th>
<th id="76895DEAFFAD6AB7FD5BFC97FF2A47B7" box="[750,808,882,907]" gridcol="5" gridrow="0" pageId="4" pageNumber="567">TAW</th>
<th id="76895DEAFFAD6AB7FC82FC97FF6747B7" box="[823,869,882,907]" gridcol="6" gridrow="0" pageId="4" pageNumber="567">HL2</th>
<th id="76895DEAFFAD6AB7FCC7FC97FFAB47B7" box="[882,937,882,907]" gridcol="7" gridrow="0" pageId="4" pageNumber="567">AGD</th>
<th id="76895DEAFFAD6AB7FC02FC97FFDB47B7" box="[951,985,882,907]" gridcol="8" gridrow="0" pageId="4" pageNumber="567">ED</th>
<th id="76895DEAFFAD6AB7FC50FC97F82847B7" box="[997,1066,882,907]" gridcol="9" gridrow="0" pageId="4" pageNumber="567">EARD</th>
<th id="76895DEAFFAD6AB7FB8CFC97F86447B7" box="[1081,1126,882,907]" gridcol="10" gridrow="0" pageId="4" pageNumber="567">SL</th>
<th id="76895DEAFFAD6AB7FBC7FC97F8A247B7" box="[1138,1184,882,907]" gridcol="11" gridrow="0" pageId="4" pageNumber="567">HL1</th>
<th id="76895DEAFFAD6AB7FB1BFC97F8D947B7" box="[1198,1243,882,907]" gridcol="12" gridrow="0" pageId="4" pageNumber="567">HW</th>
<th id="76895DEAFFAD6AB7FB52FC97F92547B7" box="[1255,1319,882,907]" gridcol="13" gridrow="0" pageId="4" pageNumber="567">FORL</th>
<th id="76895DEAFFAD6AB7FA83FC97F96647B7" box="[1334,1380,882,907]" gridcol="14" gridrow="0" pageId="4" pageNumber="567">HIL</th>
<th id="76895DEAFFAD6AB7FADAFC97F9A347B7" box="[1391,1441,882,907]" gridcol="15" gridrow="0" pageId="4" pageNumber="567">NTL</th>
<th id="76895DEAFFAD6AB7FA1BFC97F9E747B7" box="[1454,1509,882,907]" gridcol="16" gridrow="0" pageId="4" pageNumber="567">IRPL</th>
<th id="76895DEAFFAD6AB7FA46FC97FA1E47B7" box="[1523,1564,882,907]" gridcol="17" gridrow="0" pageId="4" pageNumber="567">INS</th>
<th id="76895DEAFFAD6AB7F99CFC97FA6547B7" box="[1577,1639,882,907]" gridcol="18" gridrow="0" pageId="4" pageNumber="567">SUPL</th>
<th id="76895DEAFFAD6AB7F9C0FC97FAB247B7" box="[1653,1712,882,907]" gridcol="19" gridrow="0" pageId="4" pageNumber="567">INFL</th>
<th id="76895DEAFFAD6AB7F908FC97FB1247B7" box="[1725,1808,882,907]" gridcol="20" gridrow="0" pageId="4" pageNumber="567">LAM3F</th>
<th id="76895DEAFFAD6AB7F894FC97FB7747B7" box="[1825,1909,882,907]" gridcol="21" gridrow="0" pageId="4" pageNumber="567">LAM3T</th>
</tr>
<tr id="35583496FFAD6AB7FF26FC48FB7747FA" box="[147,1909,941,966]" gridrow="1" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FC48FD4E47FA" box="[147,332,941,966]" gridcol="0" gridrow="1" pageId="4" pageNumber="567">ZSM 458/2016</th>
<td id="76895DEAFFAD6AB7FEDDFC48FDF247FA" box="[360,496,941,966]" gridcol="1" gridrow="1" pageId="4" pageNumber="567">MSZC 0 271</td>
<td id="76895DEAFFAD6AB7FDB2FC48FE6447FA" box="[519,614,941,966]" gridcol="2" gridrow="1" pageId="4" pageNumber="567">sa F (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFC48FEAA47FA" box="[632,680,941,966]" gridcol="3" gridrow="1" pageId="4" pageNumber="567">71.2</td>
<td id="76895DEAFFAD6AB7FD03FC48FEE147FA" box="[694,739,941,966]" gridcol="4" gridrow="1" pageId="4" pageNumber="567">48.4</td>
<td id="76895DEAFFAD6AB7FD5BFC48FF2A47FA" box="[750,808,941,966]" gridcol="5" gridrow="1" pageId="4" pageNumber="567">13.8</td>
<td id="76895DEAFFAD6AB7FC82FC48FF6747FA" box="[823,869,941,966]" gridcol="6" gridrow="1" pageId="4" pageNumber="567">15.6</td>
<td id="76895DEAFFAD6AB7FCC7FC48FFAB47FA" box="[882,937,941,966]" gridcol="7" gridrow="1" pageId="4" pageNumber="567">34.2</td>
<td id="76895DEAFFAD6AB7FC02FC48FFDB47FA" box="[951,985,941,966]" gridcol="8" gridrow="1" pageId="4" pageNumber="567">5.4</td>
<td id="76895DEAFFAD6AB7FC50FC48F82847FA" box="[997,1066,941,966]" gridcol="9" gridrow="1" pageId="4" pageNumber="567">1.0</td>
<td id="76895DEAFFAD6AB7FB8CFC48F86447FA" box="[1081,1126,941,966]" gridcol="10" gridrow="1" pageId="4" pageNumber="567">9.2</td>
<td id="76895DEAFFAD6AB7FBC7FC48F8A247FA" box="[1138,1184,941,966]" gridcol="11" gridrow="1" pageId="4" pageNumber="567">21.9</td>
<td id="76895DEAFFAD6AB7FB1BFC48F8D947FA" box="[1198,1243,941,966]" gridcol="12" gridrow="1" pageId="4" pageNumber="567">15.1</td>
<td id="76895DEAFFAD6AB7FB52FC48F92547FA" box="[1255,1319,941,966]" gridcol="13" gridrow="1" pageId="4" pageNumber="567">28.5</td>
<td id="76895DEAFFAD6AB7FA83FC48F96647FA" box="[1334,1380,941,966]" gridcol="14" gridrow="1" pageId="4" pageNumber="567">43.1</td>
<td id="76895DEAFFAD6AB7FADAFC48F9A347FA" box="[1391,1441,941,966]" gridcol="15" gridrow="1" pageId="4" pageNumber="567">10.6</td>
<td id="76895DEAFFAD6AB7FA1BFC48F9E747FA" box="[1454,1509,941,966]" gridcol="16" gridrow="1" pageId="4" pageNumber="567">1.1</td>
<td id="76895DEAFFAD6AB7FA46FC48FA1E47FA" box="[1523,1564,941,966]" gridcol="17" gridrow="1" pageId="4" pageNumber="567">8</td>
<td id="76895DEAFFAD6AB7F99CFC48FA6547FA" box="[1577,1639,941,966]" gridcol="18" gridrow="1" pageId="4" pageNumber="567">28/28</td>
<td id="76895DEAFFAD6AB7F9C0FC48FAB247FA" box="[1653,1712,941,966]" gridcol="19" gridrow="1" pageId="4" pageNumber="567">23/26</td>
<td id="76895DEAFFAD6AB7F908FC48FB1247FA" box="[1725,1808,941,966]" gridcol="20" gridrow="1" pageId="4" pageNumber="567">8</td>
<td id="76895DEAFFAD6AB7F894FC48FB7747FA" box="[1825,1909,941,966]" gridcol="21" gridrow="1" pageId="4" pageNumber="567">8</td>
</tr>
<tr id="35583496FFAD6AB7FF26FC02FB77403C" box="[147,1909,999,1024]" gridrow="2" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FC02FD4E403C" box="[147,332,999,1024]" gridcol="0" gridrow="2" pageId="4" pageNumber="567">UADBA-R 70493</th>
<td id="76895DEAFFAD6AB7FEDDFC02FDF2403C" box="[360,496,999,1024]" gridcol="1" gridrow="2" pageId="4" pageNumber="567">APR 12591</td>
<td id="76895DEAFFAD6AB7FDB2FC02FE64403C" box="[519,614,999,1024]" gridcol="2" gridrow="2" pageId="4" pageNumber="567">M (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFC02FEAA403C" box="[632,680,999,1024]" gridcol="3" gridrow="2" pageId="4" pageNumber="567">91.3</td>
<td id="76895DEAFFAD6AB7FD03FC02FEE1403C" box="[694,739,999,1024]" gridcol="4" gridrow="2" pageId="4" pageNumber="567">na</td>
<td id="76895DEAFFAD6AB7FD5BFC02FF2A403C" box="[750,808,999,1024]" gridcol="5" gridrow="2" pageId="4" pageNumber="567">na</td>
<td id="76895DEAFFAD6AB7FC82FC02FF67403C" box="[823,869,999,1024]" gridcol="6" gridrow="2" pageId="4" pageNumber="567">24.2</td>
<td id="76895DEAFFAD6AB7FCC7FC02FFAB403C" box="[882,937,999,1024]" gridcol="7" gridrow="2" pageId="4" pageNumber="567">49.1</td>
<td id="76895DEAFFAD6AB7FC02FC02FFDB403C" box="[951,985,999,1024]" gridcol="8" gridrow="2" pageId="4" pageNumber="567">7.2</td>
<td id="76895DEAFFAD6AB7FC50FC02F828403C" box="[997,1066,999,1024]" gridcol="9" gridrow="2" pageId="4" pageNumber="567">0.9</td>
<td id="76895DEAFFAD6AB7FB8CFC02F864403C" box="[1081,1126,999,1024]" gridcol="10" gridrow="2" pageId="4" pageNumber="567">10.5</td>
<td id="76895DEAFFAD6AB7FBC7FC02F8A2403C" box="[1138,1184,999,1024]" gridcol="11" gridrow="2" pageId="4" pageNumber="567">25.7</td>
<td id="76895DEAFFAD6AB7FB1BFC02F8D9403C" box="[1198,1243,999,1024]" gridcol="12" gridrow="2" pageId="4" pageNumber="567">19.4</td>
<td id="76895DEAFFAD6AB7FB52FC02F925403C" box="[1255,1319,999,1024]" gridcol="13" gridrow="2" pageId="4" pageNumber="567">35.4</td>
<td id="76895DEAFFAD6AB7FA83FC02F966403C" box="[1334,1380,999,1024]" gridcol="14" gridrow="2" pageId="4" pageNumber="567">47.8</td>
<td id="76895DEAFFAD6AB7FADAFC02F9A3403C" box="[1391,1441,999,1024]" gridcol="15" gridrow="2" pageId="4" pageNumber="567">8.3</td>
<td id="76895DEAFFAD6AB7FA1BFC02F9E7403C" box="[1454,1509,999,1024]" gridcol="16" gridrow="2" pageId="4" pageNumber="567">1.5</td>
<td id="76895DEAFFAD6AB7FA46FC02FA1E403C" box="[1523,1564,999,1024]" gridcol="17" gridrow="2" pageId="4" pageNumber="567">10</td>
<td id="76895DEAFFAD6AB7F99CFC02FA65403C" box="[1577,1639,999,1024]" gridcol="18" gridrow="2" pageId="4" pageNumber="567">26/25</td>
<td id="76895DEAFFAD6AB7F9C0FC02FAB2403C" box="[1653,1712,999,1024]" gridcol="19" gridrow="2" pageId="4" pageNumber="567">22/23</td>
<td id="76895DEAFFAD6AB7F908FC02FB12403C" box="[1725,1808,999,1024]" gridcol="20" gridrow="2" pageId="4" pageNumber="567">7</td>
<td id="76895DEAFFAD6AB7F894FC02FB77403C" box="[1825,1909,999,1024]" gridcol="21" gridrow="2" pageId="4" pageNumber="567">8</td>
</tr>
<tr id="35583496FFAD6AB7FF26FBC4FB774006" box="[147,1909,1057,1082]" gridrow="3" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FBC4FD4E4006" box="[147,332,1057,1082]" gridcol="0" gridrow="3" pageId="4" pageNumber="567">UADBA-R 70492</th>
<td id="76895DEAFFAD6AB7FEDDFBC4FDF24006" box="[360,496,1057,1082]" gridcol="1" gridrow="3" pageId="4" pageNumber="567">APR 12691</td>
<td id="76895DEAFFAD6AB7FDB2FBC4FE644006" box="[519,614,1057,1082]" gridcol="2" gridrow="3" pageId="4" pageNumber="567">M (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFBC4FEAA4006" box="[632,680,1057,1082]" gridcol="3" gridrow="3" pageId="4" pageNumber="567">82.5</td>
<td id="76895DEAFFAD6AB7FD03FBC4FEE14006" box="[694,739,1057,1082]" gridcol="4" gridrow="3" pageId="4" pageNumber="567">53.8</td>
<td id="76895DEAFFAD6AB7FD5BFBC4FF2A4006" box="[750,808,1057,1082]" gridcol="5" gridrow="3" pageId="4" pageNumber="567">14.2</td>
<td id="76895DEAFFAD6AB7FC82FBC4FF674006" box="[823,869,1057,1082]" gridcol="6" gridrow="3" pageId="4" pageNumber="567">21.4</td>
<td id="76895DEAFFAD6AB7FCC7FBC4FFAB4006" box="[882,937,1057,1082]" gridcol="7" gridrow="3" pageId="4" pageNumber="567">40.5</td>
<td id="76895DEAFFAD6AB7FC02FBC4FFDB4006" box="[951,985,1057,1082]" gridcol="8" gridrow="3" pageId="4" pageNumber="567">6.8</td>
<td id="76895DEAFFAD6AB7FC50FBC4F8284006" box="[997,1066,1057,1082]" gridcol="9" gridrow="3" pageId="4" pageNumber="567">1.2</td>
<td id="76895DEAFFAD6AB7FB8CFBC4F8644006" box="[1081,1126,1057,1082]" gridcol="10" gridrow="3" pageId="4" pageNumber="567">9.5</td>
<td id="76895DEAFFAD6AB7FBC7FBC4F8A24006" box="[1138,1184,1057,1082]" gridcol="11" gridrow="3" pageId="4" pageNumber="567">22.5</td>
<td id="76895DEAFFAD6AB7FB1BFBC4F8D94006" box="[1198,1243,1057,1082]" gridcol="12" gridrow="3" pageId="4" pageNumber="567">17.7</td>
<td id="76895DEAFFAD6AB7FB52FBC4F9254006" box="[1255,1319,1057,1082]" gridcol="13" gridrow="3" pageId="4" pageNumber="567">34.5</td>
<td id="76895DEAFFAD6AB7FA83FBC4F9664006" box="[1334,1380,1057,1082]" gridcol="14" gridrow="3" pageId="4" pageNumber="567">42.9</td>
<td id="76895DEAFFAD6AB7FADAFBC4F9A34006" box="[1391,1441,1057,1082]" gridcol="15" gridrow="3" pageId="4" pageNumber="567">7.3</td>
<td id="76895DEAFFAD6AB7FA1BFBC4F9E74006" box="[1454,1509,1057,1082]" gridcol="16" gridrow="3" pageId="4" pageNumber="567">1.1</td>
<td id="76895DEAFFAD6AB7FA46FBC4FA1E4006" box="[1523,1564,1057,1082]" gridcol="17" gridrow="3" pageId="4" pageNumber="567">9</td>
<td id="76895DEAFFAD6AB7F99CFBC4FA654006" box="[1577,1639,1057,1082]" gridcol="18" gridrow="3" pageId="4" pageNumber="567">26/29</td>
<td id="76895DEAFFAD6AB7F9C0FBC4FAB24006" box="[1653,1712,1057,1082]" gridcol="19" gridrow="3" pageId="4" pageNumber="567">24/24</td>
<td id="76895DEAFFAD6AB7F908FBC4FB124006" box="[1725,1808,1057,1082]" gridcol="20" gridrow="3" pageId="4" pageNumber="567">8</td>
<td id="76895DEAFFAD6AB7F894FBC4FB774006" box="[1825,1909,1057,1082]" gridcol="21" gridrow="3" pageId="4" pageNumber="567">8</td>
</tr>
<tr id="35583496FFAD6AB7FF26FBBFFB77404F" box="[147,1909,1114,1139]" gridrow="4" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FBBFFD4E404F" box="[147,332,1114,1139]" gridcol="0" gridrow="4" pageId="4" pageNumber="567">UADBA-R 70490</th>
<td id="76895DEAFFAD6AB7FEDDFBBFFDF2404F" box="[360,496,1114,1139]" gridcol="1" gridrow="4" pageId="4" pageNumber="567">APR 12590</td>
<td id="76895DEAFFAD6AB7FDB2FBBFFE64404F" box="[519,614,1114,1139]" gridcol="2" gridrow="4" pageId="4" pageNumber="567">M (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFBBFFEAA404F" box="[632,680,1114,1139]" gridcol="3" gridrow="4" pageId="4" pageNumber="567">94.7</td>
<td id="76895DEAFFAD6AB7FD03FBBFFEE1404F" box="[694,739,1114,1139]" gridcol="4" gridrow="4" pageId="4" pageNumber="567">56.4</td>
<td id="76895DEAFFAD6AB7FD5BFBBFFF2A404F" box="[750,808,1114,1139]" gridcol="5" gridrow="4" pageId="4" pageNumber="567">15.3</td>
<td id="76895DEAFFAD6AB7FC82FBBFFF67404F" box="[823,869,1114,1139]" gridcol="6" gridrow="4" pageId="4" pageNumber="567">22.1</td>
<td id="76895DEAFFAD6AB7FCC7FBBFFFAB404F" box="[882,937,1114,1139]" gridcol="7" gridrow="4" pageId="4" pageNumber="567">42</td>
<td id="76895DEAFFAD6AB7FC02FBBFFFDB404F" box="[951,985,1114,1139]" gridcol="8" gridrow="4" pageId="4" pageNumber="567">6.6</td>
<td id="76895DEAFFAD6AB7FC50FBBFF828404F" box="[997,1066,1114,1139]" gridcol="9" gridrow="4" pageId="4" pageNumber="567">1.3</td>
<td id="76895DEAFFAD6AB7FB8CFBBFF864404F" box="[1081,1126,1114,1139]" gridcol="10" gridrow="4" pageId="4" pageNumber="567">9.4</td>
<td id="76895DEAFFAD6AB7FBC7FBBFF8A2404F" box="[1138,1184,1114,1139]" gridcol="11" gridrow="4" pageId="4" pageNumber="567">23.2</td>
<td id="76895DEAFFAD6AB7FB1BFBBFF8D9404F" box="[1198,1243,1114,1139]" gridcol="12" gridrow="4" pageId="4" pageNumber="567">18.1</td>
<td id="76895DEAFFAD6AB7FB52FBBFF925404F" box="[1255,1319,1114,1139]" gridcol="13" gridrow="4" pageId="4" pageNumber="567">33.0</td>
<td id="76895DEAFFAD6AB7FA83FBBFF966404F" box="[1334,1380,1114,1139]" gridcol="14" gridrow="4" pageId="4" pageNumber="567">46.6</td>
<td id="76895DEAFFAD6AB7FADAFBBFF9A3404F" box="[1391,1441,1114,1139]" gridcol="15" gridrow="4" pageId="4" pageNumber="567">9.6</td>
<td id="76895DEAFFAD6AB7FA1BFBBFF9E7404F" box="[1454,1509,1114,1139]" gridcol="16" gridrow="4" pageId="4" pageNumber="567">1.7</td>
<td id="76895DEAFFAD6AB7FA46FBBFFA1E404F" box="[1523,1564,1114,1139]" gridcol="17" gridrow="4" pageId="4" pageNumber="567">10</td>
<td id="76895DEAFFAD6AB7F99CFBBFFA65404F" box="[1577,1639,1114,1139]" gridcol="18" gridrow="4" pageId="4" pageNumber="567">25/27</td>
<td id="76895DEAFFAD6AB7F9C0FBBFFAB2404F" box="[1653,1712,1114,1139]" gridcol="19" gridrow="4" pageId="4" pageNumber="567">21/21</td>
<td id="76895DEAFFAD6AB7F908FBBFFB12404F" box="[1725,1808,1114,1139]" gridcol="20" gridrow="4" pageId="4" pageNumber="567">8</td>
<td id="76895DEAFFAD6AB7F894FBBFFB77404F" box="[1825,1909,1114,1139]" gridcol="21" gridrow="4" pageId="4" pageNumber="567">8</td>
</tr>
<tr id="35583496FFAD6AB7FF26FB71FB774091" box="[147,1909,1172,1197]" gridrow="5" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FB71FD4E4091" box="[147,332,1172,1197]" gridcol="0" gridrow="5" pageId="4" pageNumber="567">UADBA-R 70491</th>
<td id="76895DEAFFAD6AB7FEDDFB71FDF24091" box="[360,496,1172,1197]" gridcol="1" gridrow="5" pageId="4" pageNumber="567">APR 12589</td>
<td id="76895DEAFFAD6AB7FDB2FB71FE644091" box="[519,614,1172,1197]" gridcol="2" gridrow="5" pageId="4" pageNumber="567">F (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFB71FEAA4091" box="[632,680,1172,1197]" gridcol="3" gridrow="5" pageId="4" pageNumber="567">93.7</td>
<td id="76895DEAFFAD6AB7FD03FB71FEE14091" box="[694,739,1172,1197]" gridcol="4" gridrow="5" pageId="4" pageNumber="567">51.4</td>
<td id="76895DEAFFAD6AB7FD5BFB71FF2A4091" box="[750,808,1172,1197]" gridcol="5" gridrow="5" pageId="4" pageNumber="567">16.8</td>
<td id="76895DEAFFAD6AB7FC82FB71FF674091" box="[823,869,1172,1197]" gridcol="6" gridrow="5" pageId="4" pageNumber="567">23.4</td>
<td id="76895DEAFFAD6AB7FCC7FB71FFAB4091" box="[882,937,1172,1197]" gridcol="7" gridrow="5" pageId="4" pageNumber="567">50.5</td>
<td id="76895DEAFFAD6AB7FC02FB71FFDB4091" box="[951,985,1172,1197]" gridcol="8" gridrow="5" pageId="4" pageNumber="567">6.7</td>
<td id="76895DEAFFAD6AB7FC50FB71F8284091" box="[997,1066,1172,1197]" gridcol="9" gridrow="5" pageId="4" pageNumber="567">1.6</td>
<td id="76895DEAFFAD6AB7FB8CFB71F8644091" box="[1081,1126,1172,1197]" gridcol="10" gridrow="5" pageId="4" pageNumber="567">10.2</td>
<td id="76895DEAFFAD6AB7FBC7FB71F8A24091" box="[1138,1184,1172,1197]" gridcol="11" gridrow="5" pageId="4" pageNumber="567">24.9</td>
<td id="76895DEAFFAD6AB7FB1BFB71F8D94091" box="[1198,1243,1172,1197]" gridcol="12" gridrow="5" pageId="4" pageNumber="567">19.2</td>
<td id="76895DEAFFAD6AB7FB52FB71F9254091" box="[1255,1319,1172,1197]" gridcol="13" gridrow="5" pageId="4" pageNumber="567">32.2</td>
<td id="76895DEAFFAD6AB7FA83FB71F9664091" box="[1334,1380,1172,1197]" gridcol="14" gridrow="5" pageId="4" pageNumber="567">46.2</td>
<td id="76895DEAFFAD6AB7FADAFB71F9A34091" box="[1391,1441,1172,1197]" gridcol="15" gridrow="5" pageId="4" pageNumber="567">13.2</td>
<td id="76895DEAFFAD6AB7FA1BFB71F9E74091" box="[1454,1509,1172,1197]" gridcol="16" gridrow="5" pageId="4" pageNumber="567">1.7</td>
<td id="76895DEAFFAD6AB7FA46FB71FA1E4091" box="[1523,1564,1172,1197]" gridcol="17" gridrow="5" pageId="4" pageNumber="567">9</td>
<td id="76895DEAFFAD6AB7F99CFB71FA654091" box="[1577,1639,1172,1197]" gridcol="18" gridrow="5" pageId="4" pageNumber="567">25/26</td>
<td id="76895DEAFFAD6AB7F9C0FB71FAB24091" box="[1653,1712,1172,1197]" gridcol="19" gridrow="5" pageId="4" pageNumber="567">21/21</td>
<td id="76895DEAFFAD6AB7F908FB71FB124091" box="[1725,1808,1172,1197]" gridcol="20" gridrow="5" pageId="4" pageNumber="567">7</td>
<td id="76895DEAFFAD6AB7F894FB71FB774091" box="[1825,1909,1172,1197]" gridcol="21" gridrow="5" pageId="4" pageNumber="567">8</td>
</tr>
<tr id="35583496FFAD6AB7FF26FB2BFB7740DB" box="[147,1909,1230,1255]" gridrow="6" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FB2BFD4E40DB" box="[147,332,1230,1255]" gridcol="0" gridrow="6" pageId="4" pageNumber="567">UADBA-R 70489</th>
<td id="76895DEAFFAD6AB7FEDDFB2BFDF240DB" box="[360,496,1230,1255]" gridcol="1" gridrow="6" pageId="4" pageNumber="567">APR 12522</td>
<td id="76895DEAFFAD6AB7FDB2FB2BFE6440DB" box="[519,614,1230,1255]" gridcol="2" gridrow="6" pageId="4" pageNumber="567">F (HT)</td>
<td id="76895DEAFFAD6AB7FDCDFB2BFEAA40DB" box="[632,680,1230,1255]" gridcol="3" gridrow="6" pageId="4" pageNumber="567">95.3</td>
<td id="76895DEAFFAD6AB7FD03FB2BFEE140DB" box="[694,739,1230,1255]" gridcol="4" gridrow="6" pageId="4" pageNumber="567">50.9</td>
<td id="76895DEAFFAD6AB7FD5BFB2BFF2A40DB" box="[750,808,1230,1255]" gridcol="5" gridrow="6" pageId="4" pageNumber="567">17.0</td>
<td id="76895DEAFFAD6AB7FC82FB2BFF6740DB" box="[823,869,1230,1255]" gridcol="6" gridrow="6" pageId="4" pageNumber="567">25.3</td>
<td id="76895DEAFFAD6AB7FCC7FB2BFFAB40DB" box="[882,937,1230,1255]" gridcol="7" gridrow="6" pageId="4" pageNumber="567">50.2</td>
<td id="76895DEAFFAD6AB7FC02FB2BFFDB40DB" box="[951,985,1230,1255]" gridcol="8" gridrow="6" pageId="4" pageNumber="567">6.5</td>
<td id="76895DEAFFAD6AB7FC50FB2BF82840DB" box="[997,1066,1230,1255]" gridcol="9" gridrow="6" pageId="4" pageNumber="567">1.1</td>
<td id="76895DEAFFAD6AB7FB8CFB2BF86440DB" box="[1081,1126,1230,1255]" gridcol="10" gridrow="6" pageId="4" pageNumber="567">10.8</td>
<td id="76895DEAFFAD6AB7FBC7FB2BF8A240DB" box="[1138,1184,1230,1255]" gridcol="11" gridrow="6" pageId="4" pageNumber="567">26.1</td>
<td id="76895DEAFFAD6AB7FB1BFB2BF8D940DB" box="[1198,1243,1230,1255]" gridcol="12" gridrow="6" pageId="4" pageNumber="567">20.2</td>
<td id="76895DEAFFAD6AB7FB52FB2BF92540DB" box="[1255,1319,1230,1255]" gridcol="13" gridrow="6" pageId="4" pageNumber="567">33.4</td>
<td id="76895DEAFFAD6AB7FA83FB2BF96640DB" box="[1334,1380,1230,1255]" gridcol="14" gridrow="6" pageId="4" pageNumber="567">41.4</td>
<td id="76895DEAFFAD6AB7FADAFB2BF9A340DB" box="[1391,1441,1230,1255]" gridcol="15" gridrow="6" pageId="4" pageNumber="567">9.9</td>
<td id="76895DEAFFAD6AB7FA1BFB2BF9E740DB" box="[1454,1509,1230,1255]" gridcol="16" gridrow="6" pageId="4" pageNumber="567">0.7</td>
<td id="76895DEAFFAD6AB7FA46FB2BFA1E40DB" box="[1523,1564,1230,1255]" gridcol="17" gridrow="6" pageId="4" pageNumber="567">9</td>
<td id="76895DEAFFAD6AB7F99CFB2BFA6540DB" box="[1577,1639,1230,1255]" gridcol="18" gridrow="6" pageId="4" pageNumber="567">25/26</td>
<td id="76895DEAFFAD6AB7F9C0FB2BFAB240DB" box="[1653,1712,1230,1255]" gridcol="19" gridrow="6" pageId="4" pageNumber="567">25/25</td>
<td id="76895DEAFFAD6AB7F908FB2BFB1240DB" box="[1725,1808,1230,1255]" gridcol="20" gridrow="6" pageId="4" pageNumber="567">7</td>
<td id="76895DEAFFAD6AB7F894FB2BFB7740DB" box="[1825,1909,1230,1255]" gridcol="21" gridrow="6" pageId="4" pageNumber="567">8</td>
</tr>
<tr id="35583496FFAD6AB7FF26FAE2FB77411C" box="[147,1909,1287,1312]" gridrow="7" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FAE2FD4E411C" box="[147,332,1287,1312]" gridcol="0" gridrow="7" pageId="4" pageNumber="567">UADBA-R 70494</th>
<td id="76895DEAFFAD6AB7FEDDFAE2FDF2411C" box="[360,496,1287,1312]" gridcol="1" gridrow="7" pageId="4" pageNumber="567">APR 12692</td>
<td id="76895DEAFFAD6AB7FDB2FAE2FE64411C" box="[519,614,1287,1312]" gridcol="2" gridrow="7" pageId="4" pageNumber="567">j F (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFAE2FEAA411C" box="[632,680,1287,1312]" gridcol="3" gridrow="7" pageId="4" pageNumber="567">47.1</td>
<td id="76895DEAFFAD6AB7FD03FAE2FEE1411C" box="[694,739,1287,1312]" gridcol="4" gridrow="7" pageId="4" pageNumber="567">25.7</td>
<td id="76895DEAFFAD6AB7FD5BFAE2FF2A411C" box="[750,808,1287,1312]" gridcol="5" gridrow="7" pageId="4" pageNumber="567">7.6</td>
<td id="76895DEAFFAD6AB7FC82FAE2FF67411C" box="[823,869,1287,1312]" gridcol="6" gridrow="7" pageId="4" pageNumber="567">12.1</td>
<td id="76895DEAFFAD6AB7FCC7FAE2FFAB411C" box="[882,937,1287,1312]" gridcol="7" gridrow="7" pageId="4" pageNumber="567">26.6</td>
<td id="76895DEAFFAD6AB7FC02FAE2FFDB411C" box="[951,985,1287,1312]" gridcol="8" gridrow="7" pageId="4" pageNumber="567">4.4</td>
<td id="76895DEAFFAD6AB7FC50FAE2F828411C" box="[997,1066,1287,1312]" gridcol="9" gridrow="7" pageId="4" pageNumber="567">0.6</td>
<td id="76895DEAFFAD6AB7FB8CFAE2F864411C" box="[1081,1126,1287,1312]" gridcol="10" gridrow="7" pageId="4" pageNumber="567">5.5</td>
<td id="76895DEAFFAD6AB7FBC7FAE2F8A2411C" box="[1138,1184,1287,1312]" gridcol="11" gridrow="7" pageId="4" pageNumber="567">13.6</td>
<td id="76895DEAFFAD6AB7FB1BFAE2F8D9411C" box="[1198,1243,1287,1312]" gridcol="12" gridrow="7" pageId="4" pageNumber="567">10.3</td>
<td id="76895DEAFFAD6AB7FB52FAE2F925411C" box="[1255,1319,1287,1312]" gridcol="13" gridrow="7" pageId="4" pageNumber="567">17.7</td>
<td id="76895DEAFFAD6AB7FA83FAE2F966411C" box="[1334,1380,1287,1312]" gridcol="14" gridrow="7" pageId="4" pageNumber="567">26.8</td>
<td id="76895DEAFFAD6AB7FADAFAE2F9A3411C" box="[1391,1441,1287,1312]" gridcol="15" gridrow="7" pageId="4" pageNumber="567">nm</td>
<td id="76895DEAFFAD6AB7FA1BFAE2F9E7411C" box="[1454,1509,1287,1312]" gridcol="16" gridrow="7" pageId="4" pageNumber="567">nm</td>
<td id="76895DEAFFAD6AB7FA46FAE2FA1E411C" box="[1523,1564,1287,1312]" gridcol="17" gridrow="7" pageId="4" pageNumber="567">10</td>
<td id="76895DEAFFAD6AB7F99CFAE2FA65411C" box="[1577,1639,1287,1312]" gridcol="18" gridrow="7" pageId="4" pageNumber="567">26/27</td>
<td id="76895DEAFFAD6AB7F9C0FAE2FAB2411C" box="[1653,1712,1287,1312]" gridcol="19" gridrow="7" pageId="4" pageNumber="567">23/21</td>
<td id="76895DEAFFAD6AB7F908FAE2FB12411C" box="[1725,1808,1287,1312]" gridcol="20" gridrow="7" pageId="4" pageNumber="567">6</td>
<td id="76895DEAFFAD6AB7F894FAE2FB77411C" box="[1825,1909,1287,1312]" gridcol="21" gridrow="7" pageId="4" pageNumber="567">7</td>
</tr>
<tr id="35583496FFAD6AB7FF26FAA4FB774166" box="[147,1909,1345,1370]" gridrow="8" pageId="4" pageNumber="567">
<th id="76895DEAFFAD6AB7FF26FAA4FD4E4166" box="[147,332,1345,1370]" gridcol="0" gridrow="8" pageId="4" pageNumber="567">UADBA-R 70501</th>
<td id="76895DEAFFAD6AB7FEDDFAA4FDF24166" box="[360,496,1345,1370]" gridcol="1" gridrow="8" pageId="4" pageNumber="567">MSZC 0 253</td>
<td id="76895DEAFFAD6AB7FDB2FAA4FE644166" box="[519,614,1345,1370]" gridcol="2" gridrow="8" pageId="4" pageNumber="567">sa F (PT)</td>
<td id="76895DEAFFAD6AB7FDCDFAA4FEAA4166" box="[632,680,1345,1370]" gridcol="3" gridrow="8" pageId="4" pageNumber="567">59.5</td>
<td id="76895DEAFFAD6AB7FD03FAA4FEE14166" box="[694,739,1345,1370]" gridcol="4" gridrow="8" pageId="4" pageNumber="567">38.2</td>
<td id="76895DEAFFAD6AB7FD5BFAA4FF2A4166" box="[750,808,1345,1370]" gridcol="5" gridrow="8" pageId="4" pageNumber="567">9.8</td>
<td id="76895DEAFFAD6AB7FC82FAA4FF674166" box="[823,869,1345,1370]" gridcol="6" gridrow="8" pageId="4" pageNumber="567">16.2</td>
<td id="76895DEAFFAD6AB7FCC7FAA4FFAB4166" box="[882,937,1345,1370]" gridcol="7" gridrow="8" pageId="4" pageNumber="567">27.6</td>
<td id="76895DEAFFAD6AB7FC02FAA4FFDB4166" box="[951,985,1345,1370]" gridcol="8" gridrow="8" pageId="4" pageNumber="567">5.8</td>
<td id="76895DEAFFAD6AB7FC50FAA4F8284166" box="[997,1066,1345,1370]" gridcol="9" gridrow="8" pageId="4" pageNumber="567">0.8</td>
<td id="76895DEAFFAD6AB7FB8CFAA4F8644166" box="[1081,1126,1345,1370]" gridcol="10" gridrow="8" pageId="4" pageNumber="567">7.5</td>
<td id="76895DEAFFAD6AB7FBC7FAA4F8A24166" box="[1138,1184,1345,1370]" gridcol="11" gridrow="8" pageId="4" pageNumber="567">18.1</td>
<td id="76895DEAFFAD6AB7FB1BFAA4F8D94166" box="[1198,1243,1345,1370]" gridcol="12" gridrow="8" pageId="4" pageNumber="567">13.7</td>
<td id="76895DEAFFAD6AB7FB52FAA4F9254166" box="[1255,1319,1345,1370]" gridcol="13" gridrow="8" pageId="4" pageNumber="567">23.9</td>
<td id="76895DEAFFAD6AB7FA83FAA4F9664166" box="[1334,1380,1345,1370]" gridcol="14" gridrow="8" pageId="4" pageNumber="567">35.0</td>
<td id="76895DEAFFAD6AB7FADAFAA4F9A34166" box="[1391,1441,1345,1370]" gridcol="15" gridrow="8" pageId="4" pageNumber="567">nm</td>
<td id="76895DEAFFAD6AB7FA1BFAA4F9E74166" box="[1454,1509,1345,1370]" gridcol="16" gridrow="8" pageId="4" pageNumber="567">nm</td>
<td id="76895DEAFFAD6AB7FA46FAA4FA1E4166" box="[1523,1564,1345,1370]" gridcol="17" gridrow="8" pageId="4" pageNumber="567">10</td>
<td id="76895DEAFFAD6AB7F99CFAA4FA654166" box="[1577,1639,1345,1370]" gridcol="18" gridrow="8" pageId="4" pageNumber="567">29/29</td>
<td id="76895DEAFFAD6AB7F9C0FAA4FAB24166" box="[1653,1712,1345,1370]" gridcol="19" gridrow="8" pageId="4" pageNumber="567">25/21</td>
<td id="76895DEAFFAD6AB7F908FAA4FB124166" box="[1725,1808,1345,1370]" gridcol="20" gridrow="8" pageId="4" pageNumber="567">7</td>
<td id="76895DEAFFAD6AB7F894FAA4FB774166" box="[1825,1909,1345,1370]" gridcol="21" gridrow="8" pageId="4" pageNumber="567">8</td>
</tr>
</table>
</paragraph>
<caption id="DF17665CFFAC954DFF22F898F83543D3" httpUri="https://zenodo.org/record/2618343/files/figure.png" pageId="5" pageNumber="568" startId="5.[151,250,1917,1939]" targetBox="[228,1370,195,1901]" targetPageId="5">
<paragraph id="8BD736D4FFAC954DFF22F898F83543D3" blockId="5.[151,1437,1917,2031]" pageId="5" pageNumber="568">
<emphasis id="B91CEAC6FFAC954DFF22F898FD1743AE" bold="true" box="[151,277,1917,1939]" pageId="5" pageNumber="568">FIGURE 1.</emphasis>
(a) Bayesian inference phylogenetic tree of the
<taxonomicName id="4C684D57FFAC954DFCBFF898FFCA43AF" baseAuthorityName="Boettger" baseAuthorityYear="1879" box="[778,968,1917,1939]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="568" phylum="Chordata" rank="species" species="ebenaui">
<emphasis id="B91CEAC6FFAC954DFCBFF898FFCA43AF" box="[778,968,1917,1939]" italics="true" pageId="5" pageNumber="568">Uroplatus ebenaui</emphasis>
</taxonomicName>
group based on a 2326 bp alignment of four concatenated mitochondrial gene fragments (ND4, 12S rRNA, 16S rRNA, COX1). Only posterior probability values of 0.94 or greater are displayed at nodes. An outgroup (
<taxonomicName id="4C684D57FFAC954DFDD7F859FED743ED" authorityName="Mocquard" authorityYear="1894" box="[610,725,1979,2001]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="568" phylum="Chordata" rank="species" species="alluaudi">
<emphasis id="B91CEAC6FFAC954DFDD7F859FED743ED" box="[610,725,1979,2001]" italics="true" pageId="5" pageNumber="568">U. alluaudi</emphasis>
</taxonomicName>
) was used for rooting but omitted from the tree for graphical reasons. (b) Haplotype network independently calculated from sequences of the nuclear gene c-mos.
</paragraph>
</caption>
<caption id="DF17665CFFAF954EFF22F9B2F81C42B0" httpUri="https://zenodo.org/record/2618345/files/figure.png" pageId="6" pageNumber="569" startId="6.[151,250,1623,1645]" targetBox="[154,1433,195,1601]" targetPageId="6">
<paragraph id="8BD736D4FFAF954EFF22F9B2F81C42B0" blockId="6.[151,1436,1623,1676]" pageId="6" pageNumber="569">
<emphasis id="B91CEAC6FFAF954EFF22F9B2FD174250" bold="true" box="[151,277,1623,1645]" pageId="6" pageNumber="569">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="4C684D57FFAF954EFE16F9B2FE6A4251" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[419,616,1623,1645]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="569" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAF954EFE16F9B2FE6A4251" box="[419,616,1623,1645]" italics="true" pageId="6" pageNumber="569">Uroplatus finaritra</emphasis>
</taxonomicName>
<emphasis id="B91CEAC6FFAF954EFDC5F9BDFEC14251" bold="true" box="[624,707,1624,1645]" pageId="6" pageNumber="569">
<taxonomicNameLabel id="A22F57BDFFAF954EFDC5F9BDFEC14251" box="[624,707,1624,1645]" pageId="6" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UADBA-R 70489, APR 12522) in dorsal view (left) and a paratype (UADBA-R 70491, APR 12589) in ventral view (right) view. Scale bar indicates 10 mm.
</paragraph>
</caption>
<subSubSection id="C372655FFFAF9541FF72F95CF9304521" lastPageId="9" lastPageNumber="572" pageId="6" pageNumber="569" type="description">
<paragraph id="8BD736D4FFAF954FFF72F95CFFF44559" blockId="6.[151,1436,1720,2033]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="570" pageId="6" pageNumber="569">
<emphasis id="B91CEAC6FFAF954EFF72F95CFE1342ED" bold="true" box="[199,529,1720,1745]" pageId="6" pageNumber="569">
<emphasis id="B91CEAC6FFAF954EFF72F95CFE0842ED" bold="true" box="[199,522,1720,1745]" italics="true" pageId="6" pageNumber="569">
Description of the
<typeStatus id="54D38876FFAF954EFE1DF95DFE0842ED" box="[424,522,1720,1745]" pageId="6" pageNumber="569" type="holotype">holotype</typeStatus>
</emphasis>
.
</emphasis>
Adult female in fair condition, with a fully intact tail, both of its hind limbs fractured (
<figureCitation id="13532A51FFAF954EFEBEF939FD5242C9" box="[267,336,1756,1781]" captionStart="FIGURE 2" captionStartId="6.[151,250,1623,1645]" captionTargetBox="[154,1433,195,1601]" captionTargetId="figure@6.[151,1436,193,1602]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Uroplatus finaritra sp. nov. (UADBA-R 70489, APR 12522) in dorsal view (left) and a paratype (UADBA-R 70491, APR 12589) in ventral view (right) view. Scale bar indicates 10 mm." httpUri="https://zenodo.org/record/2618345/files/figure.png" pageId="6" pageNumber="569">Fig. 2</figureCitation>
). SVL
<quantity id="4C909B31FFAF954EFE15F939FE0442C9" box="[416,518,1756,1781]" metricMagnitude="-2" metricUnit="m" metricValue="9.53" pageId="6" pageNumber="569" unit="mm" value="95.3">95.3 mm</quantity>
, tail length
<quantity id="4C909B31FFAF954EFD3CF939FEED42C9" box="[649,751,1756,1781]" metricMagnitude="-2" metricUnit="m" metricValue="5.09" pageId="6" pageNumber="569" unit="mm" value="50.9">50.9 mm</quantity>
, maximum tail width 17.0 mm; for further measurements see Table 1. Head triangular in dorsal view; canthus rostralis indistinct and concave; snout sloping weakly downwards anteriorly; snout weakly depressed, fairly long (1.7 times longer than eye diameter); eyes large (eye diameter
<quantity id="4C909B31FFAF954EFACCF8C0FCC5435D" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="569" unit="mm" value="6.5">6.5 mm</quantity>
), bulging slightly above dorsal surface of cranium, directed laterally, pupil vertical with crenate borders; ear opening very small (horizontal diameter
<quantity id="4C909B31FFAF954EFDEAF888FEBB43B9" box="[607,697,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="569" unit="mm" value="1.1">1.1 mm</quantity>
), its opening facing posterolaterally, but also posteroventrally (ear opening clearly visible in ventral view but not in dorsal view); nostrils laterally oriented; body somewhat laterally compressed, without lateral dermal fringes; limbs without fringes and very few, miniscule spines on the hind limbs (one on the knee, as well as a small flap of skin), and two miniscule spines on the forelimb (on the elbow and near the wrist); forelimb reaches the canthus rostralis when adpressed forward and midbody when adpressed backwards along body (forelimb length/axillagroin distance 33.4/
<quantity id="4C909B31FFAE954FFCBAFF59FF7744E8" box="[783,885,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.02" pageId="7" pageNumber="570" unit="mm" value="50.2">50.2 mm</quantity>
= 67%), hindlimb reaches ¾ up the body when adpressed forward along body (hindlimb length/axillagroin distance 41.4/
<quantity id="4C909B31FFAE954FFC44FF3AF85544CB" box="[1009,1111,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.02" pageId="7" pageNumber="570" unit="mm" value="50.2">50.2 mm</quantity>
= 83%); original tail length 53% of snoutvent length, membranous borders of the tail symmetrical, without any emarginations, long and leafshaped, with a thin, borderless spatulate tip. Toes bear small claws, with the distal phalange not much wider than the rest of the digit; the third finger bears 7 lamellae, the third toe 8 lamellae.
</paragraph>
<caption id="DF17665CFFAE954FFF22F935FF20435D" httpUri="https://zenodo.org/record/2618347/files/figure.png" pageId="7" pageNumber="570" startId="7.[151,250,1744,1766]" targetBox="[151,1407,409,1723]" targetPageId="7">
<paragraph id="8BD736D4FFAE954FFF22F935FF20435D" blockId="7.[151,1436,1744,1889]" pageId="7" pageNumber="570">
<emphasis id="B91CEAC6FFAE954FFF22F935FD1442D9" bold="true" box="[151,278,1744,1766]" pageId="7" pageNumber="570">FIGURE 3.</emphasis>
Variation among specimens of
<taxonomicName id="4C684D57FFAE954FFDDDF935FF2C42DA" authority="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences, 2019" authorityName="Ratsoavina &amp; Raselimanana &amp; Scherz &amp; Rakotoarison &amp; Razafindraibe &amp; Glaw &amp; Vences" authorityYear="2019" box="[616,814,1744,1766]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="570" phylum="Chordata" rank="species" species="finaritra" status="sp. nov.">
<emphasis id="B91CEAC6FFAE954FFDDDF935FF2C42DA" box="[616,814,1744,1766]" italics="true" pageId="7" pageNumber="570">Uroplatus finaritra</emphasis>
</taxonomicName>
<emphasis id="B91CEAC6FFAE954FFC8DF934FF8E42DA" bold="true" box="[824,908,1745,1766]" pageId="7" pageNumber="570">
<taxonomicNameLabel id="A22F57BDFFAE954FFC8DF934FF8E42DA" box="[824,908,1745,1766]" pageId="7" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Marojejy National Park. (a) photo probably showing paratype UADBA-R 70494 (APR 12692), (b) paratype UADBA-R 70491 (APR 12589), (c) paratype UADBA-R 70490 (APR 12590), (d) holotype UADBA-R 70489 (APR 12522), (e) paratype UADBA-R 70501 (MSZC 0253), (f) paratype ZSM 458/2016 (MSZC 0271). All specimens shown except (c) are females; note the differences in tail notching between this specimen and the others, and the presence of hemipenial bulges.
</paragraph>
</caption>
<paragraph id="8BD736D4FFAE9540FF72F86AFFD845F3" blockId="7.[151,1436,1933,2031]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="571" pageId="7" pageNumber="570">Nares separated from each other by at least 10 small granular scales, from the first supralabial by 3 scales, and from the rostral scale by 5 scales; first supralabial not taller than the others; rostral entire, much wider than tall; mental scale very small, not differentiated from infralabial scales; 25/26 (right/left) supralabials and 25/25 infralabials; no enlarged postmental scales or chin shields; dorsal and ventral scales of head, neck, body, limbs, and tail small, granular, juxtaposed and largely of uniform size, except on the posterior ventral abdomen, where they are slightly larger than the rest of the body, and arranged almost uniformly. Two curved lines (rows of slightly enlarged scales) extending from the posterolateral parts of the head (nuchal region) converge on the neck forming a V-shaped pattern (neck triangular line). A similar, moderately distinct, fairly straight line (also formed by a row of slightly enlarged scales) is present between the eyes and connects the supraocular spines. Another, less distinct line is present connecting the anterior angle of the eyes over the snout in a bowed line. The body possesses very few dermal spines—a prominent pointed supraocular spine, a single spine at the posterior angle of the head, and a small number of spines on each limb (described above). No axillary pits present.</paragraph>
<paragraph id="8BD736D4FFA19540FF72FE39FD4247F4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="571">
<emphasis id="B91CEAC6FFA19540FF72FE39FD5245C8" box="[199,336,476,500]" italics="true" pageId="8" pageNumber="571">Colouration</emphasis>
. After two years of preservation in 70% ethanol the colour is slightly faded (
<figureCitation id="13532A51FFA19540FB79FE39F91645C8" box="[1228,1300,476,501]" captionStart="FIGURE 2" captionStartId="6.[151,250,1623,1645]" captionTargetBox="[154,1433,195,1601]" captionTargetId="figure@6.[151,1436,193,1602]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Uroplatus finaritra sp. nov. (UADBA-R 70489, APR 12522) in dorsal view (left) and a paratype (UADBA-R 70491, APR 12589) in ventral view (right) view. Scale bar indicates 10 mm." httpUri="https://zenodo.org/record/2618345/files/figure.png" pageId="8" pageNumber="571">Fig. 2</figureCitation>
). All dorsal surfaces of the specimen are a ruddy brown, fading to a salmon brown in the neck region and at the insertion of the hind limbs. A fine network of blackish lines is present across the body, with the following notable features: a somewhat straight line connecting the suprocular spines (described above); a v-shaped marking on the neck, which extends into a vertebral stripe that reaches all the way to the tail tip; a number of posteriorly oriented chevrons from the mid-dorsum to the belly, becoming thinner posteroventrally, three of which are most distinct. The exterior of the shank and outermost two toes are grey-black, whereas the inner foot is brown. The dorsal snout is darker than the rest of the head. The tail is not different in colour or pattern from the dorsal body. A distinct lateral line runs along the ventrolateral edges of the abdomen from the insertion of the leg to the axilla. The ventral colouration is uniformly orange-brown, with a faint, symmetrical pattern of dark lines on the chin. The soles of the hands and feet are grey. The ventral tail is the colour of the trunk mottled with grey splotches. A faint cream tear-like marking is present at the posterior corner of the eye above the supralabial. The oral mucosa is dark red, with a pink tongue and cream under the eyes and along the lower jaw. In life, the iris colour was silver externally and rusty around the pupil (
<figureCitation id="13532A51FFA19540FF55FC4AFD3147F4" box="[224,307,943,968]" captionStart="FIGURE 3" captionStartId="7.[151,250,1744,1766]" captionTargetBox="[151,1407,409,1723]" captionTargetId="figure@7.[151,1436,409,1723]" captionTargetPageId="7" captionText="FIGURE 3. Variation among specimens of Uroplatus finaritra sp. nov. from Marojejy National Park. (a) photo probably showing paratype UADBA-R 70494 (APR 12692), (b) paratype UADBA-R 70491 (APR 12589), (c) paratype UADBA-R 70490 (APR 12590), (d) holotype UADBA-R 70489 (APR 12522), (e) paratype UADBA-R 70501 (MSZC 0253), (f) paratype ZSM 458/2016 (MSZC 0271). All specimens shown except (c) are females; note the differences in tail notching between this specimen and the others, and the presence of hemipenial bulges." httpUri="https://zenodo.org/record/2618347/files/figure.png" pageId="8" pageNumber="571">Fig. 3d</figureCitation>
).
</paragraph>
<paragraph id="8BD736D4FFA19540FF72FC30F8084009" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="571">
<emphasis id="B91CEAC6FFA19540FF72FC30FD2D47D0" box="[199,303,981,1004]" italics="true" pageId="8" pageNumber="571">Variation</emphasis>
. Molecular data are also available from a previous paper (
<bibRefCitation id="EFF94B25FFA19540FC72FC31F8CB47D0" author="Raxworthy, C. J. &amp; Pearson, R. G. &amp; Zimkus, B. M. &amp; Reddy, S. &amp; Deo, A. J. &amp; Nussbaum, R. A. &amp; Ingram, C. M." box="[967,1225,980,1005]" pageId="8" pageNumber="571" pagination="423 - 440" refId="ref8784" refString="Raxworthy, C. J., Pearson, R. G., Zimkus, B. M., Reddy, S., Deo, A. J., Nussbaum, R. A. &amp; Ingram, C. M. (2008) Continental speciation in the tropics: contrasting biogeographic patterns of divergence in the Uroplatus leaf-tailed gecko radiation of Madagascar. Journal of Zoology, 275, 423 - 440. https: // doi. org / 10.1111 / j. 1469 - 7998.2008.00460. x" type="journal article" year="2008">
Raxworthy
<emphasis id="B91CEAC6FFA19540FBFBFC30F88547D0" box="[1102,1159,980,1004]" italics="true" pageId="8" pageNumber="571">et al.</emphasis>
2008
</bibRefCitation>
) for one specimen (RAN 42274) from Marojejy as
<taxonomicName id="4C684D57FFA19540FDB4FC1DFE70402C" box="[513,626,1016,1040]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="571" phylum="Chordata" rank="genus">
<emphasis id="B91CEAC6FFA19540FDB4FC1DFE70402C" box="[513,626,1016,1040]" italics="true" pageId="8" pageNumber="571">Uroplatus</emphasis>
</taxonomicName>
sp. G, included for 12S and cytochrome
<emphasis id="B91CEAC6FFA19540FB8DFC1DF844402C" box="[1080,1094,1016,1040]" italics="true" pageId="8" pageNumber="571">b</emphasis>
as
<emphasis id="B91CEAC6FFA19540FBDEFC1CF87D402C" box="[1131,1151,1017,1040]" italics="true" pageId="8" pageNumber="571">U</emphasis>
. sp. Ca
<quantity id="4C909B31FFA19540FB61FC1DF8FC402C" box="[1236,1278,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="571" unit="in" value="9.0">9 in</quantity>
<bibRefCitation id="EFF94B25FFA19540FAB0FC12FD0A4008" author="Ratsoavina, F. M. &amp; Raminosoa, N. R. &amp; Louis, E. E. Jr. &amp; Raselimanana, A. P. &amp; Glaw, F. &amp; Vences, M." pageId="8" pageNumber="571" pagination="115 - 148" refId="ref8519" refString="Ratsoavina, F. M., Raminosoa, N. R., Louis, E. E. Jr., Raselimanana, A. P., Glaw, F. &amp; Vences, M. (2013) An overview of Madagascar's leaf tailed geckos (genus Uroplatus): species boundaries, candidate species and review of geographical distribution based on molecular data. Salamandra, 49, 115 - 148." type="journal article" year="2013">
Ratsoavina
<emphasis id="B91CEAC6FFA19540FA3DFC1CFCB64008" italics="true" pageId="8" pageNumber="571">et al.</emphasis>
(2013)
</bibRefCitation>
. The available sequence for 12S agrees with those of our specimens.
</paragraph>
<paragraph id="8BD736D4FFA19540FF72FBDAFF364169" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="571">
The
<typeStatus id="54D38876FFA19540FEB5FBA5FD694064" box="[256,363,1088,1112]" pageId="8" pageNumber="571" type="paratype">paratypes</typeStatus>
were sexed as adult males by the presence of everted hemipenes (UADBA-R 70493) or presence of distinctly enlarged tail base (UADBA-R 70490 and 70492), and distinctly notched tail edges; or as females based on the absence of hemipenes or hemipenial bulges, presence of visible oviduct or ovary structures after dissection, and generally smooth tail edges. In the smallest specimen (UADBA-R 70494) female inner reproductive organs were observed but were only weakly developed, and we therefore consider this specimen as juvenile. The sexual maturity of two further specimens (UADBA-R 70501 and ZSM 458/2016) is unclear, as they are considerably smaller than the large adult females (UADBA-R 70491 and 70489) but have similar gonad development. We tentatively consider these to be subadults.
</paragraph>
<paragraph id="8BD736D4FFA19540FF72FA84FDD442A4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="571">
In general, the
<typeStatus id="54D38876FFA19540FECDFA84FDE14145" box="[376,483,1377,1401]" pageId="8" pageNumber="571" type="paratype">paratypes</typeStatus>
agree well with the
<typeStatus id="54D38876FFA19540FD63FABAFF3A4144" box="[726,824,1375,1400]" pageId="8" pageNumber="571" type="holotype">holotype</typeStatus>
in morphology. For measurements, see Table 1. The number of head spines varies from 29, and is more in males than in females, as males are more spiny than females in general, and have more heterogeneous scalation. Forelimbs can bear a minimum of two spines, or as many as eight, with a small spine present on the elbow. Hind limbs can bear 215 spines, and the dermal knee flap almost always bears at least a small spine, which is smaller in females than males. In males the tail is moderately emarginated, with semicircular emarginations that tend to be asymmetrical. In females the tail is without emarginations and has smooth edges. Internarial scales range from nine to ten, supralabials from 2529, and infralabials from 2125. The third finger bears 78 lamellae (
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one juvenile specimen, UADBA-R 70494), the third toe bears 78 lamellae.
</paragraph>
<paragraph id="8BD736D4FFA19541FF72F941F9304521" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1436,151,827]" lastPageId="9" lastPageNumber="572" pageId="8" pageNumber="571">
The colouration is varied, as is commonly the case in
<taxonomicName id="4C684D57FFA19540FCF1F941FFB74280" box="[836,949,1700,1724]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="571" phylum="Chordata" rank="genus">
<emphasis id="B91CEAC6FFA19540FCF1F941FFB74280" box="[836,949,1700,1724]" italics="true" pageId="8" pageNumber="571">Uroplatus</emphasis>
</taxonomicName>
species (
<figureCitation id="13532A51FFA19540FB97F941F8914280" box="[1058,1171,1700,1725]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="7.[151,250,1744,1766]" captionStartId-1="11.[151,250,1965,1987]" captionTargetBox-0="[151,1407,409,1723]" captionTargetBox-1="[276,1315,205,1941]" captionTargetId-0="figure@7.[151,1436,409,1723]" captionTargetId-1="figure@11.[268,1322,193,1949]" captionTargetPageId-0="7" captionTargetPageId-1="11" captionText-0="FIGURE 3. Variation among specimens of Uroplatus finaritra sp. nov. from Marojejy National Park. (a) photo probably showing paratype UADBA-R 70494 (APR 12692), (b) paratype UADBA-R 70491 (APR 12589), (c) paratype UADBA-R 70490 (APR 12590), (d) holotype UADBA-R 70489 (APR 12522), (e) paratype UADBA-R 70501 (MSZC 0253), (f) paratype ZSM 458/2016 (MSZC 0271). All specimens shown except (c) are females; note the differences in tail notching between this specimen and the others, and the presence of hemipenial bulges." captionText-1="FIGURE 4. Subadult female specimens of Uroplatus finaritra sp. nov. (ab) ZSM 458/2016 (MSZC 0271) and (cd) UADBA-R 70501 (MSZC 0253) in (a, d) dorsolateral and (bc) dorsal view." httpUri-0="https://zenodo.org/record/2618347/files/figure.png" httpUri-1="https://zenodo.org/record/2618349/files/figure.png" pageId="8" pageNumber="571">Figs. 34</figureCitation>
). The line on the head connecting the supraocular spines is generally bowed, but can be slightly pointed posteriorly, and in UADBA-R 70492, it is irregular at its midpoint. The dark vertebral stripe is only present in the
<typeStatus id="54D38876FFA19540FBE9F909F8C34339" box="[1116,1217,1772,1797]" pageId="8" pageNumber="571" type="holotype">holotype</typeStatus>
, UADBA-R 70491 and 70494, and in other specimens is either absent, or is cream and not dark (in UADBA-R 70492 and 70490). Overall dorsal colouration differs dramatically, from almost completely reddish brown (UADBA-R 70501) to highly mottled greys and blacks (UADBA-R 70493). Irregular flecks can be present or absent on any part of the body. The different colouration of the outer shank and foot is always present, and a cream tear-like marking below the eye is almost always present (absent in UADBA-R 70494 and 70501). The ventrolateral stripe is always present but not always distinct (indistinct in UADBA-R 70493). Ventral colouration is as variable as dorsal colouration. UADBA-R 70491 is extremely similar to the
<typeStatus id="54D38876FFA09541FD2BFF72FF01448C" box="[670,771,151,176]" pageId="9" pageNumber="572" type="holotype">holotype</typeStatus>
, and to a lesser degree 70492, 70494, and 70501 resemble the
<typeStatus id="54D38876FFA09541FF75FF59FD2044E9" box="[192,290,188,213]" pageId="9" pageNumber="572" type="holotype">holotype</typeStatus>
ventrally. UADBA-R 70490 and 70493 differ dramatically in being darker in colour, having irregular flecks of cream and reddish brown, and lacking the symmetrical markings on the chin. In life, the colouration of the iris is typically silvery but can be red (
<figureCitation id="13532A51FFA09541FDF0FEE1FEBA4521" box="[581,696,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,1744,1766]" captionTargetBox="[151,1407,409,1723]" captionTargetId="figure@7.[151,1436,409,1723]" captionTargetPageId="7" captionText="FIGURE 3. Variation among specimens of Uroplatus finaritra sp. nov. from Marojejy National Park. (a) photo probably showing paratype UADBA-R 70494 (APR 12692), (b) paratype UADBA-R 70491 (APR 12589), (c) paratype UADBA-R 70490 (APR 12590), (d) holotype UADBA-R 70489 (APR 12522), (e) paratype UADBA-R 70501 (MSZC 0253), (f) paratype ZSM 458/2016 (MSZC 0271). All specimens shown except (c) are females; note the differences in tail notching between this specimen and the others, and the presence of hemipenial bulges." httpUri="https://zenodo.org/record/2618347/files/figure.png" pageId="9" pageNumber="572">Figs. 3c, f</figureCitation>
), and the oral mucosa was consistently dark red (
<figureCitation id="13532A51FFA09541FB55FEE1F9264521" box="[1248,1316,260,285]" captionStart="FIGURE 5" captionStartId="12.[151,250,1534,1556]" captionTargetBox="[150,1436,483,1512]" captionTargetId="figure@12.[150,1436,483,1512]" captionTargetPageId="12" captionText="FIGURE 5. Oral mucosa colouration differences between (ac) Uroplatus finaritra sp. nov. from Marojejy and (df) U. phantasticus from (d) Fierenana, (e) Ranomafana, and (f) Kianjavato." httpUri="https://zenodo.org/record/2618351/files/figure.png" pageId="9" pageNumber="572">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C372655FFFA09541FF72FEC2FE784707" pageId="9" pageNumber="572" type="distribution">
<paragraph id="8BD736D4FFA09541FF72FEC2FCEA4559" blockId="9.[151,1436,151,827]" pageId="9" pageNumber="572">
<emphasis id="B91CEAC6FFA09541FF72FEC2FD58457C" bold="true" box="[199,346,295,320]" pageId="9" pageNumber="572">
<emphasis id="B91CEAC6FFA09541FF72FEC2FD51457C" bold="true" box="[199,339,295,320]" italics="true" pageId="9" pageNumber="572">Distribution</emphasis>
.
</emphasis>
At present this species has only been collected on the Marojejy Massif, at altitudes from
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a.s.l.
</paragraph>
<paragraph id="8BD736D4FFA09541FF72FE8AF9804601" blockId="9.[151,1436,151,827]" pageId="9" pageNumber="572">
<emphasis id="B91CEAC6FFA09541FF72FE8AFD8745B4" bold="true" box="[199,389,367,392]" italics="true" pageId="9" pageNumber="572">Natural History.</emphasis>
The
<typeStatus id="54D38876FFA09541FE77FE8AFE2645B4" box="[450,548,367,392]" pageId="9" pageNumber="572" type="holotype">holotype</typeStatus>
was collected active at night on a stem
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above the forest floor in nearly closed canopy humid forest at the bottom of a slope, in the vicinity of a
<taxonomicName id="4C684D57FFA09541FC7EFE71F89E4590" baseAuthorityName="Duméril &amp; Bibron" baseAuthorityYear="1836" box="[971,1180,404,428]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="572" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B91CEAC6FFA09541FC7EFE71F89E4590" box="[971,1180,404,428]" italics="true" pageId="9" pageNumber="572">Uroplatus lineatus</emphasis>
</taxonomicName>
specimen. UADBA-R 70490, 70491, and 70493 were collected on small branches and leaves between 1.5 and
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above the ground in similar forest. UADBA-R 70492 and 70494 were also active on branches
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from the forest floor in nearly closed-canopy humid forest with a thick understory and ferns. UADBA-R 70501 and ZSM 458/2016 were collected active at night on thin branches (up to
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above the ground) in closed-canopy, low-altitude rainforest.
</paragraph>
<paragraph id="8BD736D4FFA09541FF72FDADFE784707" blockId="9.[151,1436,151,827]" pageId="9" pageNumber="572">
We consider the various collection localities to represent a single Threat-Defined Location in the sense of the International Union for the Conservation of Nature (
<bibRefCitation id="EFF94B25FFA09541FD6BFD89FF6546B8" author="IUCN" box="[734,871,620,644]" pageId="9" pageNumber="572" refId="ref7953" refString="IUCN. (2012) IUCN Red List Categories and Criteria: Fersion 3.1. IUCN, Gland and Cambridge." type="book" year="2012">IUCN 2012</bibRefCitation>
), as we have recently done for several frog species from low- to mid-elevation forests in Marojejy (
<bibRefCitation id="EFF94B25FFA09541FD75FD6AFF964694" author="Scherz, M. D. &amp; Glaw, F. &amp; Vences, M. &amp; Andreone, F. &amp; Crottini, A." box="[704,916,655,680]" pageId="9" pageNumber="572" pagination="91 - 106" refId="ref9139" refString="Scherz, M. D., Glaw, F., Vences, M., Andreone, F. &amp; Crottini, A. (2016) Two new species of terrestrial microhylid frogs (Microhylidae: Cophylinae: Rhombophryne) from northeastern Madagascar. Salamandra, 52, 91 - 106." type="journal article" year="2016">
Scherz
<emphasis id="B91CEAC6FFA09541FCA3FD74FF534694" box="[790,849,656,680]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="EFF94B25FFA09541FC14FD6AF8C24694" author="Rakotoarison, A. &amp; Scherz, M. D. &amp; Glaw, F. &amp; Kohler, J. &amp; Andreone, F. &amp; Franzen, M. &amp; Glos, J. &amp; Hawlitschek, O. &amp; Jono, T. &amp; Mori, A. &amp; Ndriantsoa, S. H. &amp; Raminosoa Rasoamampionona, N. &amp; Riemann, J. C. &amp; Rodel, M. - O. &amp; Rosa, G. M. &amp; Vieites, D. R. &amp; Crottini, A. &amp; Vences, M." box="[929,1216,655,680]" pageId="9" pageNumber="572" pagination="271 - 398" refId="ref8228" refString="Rakotoarison, A., Scherz, M. D., Glaw, F., Kohler, J., Andreone, F., Franzen, M., Glos, J., Hawlitschek, O., Jono, T., Mori, A., Ndriantsoa, S. H., Raminosoa Rasoamampionona, N., Riemann, J. C., Rodel, M. - O., Rosa, G. M., Vieites, D. R., Crottini, A. &amp; Vences, M. (2017) Describing the smaller majority: integrative taxonomy reveals twenty-six new species of tiny microhylid frogs (genus Stumpffia) from Madagascar. Fertebrate Zoology, 67, 271 - 398." type="journal article" year="2017">
Rakotoarison
<emphasis id="B91CEAC6FFA09541FBF5FD74F8784694" box="[1088,1146,656,680]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
2017
</bibRefCitation>
). In line with these assessments, we recognise that there is on-going habitat degradation due to illegal logging activity, especially at low altitudes in and around Marojejy National Park. The Extent of Occurrence and Area of Occupancy of the new species is certainly greater than
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, but is likely less than
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, and as such we recommend a status of Endangered for it under criterion B1ab(iii).
</paragraph>
</subSubSection>
</treatment>
</document>