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<document id="C500B9674766053F1F46370D41942FCB" ID-DOI="10.11646/zootaxa.5424.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10808003" ID-ZooBank="D3515496-31E1-48A4-B214-1AC668FB38D5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710236539104" checkinUser="plazi" docAuthor="Tan, Ming Kai, Grumo, Kim C., Gono, Alfredo Iii A., Bahoy, Daphne Cayle M., Rivera, Rodrin R., Nuñeza, Olga Macas, Japir, Razy &amp; Chung, Arthur Y. C." docDate="2024" docId="03C15030FFF1FFC943C6FA7EFE392449" docLanguage="en" docName="zootaxa.5424.1.3.pdf" docOrigin="Zootaxa 5424 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Itara (Bornitara) tenompok Tan, Japir &amp; Chung 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="74" masterDocId="FFF82848FFF9FFC44351FFB8FFD9252C" masterDocTitle="New species of Itara (Orthoptera: Grylloidea: Itarinae) and descriptions of calling songs from Mindanao and Sabah" masterLastPageNumber="79" masterPageNumber="61" pageNumber="69" updateTime="1710237287157" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="800FB4D7E89E90A5C4C128918F865E42">New species of Itara (Orthoptera: Grylloidea: Itarinae) and descriptions of calling songs from Mindanao and Sabah</mods:title>
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<mods:namePart id="B780D9683D15704A5C5E2D0066B248DF">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="DC2F545FE727AB5893FBEF93BFC23312">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore.</mods:affiliation>
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<mods:namePart id="059A122B028401258A22CEE33B138E79">Grumo, Kim C.</mods:namePart>
<mods:nameIdentifier id="323E59C75ECDC8E6EB28BE1468A6934C" type="ORCID">0009-0002-1091-9355</mods:nameIdentifier>
<mods:affiliation id="E4FD428BFF416E95368AD0E9E0DC31EE">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines.</mods:affiliation>
<mods:nameIdentifier id="B748B10C72CF3CCEEBDB56AB29081D62" type="email">kim.grumo@g.msuiit.edu.ph</mods:nameIdentifier>
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<mods:namePart id="A015C7803565B5C7E4A1923BAD602B55">Gono, Alfredo Iii A.</mods:namePart>
<mods:nameIdentifier id="9EBC30F0085A5E414946370EB037F476" type="ORCID">0009-0000-4180-5970</mods:nameIdentifier>
<mods:affiliation id="22C4BA89EC46FEFA1A802E08F055C06D">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines.</mods:affiliation>
<mods:nameIdentifier id="C32212FE3434B98B516DE653C0C78608" type="email">alfredoiii.gono@g.msuiit.edu.ph</mods:nameIdentifier>
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<mods:namePart id="0ED29555D575D3BFC55C7B7A62311AE9">Bahoy, Daphne Cayle M.</mods:namePart>
<mods:nameIdentifier id="FE19AB4BE247FBDDA15E7D73CFA73B32" type="ORCID">0009-0006-9609-0312</mods:nameIdentifier>
<mods:affiliation id="0E359FAA0C96E593E251301E0AD17663">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines.</mods:affiliation>
<mods:nameIdentifier id="E9DE0CB46F36F556E0AE11D0EAE9A2D5" type="email">daphnecayle.bahoy@g.msuiit.edu.ph</mods:nameIdentifier>
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<mods:namePart id="2EBD3ACAEFCCEC51A772E56CE392E811">Rivera, Rodrin R.</mods:namePart>
<mods:nameIdentifier id="8B24A6EBFF9451298B390691D8D914BC" type="ORCID">0000-0003-3019-0258</mods:nameIdentifier>
<mods:affiliation id="61902CF7654071C8874539B2C7C2D843">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines.</mods:affiliation>
<mods:nameIdentifier id="63C071C76296952386D7C6EE84D3147E" type="email">rodrin.rivera@nmsc.edu.ph</mods:nameIdentifier>
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<mods:namePart id="2A33F390D0FF155CDBC76D0B0D64C169">Nuñeza, Olga Macas</mods:namePart>
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<mods:affiliation id="D07032655021054E4809C84D7DA73B38">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines.</mods:affiliation>
<mods:nameIdentifier id="431A2D1A1E17C6BE154A44D93C12BA73" type="email">olga.nuneza@g.msuiit.edu.ph</mods:nameIdentifier>
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<mods:namePart id="45C35A280AE53B6976C6CCB39404B6B5">Japir, Razy</mods:namePart>
<mods:affiliation id="F25275E12CE6EE3E5F07608AB425CECF">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
<mods:nameIdentifier id="475182E525F5B8C7A770EFE9C067AEED" type="email">razy.japir@sabah.gov.my</mods:nameIdentifier>
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<mods:namePart id="150CB3E47615B7D1533FC5BF62067B7F">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="5BB1F35E78EA389C33091616EA4CBA15">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
<mods:nameIdentifier id="4D098588D90C250FAEA4CF824676F5B3" type="email">arthur.chung@sabah.gov.my</mods:nameIdentifier>
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<mods:title id="32E0A77C522A7BD5BD1D7B71C54AE140">Zootaxa</mods:title>
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<mods:part id="F6D84D71AB71C484A87B4D7C834F8178">
<mods:date id="4E9E0DD544161E3CEA3A0D0ABBF5294A">2024</mods:date>
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<mods:number id="69BBD6EC63EC73B1ADC2A9F74A6901BA">2024-03-12</mods:number>
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<mods:number id="A55C2B7890442A2F55C7CA9CB1C0C756">5424</mods:number>
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<treatment id="03C15030FFF1FFC943C6FA7EFE392449" LSID="urn:lsid:plazi:treatment:03C15030FFF1FFC943C6FA7EFE392449" httpUri="http://treatment.plazi.org/id/03C15030FFF1FFC943C6FA7EFE392449" lastPageId="13" lastPageNumber="74" pageId="8" pageNumber="69">
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<heading id="D09F564AFFF1FFCC43C6FA7EFC8D20F3" bold="true" box="[151,852,1477,1504]" fontSize="11" level="1" pageId="8" pageNumber="69" reason="1">
<emphasis id="B91C3D34FFF1FFCC43C6FA7EFC8D20F3" bold="true" box="[151,852,1477,1504]" pageId="8" pageNumber="69">
<taxonomicName id="4C689AA5FFF1FFCC43C6FA7EFD3020F3" authority="Tan, Japir &amp; Chung" authorityName="Tan, Japir &amp; Chung" authorityYear="2024" box="[151,745,1477,1504]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="tenompok" status="sp. nov." subGenus="Bornitara">
<emphasis id="B91C3D34FFF1FFCC43C6FA7EFF0A20CC" bold="true" box="[151,211,1478,1504]" italics="true" pageId="8" pageNumber="69">Itara</emphasis>
(
<emphasis id="B91C3D34FFF1FFCC43B4FA7DFE8520F3" bold="true" box="[229,348,1477,1503]" italics="true" pageId="8" pageNumber="69">Bornitara</emphasis>
)
<emphasis id="B91C3D34FFF1FFCC423CFA7DFE3D20CC" bold="true" box="[365,484,1477,1504]" italics="true" pageId="8" pageNumber="69">tenompok</emphasis>
Tan, Japir &amp; Chung
</taxonomicName>
,
<taxonomicNameLabel id="A22F804FFFF1FFCC41A7FA7DFC8D20F3" box="[758,852,1477,1503]" pageId="8" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C372B2ADFFF1FFCC43C6FA51FEFE232F" box="[151,295,1513,1539]" pageId="8" pageNumber="69" type="description">
<paragraph id="8BD7E126FFF1FFCC43C6FA51FEFE232F" blockId="8.[151,852,1477,1539]" box="[151,295,1513,1539]" pageId="8" pageNumber="69">
(
<figureCitation id="1353FDA3FFF1FFCC43CEFA51FEC6232F" box="[159,287,1513,1539]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="12.[151,250,1061,1085]" captionStartId-1="13.[151,250,1932,1956]" captionStartId-2="14.[151,250,1107,1131]" captionTargetBox-0="[151,1435,181,1037]" captionTargetBox-1="[151,1436,381,1909]" captionTargetBox-2="[151,1436,181,1083]" captionTargetId-0="figure-21@12.[151,1435,181,1037]" captionTargetId-1="figure-77@13.[151,1436,381,1909]" captionTargetId-2="figure-21@14.[151,1436,181,1083]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 10. Itara (Bornitara) tenompok sp. nov. ♂ holotype in its natural environment at Tenompok Forest Reserve, Sabah." captionText-1="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." captionText-2="FIGURE 12. Itara (Bornitara) tenompok sp. nov. ♂ holotype genitalia in dorsal (A), ventral (B) and lateral (C) views; basal part of the genitalia in ventral view (D). Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10808026" figureDoi-1="http://doi.org/10.5281/zenodo.10808028" figureDoi-2="http://doi.org/10.5281/zenodo.10808032" httpUri-0="https://zenodo.org/record/10808026/files/figure.png" httpUri-1="https://zenodo.org/record/10808028/files/figure.png" httpUri-2="https://zenodo.org/record/10808032/files/figure.png" pageId="8" pageNumber="69">Figs 10–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372B2ADFFF1FFCC43C6F989FD0B23BF" pageId="8" pageNumber="69" type="materials_examined">
<paragraph id="8BD7E126FFF1FFCC43C6F989FD0B23BF" blockId="8.[151,1437,1585,2008]" pageId="8" pageNumber="69">
<emphasis id="B91C3D34FFF1FFCC43C6F989FE5F2367" bold="true" box="[151,390,1585,1611]" pageId="8" pageNumber="69">Material examined.</emphasis>
<materialsCitation id="3B00EB7BFFF1FFCC42C3F989FD0B23BF" collectingDate="2023-11-06" collectionCode="FRC" collectorName="M. K. Tan &amp; Tan, R &amp; Japir, M. A. B &amp; D. F. A. Damit" country="Malaysia" elevation="14" latitude="6.01287" location="Tenompok Forest Reserve" longLatPrecision="1" longitude="116.50288" pageId="8" pageNumber="69" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B91C3D34FFF1FFCC42C3F989FDD02367" bold="true" box="[402,521,1585,1611]" pageId="8" pageNumber="69">
<typeStatus id="54D35F84FFF1FFCC42C3F989FDDA2367" box="[402,515,1585,1611]" pageId="8" pageNumber="69" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="F37FA1B6FFF1FFCC4136F989FD242367" box="[615,765,1585,1611]" name="Malaysia" pageId="8" pageNumber="69">MALAYSIA</collectingCountry>
:
<collectingRegion id="49AC2FC4FFF1FFCC405CF989FC4F2367" box="[781,918,1585,1611]" country="Malaysia" name="Sabah" pageId="8" pageNumber="69">Sabah State</collectingRegion>
•
<specimenCount id="9D6E2AAFFFF1FFCC40E9F98AFC092360" box="[952,976,1586,1612]" count="1" pageId="8" pageNumber="69" type="male">♂</specimenCount>
;
<location id="8EB7B7FDFFF1FFCC40B0F989FACF2360" LSID="urn:lsid:plazi:treatment:03C15030FFF1FFC943C6FA7EFE392449:8EB7B7FDFFF1FFCC40B0F989FACF2360" box="[993,1302,1585,1612]" country="Malaysia" latitude="6.01287" longLatPrecision="1" longitude="116.50288" name="Tenompok Forest Reserve" pageId="8" pageNumber="69" stateProvince="Sabah">Tenompok Forest Reserve</location>
;
<geoCoordinate id="EE5C87E1FFF1FFCC4677F989FA402367" box="[1318,1433,1585,1611]" degrees="6.01287" direction="north" orientation="latitude" pageId="8" pageNumber="69" precision="1" value="6.01287">N6.01287</geoCoordinate>
,
<geoCoordinate id="EE5C87E1FFF1FFCC43C6F9EDFEF8235C" box="[151,289,1621,1648]" degrees="116.50288" direction="east" orientation="longitude" pageId="8" pageNumber="69" precision="1" value="116.50288">E116.50288</geoCoordinate>
, 1366.0±
<quantity id="4C904CC3FFF1FFCC42DDF9EDFE3B2343" box="[396,482,1621,1648]" metricMagnitude="1" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="69" unit="m" value="13.5">
<elevation id="00450615FFF1FFCC42DDF9EDFE3B2343" box="[396,482,1621,1648]" metricMagnitude="1" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="69" unit="m" value="13.5">13.5 m</elevation>
</quantity>
.a.s.l.;
<date id="FFD6C7E6FFF1FFCC4177F9EDFD212343" box="[550,760,1621,1647]" pageId="8" pageNumber="69" value="2023-11-06">
<collectingDate id="EF923E0EFFF1FFCC4177F9EDFD212343" box="[550,760,1621,1647]" pageId="8" pageNumber="69" value="2023-11-06">6 November 2023</collectingDate>
</date>
, 20h15; on a ginger plant foliage; coll.
<collectorName id="269D84F0FFF1FFCC4791F9EEFAE9235C" box="[1216,1328,1622,1648]" pageId="8" pageNumber="69">M.K. Tan</collectorName>
, R.
<collectorName id="269D84F0FFF1FFCC4631F9EDFF3123B8" pageId="8" pageNumber="69">Japir, M.A.B</collectorName>
. Asidi &amp;
<collectorName id="269D84F0FFF1FFCC4203F9C2FE3723BF" box="[338,494,1657,1684]" pageId="8" pageNumber="69">D.F.A. Damit</collectorName>
; SBH.23.189 (
<collectionCode id="ED7979E3FFF1FFCC41C3F9C1FD1323BF" box="[658,714,1657,1683]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="8" pageNumber="69">FRC</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C372B2ADFFF1FFCC4396F925FC8B22FB" pageId="8" pageNumber="69" type="diagnosis">
<paragraph id="8BD7E126FFF1FFCC4396F925FBCA22A3" blockId="8.[151,1437,1585,2008]" pageId="8" pageNumber="69">
<emphasis id="B91C3D34FFF1FFCC4396F925FE9B239B" bold="true" box="[199,322,1693,1719]" pageId="8" pageNumber="69">Diagnosis.</emphasis>
This new species is most similar to
<taxonomicName id="4C689AA5FFF1FFCC4180F926FB0E239B" authority="Gorochov, 1997" authorityName="Gorochov" authorityYear="1997" box="[721,1239,1693,1719]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="borneoensis" subGenus="Bornitara">
<emphasis id="B91C3D34FFF1FFCC4180F926FCD0239B" box="[721,777,1694,1719]" italics="true" pageId="8" pageNumber="69">Itara</emphasis>
(
<emphasis id="B91C3D34FFF1FFCC4049F926FC5F239B" box="[792,902,1694,1719]" italics="true" pageId="8" pageNumber="69">Bornitara</emphasis>
)
<emphasis id="B91C3D34FFF1FFCC40C7F925FBC4239B" box="[918,1053,1693,1719]" italics="true" pageId="8" pageNumber="69">borneoensis</emphasis>
<bibRefCitation id="EFF99CD7FFF1FFCC4775F925FB0E239B" author="Gorochov, A. V." box="[1060,1239,1693,1719]" pageId="8" pageNumber="69" pagination="47 - 75" refId="ref7092" refString="Gorochov, A. V. (1997) Partial revision of the subfamily Itarinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 6 (1 - 2), 47 - 75." type="journal article" year="1997">Gorochov, 1997</bibRefCitation>
</taxonomicName>
from Kalimantan and
<taxonomicName id="4C689AA5FFF1FFCC4396F97AFD7823F7" authority="Gorochov, 2007" authorityName="Gorochov" authorityYear="2007" box="[199,673,1729,1755]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="copiosa" subGenus="Bornitara">
<emphasis id="B91C3D34FFF1FFCC4396F97AFF2623F7" box="[199,255,1730,1755]" italics="true" pageId="8" pageNumber="69">Itara</emphasis>
(
<emphasis id="B91C3D34FFF1FFCC425EF97AFEA423F7" box="[271,381,1730,1755]" italics="true" pageId="8" pageNumber="69">Bornitara</emphasis>
)
<emphasis id="B91C3D34FFF1FFCC42DFF97AFE3C23F7" box="[398,485,1730,1755]" italics="true" pageId="8" pageNumber="69">copiosa</emphasis>
<bibRefCitation id="EFF99CD7FFF1FFCC42BCF979FD7823F7" author="Gorochov, A. V." box="[493,673,1729,1755]" pageId="8" pageNumber="69" pagination="201 - 207" refId="ref7158" refString="Gorochov, A. V. (2007) Fourth addition to the revision of Itarinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 16 (2), 201 - 207. https: // doi. org / 10.31610 / zsr / 2007.16.2.201" type="journal article" year="2007">Gorochov, 2007</bibRefCitation>
</taxonomicName>
from Mount Trus Madi (
<collectingRegion id="49AC2FC4FFF1FFCC40EAF979FBDA23F7" box="[955,1027,1729,1755]" country="Malaysia" name="Sabah" pageId="8" pageNumber="69">Sabah</collectingRegion>
) in the male genitalia; but differs by the pseudepiphallus having its apex narrower in dorsal view and not as strongly curved in lateral view, and by the pseudepiphallic parameres in ventral view having the distal part distinctly longer than the proximal part (instead of the proximal part longer) and a distinctly longer concavity along the inner margin (
<figureCitation id="1353FDA3FFF1FFCC477DF895FB0A226B" box="[1068,1235,1837,1863]" captionStart="FIGURE 13" captionStartId="15.[151,250,1982,2006]" captionTargetBox="[155,1436,491,1952]" captionTargetId="figure-111@15.[151,1436,477,1960]" captionTargetPageId="15" captionText="FIGURE 13. Itara (Bornitara) copiosa (A, B) and Itara (Maxitara) kinabalu (C–E) ♂ genitalia in lateral (A, C), dorsal (D) and ventral (B, E) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10808034" httpUri="https://zenodo.org/record/10808034/files/figure.png" pageId="8" pageNumber="69">Figs 13A, 13B</figureCitation>
). The new species also differs from
<taxonomicName id="4C689AA5FFF1FFCC420CF8EAFDA72247" authorityName="Gorochov" authorityYear="2007" box="[349,638,1873,1899]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="copiosa" subGenus="Bornitara">
<emphasis id="B91C3D34FFF1FFCC420CF8EAFE4C2247" box="[349,405,1874,1899]" italics="true" pageId="8" pageNumber="69">Itara</emphasis>
(
<emphasis id="B91C3D34FFF1FFCC42F6F8EAFDCC2247" box="[423,533,1874,1899]" italics="true" pageId="8" pageNumber="69">Bornitara</emphasis>
)
<emphasis id="B91C3D34FFF1FFCC4176F8EAFDA72247" box="[551,638,1874,1899]" italics="true" pageId="8" pageNumber="69">copiosa</emphasis>
</taxonomicName>
by the pseudepiphallus longer and at apical half narrower; from
<taxonomicName id="4C689AA5FFF1FFCC4635F8EAFE7F22A3" authorityName="Gorochov" authorityYear="1997" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="borneoensis" subGenus="Bornitara">
<emphasis id="B91C3D34FFF1FFCC4635F8EAFA452247" box="[1380,1436,1874,1899]" italics="true" pageId="8" pageNumber="69">Itara</emphasis>
(
<emphasis id="B91C3D34FFF1FFCC43F0F8CEFED622A3" box="[161,271,1910,1935]" italics="true" pageId="8" pageNumber="69">Bornitara</emphasis>
)
<emphasis id="B91C3D34FFF1FFCC424EF8CDFE7F22A3" box="[287,422,1909,1935]" italics="true" pageId="8" pageNumber="69">borneoensis</emphasis>
</taxonomicName>
by the posterior-most harp vein forming a zig-zag line.
</paragraph>
<paragraph id="8BD7E126FFF1FFCC4396F822FC8B22FB" blockId="8.[151,1437,1585,2008]" pageId="8" pageNumber="69">
From the syntopic species
<taxonomicName id="4C689AA5FFF1FFCC42AFF822FC35229F" authority="Gorochov, 2013" authorityName="Gorochov" authorityYear="2013" box="[510,1004,1945,1971]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="kinabalu" subGenus="Maxitara">
<emphasis id="B91C3D34FFF1FFCC42AFF822FDEF229F" box="[510,566,1946,1971]" italics="true" pageId="8" pageNumber="69">Itara</emphasis>
(
<emphasis id="B91C3D34FFF1FFCC411BF822FD6B229F" box="[586,690,1946,1971]" italics="true" pageId="8" pageNumber="69">Maxitara</emphasis>
)
<emphasis id="B91C3D34FFF1FFCC4197F821FCF1229F" box="[710,808,1945,1971]" italics="true" pageId="8" pageNumber="69">kinabalu</emphasis>
<bibRefCitation id="EFF99CD7FFF1FFCC4063F821FC35229F" author="Gorochov, A. V." box="[818,1004,1945,1971]" pageId="8" pageNumber="69" pagination="224 - 229" refId="ref7250" refString="Gorochov, A. V. (2013) Eighth addition to the revision of Itarinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 22 (2), 224 - 229. https: // doi. org / 10.31610 / zsr / 2013.22.2.224" type="journal article" year="2013">Gorochov, 2013</bibRefCitation>
</taxonomicName>
, this new species differs by the male genitalia (
<figureCitation id="1353FDA3FFF1FFCC4257F805FE5722FB" box="[262,398,1981,2007]" captionStart="FIGURE 13" captionStartId="15.[151,250,1982,2006]" captionTargetBox="[155,1436,491,1952]" captionTargetId="figure-111@15.[151,1436,477,1960]" captionTargetPageId="15" captionText="FIGURE 13. Itara (Bornitara) copiosa (A, B) and Itara (Maxitara) kinabalu (C–E) ♂ genitalia in lateral (A, C), dorsal (D) and ventral (B, E) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10808034" httpUri="https://zenodo.org/record/10808034/files/figure.png" pageId="8" pageNumber="69">Figs 13C–E</figureCitation>
) and by a smaller and slenderer habitus.
</paragraph>
</subSubSection>
<caption id="DF17B1AEFFF0FFCD43C6F83DFC2622B1" ID-DOI="http://doi.org/10.5281/zenodo.10808017" ID-Zenodo-Dep="10808017" box="[151,1023,1925,1949]" httpUri="https://zenodo.org/record/10808017/files/figure.png" pageId="9" pageNumber="70" startId="9.[151,250,1925,1949]" targetBox="[151,1436,181,1901]" targetPageId="9" targetType="figure">
<paragraph id="8BD7E126FFF0FFCD43C6F83DFC2622B1" blockId="9.[151,1023,1925,1949]" box="[151,1023,1925,1949]" pageId="9" pageNumber="70">
<emphasis id="B91C3D34FFF0FFCD43C6F83DFECA22B1" bold="true" box="[151,275,1925,1949]" pageId="9" pageNumber="70">FIGURE 6.</emphasis>
<taxonomicName id="4C689AA5FFF0FFCD4248F83EFDF422B1" authorityName="Gorochov" authorityYear="1997" box="[281,557,1925,1949]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="singularis" subGenus="Singitara">
<emphasis id="B91C3D34FFF0FFCD4248F83EFE9522B1" box="[281,332,1926,1949]" italics="true" pageId="9" pageNumber="70">Itara</emphasis>
(
<emphasis id="B91C3D34FFF0FFCD420AF83DFE6022B0" box="[347,441,1925,1948]" italics="true" pageId="9" pageNumber="70">Singitara</emphasis>
)
<emphasis id="B91C3D34FFF0FFCD4299F83DFDF422B1" box="[456,557,1925,1949]" italics="true" pageId="9" pageNumber="70">singularis</emphasis>
</taxonomicName>
♂ calling among shrubs at Mount Trus Madi.
</paragraph>
</caption>
<caption id="DF17B1AEFFF3FFCE43C6FD04FDCA27DB" ID-DOI="http://doi.org/10.5281/zenodo.10808019" ID-Zenodo-Dep="10808019" httpUri="https://zenodo.org/record/10808019/files/figure.png" pageId="10" pageNumber="71" startId="10.[151,250,700,724]" targetBox="[160,1426,181,675]" targetPageId="10" targetType="figure">
<paragraph id="8BD7E126FFF3FFCE43C6FD04FDCA27DB" blockId="10.[151,1437,700,760]" pageId="10" pageNumber="71">
<emphasis id="B91C3D34FFF3FFCE43C6FD04FECA27F8" bold="true" box="[151,275,700,724]" pageId="10" pageNumber="71">FIGURE 7.</emphasis>
<taxonomicName id="4C689AA5FFF3FFCE4248FD05FDF527F8" authorityName="Gorochov" authorityYear="1997" box="[281,556,700,724]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="species" species="singularis" subGenus="Singitara">
<emphasis id="B91C3D34FFF3FFCE4248FD05FE9527F8" box="[281,332,701,724]" italics="true" pageId="10" pageNumber="71">Itara</emphasis>
(
<emphasis id="B91C3D34FFF3FFCE420BFD04FE6127FF" box="[346,440,700,723]" italics="true" pageId="10" pageNumber="71">Singitara</emphasis>
)
<emphasis id="B91C3D34FFF3FFCE4296FD04FDF527F8" box="[455,556,700,724]" italics="true" pageId="10" pageNumber="71">singularis</emphasis>
</taxonomicName>
♂ male genitalia in ventral (only apical part) (A) and lateral (B) views; spermatophore in lateral view (C). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF17B1AEFFF3FFCE43C6F91CFC152390" ID-DOI="http://doi.org/10.5281/zenodo.10808021" ID-Zenodo-Dep="10808021" box="[151,972,1700,1724]" httpUri="https://zenodo.org/record/10808021/files/figure.png" pageId="10" pageNumber="71" startId="10.[151,250,1700,1724]" targetBox="[151,1435,820,1676]" targetPageId="10" targetType="figure">
<paragraph id="8BD7E126FFF3FFCE43C6F91CFC152390" blockId="10.[151,972,1700,1724]" box="[151,972,1700,1724]" pageId="10" pageNumber="71">
<emphasis id="B91C3D34FFF3FFCE43C6F91CFECA2390" bold="true" box="[151,275,1700,1724]" pageId="10" pageNumber="71">FIGURE 8.</emphasis>
<taxonomicName id="4C689AA5FFF3FFCE4248F91DFDF42390" authorityName="Gorochov" authorityYear="1997" box="[281,557,1700,1724]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="species" species="singularis" subGenus="Singitara">
<emphasis id="B91C3D34FFF3FFCE4248F91DFE952390" box="[281,332,1701,1724]" italics="true" pageId="10" pageNumber="71">Itara</emphasis>
(
<emphasis id="B91C3D34FFF3FFCE420AF91DFE602390" box="[347,441,1701,1724]" italics="true" pageId="10" pageNumber="71">Singitara</emphasis>
)
<emphasis id="B91C3D34FFF3FFCE4299F91CFDF42390" box="[456,557,1700,1724]" italics="true" pageId="10" pageNumber="71">singularis</emphasis>
</taxonomicName>
♂ attracted to light at Mount Trus Madi.
</paragraph>
</caption>
<subSubSection id="C372B2ADFFF3FFCE4396F8B7FBAB2205" box="[199,1138,1806,1833]" pageId="10" pageNumber="71" type="etymology">
<paragraph id="8BD7E126FFF3FFCE4396F8B7FBAB2205" blockId="10.[151,1436,1806,2013]" box="[199,1138,1806,1833]" pageId="10" pageNumber="71">
<emphasis id="B91C3D34FFF3FFCE4396F8B7FE972205" bold="true" box="[199,334,1807,1833]" pageId="10" pageNumber="71">Etymology.</emphasis>
The species is named after the
<typeStatus id="54D35F84FFF3FFCE41FFF8B7FD072205" box="[686,734,1807,1833]" pageId="10" pageNumber="71">type</typeStatus>
locality, Tenompok Forest Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C372B2ADFFF3FFC84396F88AFC7722CA" lastPageId="12" lastPageNumber="73" pageId="10" pageNumber="71" type="description">
<paragraph id="8BD7E126FFF3FFCF4396F88AFC982711" blockId="10.[151,1436,1806,2013]" lastBlockId="11.[151,1437,151,573]" lastPageId="11" lastPageNumber="72" pageId="10" pageNumber="71">
<emphasis id="B91C3D34FFF3FFCE4396F88AFE802260" bold="true" box="[199,345,1842,1868]" pageId="10" pageNumber="71">Description.</emphasis>
Medium sized among congeners. Head dorsum shiny dark brown (
<figureCitation id="1353FDA3FFF3FFCE476EF88AFB7C2261" box="[1087,1189,1842,1869]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="10" pageNumber="71">Fig. 11A</figureCitation>
). Fastigium darkened; 1.8 times wider than scapes (
<figureCitation id="1353FDA3FFF3FFCE4284F8EEFDE5225D" box="[469,572,1878,1905]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="10" pageNumber="71">Fig. 11A</figureCitation>
). Scapes brown with tint of dark brown (
<figureCitation id="1353FDA3FFF3FFCE40ADF8EEFBBB225D" box="[1020,1122,1878,1905]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="10" pageNumber="71">Fig. 11A</figureCitation>
). Eyes in dorsal view faintly protruding anteriorly (
<figureCitation id="1353FDA3FFF3FFCE42C7F8C2FE2722B9" box="[406,510,1914,1941]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="10" pageNumber="71">Fig. 11A</figureCitation>
) and in profile view taller than long (
<figureCitation id="1353FDA3FFF3FFCE40E6F8C2FBC622B9" box="[951,1055,1914,1941]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="10" pageNumber="71">Fig. 11B</figureCitation>
). Median ocellus hermispherical; lateral ocelli larger and broadly oval, located dorsal of scapes (
<figureCitation id="1353FDA3FFF3FFCE4020F826FC002295" box="[881,985,1950,1977]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="10" pageNumber="71">Fig. 11C</figureCitation>
). Maxillary palpi with fine short setae; with apical segment dark brown, longest and apical half strongly oblique; with subapical segment shorter than third segment; subapical segment at apex faintly expanding; third segment cylindrical (
<figureCitation id="1353FDA3FFF2FFCF4760FF2FFB41259D" box="[1073,1176,151,177]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="11" pageNumber="72">Fig. 11D</figureCitation>
). Face in anterior view rounded, as tall as wide; area between scapes yellow brown; otherwise brown to grey brown (
<figureCitation id="1353FDA3FFF2FFCF47FFFF03FACA25F9" box="[1198,1299,187,213]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="11" pageNumber="72">Fig. 11C</figureCitation>
). Gena light brown; slightly dark brown posterior of eye (
<figureCitation id="1353FDA3FFF2FFCF41CFFF67FCDF25D5" box="[670,774,223,249]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="11" pageNumber="72">Fig. 11B</figureCitation>
). Pronotal disk unicolourous dark brown; 1.6 times wider than long, widening posteriorly (posterior margin 1.5 times wider than anterior margin); very finely pubescent, with a row of setae along posterior and anterior margins; anterior margin broadly concave; posterior margin somewhat straightened at middle (
<figureCitation id="1353FDA3FFF2FFCF42F7FEF3FDC92449" box="[422,528,331,357]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="11" pageNumber="72">Fig. 11A</figureCitation>
). Pronotal lateral lobe same colouration as dorsal disk, ventral part with tint of fainter brown; ventral margin dark and nearly straight; ventro-posterior angle sloping (
<figureCitation id="1353FDA3FFF2FFCF4738FED7FB1624A5" box="[1129,1231,367,393]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="11" pageNumber="72">Fig. 11B</figureCitation>
). TI without inner tympanum, outer tympanum with opening elongated oval (
<figureCitation id="1353FDA3FFF2FFCF407BFE2BFC572481" box="[810,910,403,429]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="11" pageNumber="72">Fig. 11E</figureCitation>
). TIII with 4 inner and 4 outer subapical spurs, 3 inner and 3 outer apical spurs; with 4 inner and 5 outer small spines before most proximal subapical spurs; without small spine between spurs. FIs and FIIs brown, with basal half paler. TIs generally brown. FIIIs pale brown; apical part including knee dark brown with tint of red brown. TIIIs generally yellow brown; spurs generally yellow brown with apices dark brown. Tergites and sternites yellow brown.
</paragraph>
<caption id="DF17B1AEFFF2FFCF43C6F8DFFC3E22EB" ID-DOI="http://doi.org/10.5281/zenodo.10808023" ID-Zenodo-Dep="10808023" httpUri="https://zenodo.org/record/10808023/files/figure.png" pageId="11" pageNumber="72" startId="11.[151,250,1895,1919]" targetBox="[162,1425,608,1867]" targetPageId="11" targetType="figure">
<paragraph id="8BD7E126FFF2FFCF43C6F8DFFC3E22EB" blockId="11.[151,1436,1895,1991]" pageId="11" pageNumber="72">
<emphasis id="B91C3D34FFF2FFCF43C6F8DFFEC92253" bold="true" box="[151,272,1895,1919]" pageId="11" pageNumber="72">FIGURE 9.</emphasis>
<taxonomicName id="4C689AA5FFF2FFCF4244F8D0FDFD2253" authorityName="Gorochov" authorityYear="1997" box="[277,548,1895,1920]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="singularis" subGenus="Singitara">
<emphasis id="B91C3D34FFF2FFCF4244F8D0FE912253" box="[277,328,1896,1919]" italics="true" pageId="11" pageNumber="72">Itara</emphasis>
(
<emphasis id="B91C3D34FFF2FFCF4205F8D0FE6B2253" box="[340,434,1896,1919]" italics="true" pageId="11" pageNumber="72">Singitara</emphasis>
)
<emphasis id="B91C3D34FFF2FFCF42EEF8DFFDFD2253" box="[447,548,1895,1919]" italics="true" pageId="11" pageNumber="72">singularis</emphasis>
</taxonomicName>
♂ calling song: oscillograms showing five echemes (A) and one echeme (B); spectrogram of the same echeme (C); power spectra of a syllable (D) where the red line represents the power [dB] (axis name and scale in red) and the black solid peak represents the relative amplitude (axis name in black).
</paragraph>
</caption>
<caption id="DF17B1AEFFF5FFC843C6FB9DFA4B2112" ID-DOI="http://doi.org/10.5281/zenodo.10808026" ID-Zenodo-Dep="10808026" box="[151,1426,1061,1086]" httpUri="https://zenodo.org/record/10808026/files/figure.png" pageId="12" pageNumber="73" startId="12.[151,250,1061,1085]" targetBox="[151,1435,181,1037]" targetPageId="12" targetType="figure">
<paragraph id="8BD7E126FFF5FFC843C6FB9DFA4B2112" blockId="12.[151,1426,1061,1086]" box="[151,1426,1061,1086]" pageId="12" pageNumber="73">
<emphasis id="B91C3D34FFF5FFC843C6FB9DFEF92111" bold="true" box="[151,288,1061,1085]" pageId="12" pageNumber="73">FIGURE 10.</emphasis>
<taxonomicName id="4C689AA5FFF5FFC84277FB9EFDE52112" authority="Tan &amp; Grumo &amp; Gono &amp; Bahoy &amp; Rivera &amp; Nuñeza &amp; Japir &amp; Chung, 2024" authorityName="Tan &amp; Grumo &amp; Gono &amp; Bahoy &amp; Rivera &amp; Nuñeza &amp; Japir &amp; Chung" authorityYear="2024" box="[294,572,1062,1086]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="tenompok" status="sp. nov." subGenus="Bornitara">
<emphasis id="B91C3D34FFF5FFC84277FB9EFE802111" box="[294,345,1062,1085]" italics="true" pageId="12" pageNumber="73">Itara</emphasis>
(
<emphasis id="B91C3D34FFF5FFC84236FB9EFE122111" box="[359,459,1062,1085]" italics="true" pageId="12" pageNumber="73">Bornitara</emphasis>
)
<emphasis id="B91C3D34FFF5FFC8428BFB9EFDE52112" box="[474,572,1062,1086]" italics="true" pageId="12" pageNumber="73">tenompok</emphasis>
</taxonomicName>
<emphasis id="B91C3D34FFF5FFC84112FB9EFD4D2112" bold="true" box="[579,660,1062,1086]" pageId="12" pageNumber="73">
<taxonomicNameLabel id="A22F804FFFF5FFC84112FB9EFD4D2112" box="[579,660,1062,1086]" pageId="12" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ holotype in its natural environment at Tenompok Forest Reserve, Sabah.
</paragraph>
</caption>
<paragraph id="8BD7E126FFF5FFC84396FB28FB3020AE" blockId="12.[151,1437,1168,2023]" pageId="12" pageNumber="73">
<emphasis id="B91C3D34FFF5FFC84396FB28FED22186" bold="true" box="[199,267,1168,1194]" pageId="12" pageNumber="73">Male.</emphasis>
FW 2.3 times longer than wide, surpassing abdominal apex, mostly yellow brown and hyalinous (
<figureCitation id="1353FDA3FFF5FFC8463EFB28FF1F21E2" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="12" pageNumber="73">Fig. 11F</figureCitation>
). FW venation (
<figureCitation id="1353FDA3FFF5FFC84226FB0CFE0021E2" box="[375,473,1204,1230]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="12" pageNumber="73">Fig. 11F</figureCitation>
): dorsal field in harp (ha) with four increasingly longer veins; posterior-most harp vein at base curved, at distal half forming zig-zag line. Mirror (mi) very large and transverse, 1.7 times as wide as long, very large and separated by one dividing vein; dividing vein broadly curved. Apical field relatively short, 1.2 times longer than length of mirror (
<figureCitation id="1353FDA3FFF5FFC842BBFA98FD892016" box="[490,592,1312,1338]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="12" pageNumber="73">Fig. 11F</figureCitation>
). Lateral field with R and M generally parallel, with numerous very faint transverse veins; M and Sc mostly parallel and more closely spaced apart; Sc with numerous inter-laced projections, with ca. 10 tranverse projecting veins (
<figureCitation id="1353FDA3FFF5FFC8411BFAD0FD6B20AE" box="[586,690,1384,1410]" captionStart="FIGURE 11" captionStartId="13.[151,250,1932,1956]" captionTargetBox="[151,1436,381,1909]" captionTargetId="figure-77@13.[151,1436,381,1909]" captionTargetPageId="13" captionText="FIGURE 11. Itara (Bornitara) tenompok sp. nov. ♂ holotype: head and pronotum in dorsal (A) and lateral (B) views; face in anterior view (C); maxillary palps in lateral view (D); outer tympanum on TI in external view (E); FW in dorsal (F) and lateral (G) views. Scale bars: 5 mm (F, G), 1 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.10808028" httpUri="https://zenodo.org/record/10808028/files/figure.png" pageId="12" pageNumber="73">Fig. 11G</figureCitation>
). Hind wings only slightly surpassing apex of FW.
</paragraph>
<paragraph id="8BD7E126FFF5FFC84396FA34FD272256" blockId="12.[151,1437,1168,2023]" pageId="12" pageNumber="73">
Male genitalia (
<figureCitation id="1353FDA3FFF5FFC84229FA34FE15208A" box="[376,460,1420,1446]" captionStart="FIGURE 12" captionStartId="14.[151,250,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-21@14.[151,1436,181,1083]" captionTargetPageId="14" captionText="FIGURE 12. Itara (Bornitara) tenompok sp. nov. ♂ holotype genitalia in dorsal (A), ventral (B) and lateral (C) views; basal part of the genitalia in ventral view (D). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10808032" httpUri="https://zenodo.org/record/10808032/files/figure.png" pageId="12" pageNumber="73">Fig. 12</figureCitation>
). Pseudepiphallus [epiphallus] typical of subgenus; in dorsal view triangular with apex acute; in lateral view flattened and at apex hooked. Pseudepiphallus in dorsal view: after middle strongly tapering, with lateral margin slightly concave. Pseudepiphallus in lateral view: with upper proximal edge slightly raised; dorsal margin at proximal half gently sloping, distal half slightly curved; at apex with a small tooth pointing dorsad; ventral margin lacking denticles on ventral surface, basally with tongue-shaped lobe pointing posteriorly, at middle slightly concave, distal half mostly straight. Pseudepiphallic parameres [ectoparameres] not surpassing apex of pseudepiphallus, sinuous. Pseudepiphallic parameres with proximal part reaching basal third with basal margin oblique and lateral margins straight and parallel; just before middle with inner margin forming 90° bent and outer margin broadly convex; thereafter with distal part having inner margin with deep and elongated concavity extending to near apex, outer margin broadly curved; at apex somewhat triangular and subobtuse. Ectophallic apodeme [endoparameral apodeme] in ventral view, straight to faintly curved. Ectophallic fold [rachis] needle-like, long and slender with posterior apex acute, tubular and strongly curved anteriorly into endophallic sclerite [formula]. Endophallic sclerite small, slender and Y-shaped, ventrad of ectophallic fold. Rami slightly curved, longer than pseudepiphallus length, at anterior ends not connected.
</paragraph>
<paragraph id="8BD7E126FFF5FFC84396F83CFE7B22B2" blockId="12.[151,1437,1168,2023]" box="[199,418,1924,1950]" pageId="12" pageNumber="73">
<emphasis id="B91C3D34FFF5FFC84396F83CFEFC22B2" bold="true" box="[199,293,1924,1950]" pageId="12" pageNumber="73">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BD7E126FFF5FFC84396F810FC7722CA" blockId="12.[151,1437,1168,2023]" pageId="12" pageNumber="73">
<emphasis id="B91C3D34FFF5FFC84396F810FE3F22ED" bold="true" box="[199,486,1959,1986]" pageId="12" pageNumber="73">Measurements (in mm).</emphasis>
<specimenCount id="9D6E2AAFFFF5FFC842BFF810FDDA22EE" box="[494,515,1960,1986]" count="1" pageId="12" pageNumber="73" type="male">♂</specimenCount>
<typeStatus id="54D35F84FFF5FFC8415BF810FDA922EE" box="[522,624,1960,1986]" pageId="12" pageNumber="73" type="holotype">holotype</typeStatus>
: BL = 16.3; BWL = 24.1; HL = 2.1; PronL = 2.7; PronW = 4.3; FWL = 17.0; FWW = 7.5; HWT = 2.3; FIIIL = 10.5; TIIIL = 6.9; TaIIIL = 3.0.
</paragraph>
</subSubSection>
<subSubSection id="C372B2ADFFF4FFC94396FF2FFE392449" pageId="13" pageNumber="74" type="biology_ecology">
<paragraph id="8BD7E126FFF4FFC94396FF2FFE392449" blockId="13.[151,1437,151,357]" pageId="13" pageNumber="74">
<emphasis id="B91C3D34FFF4FFC94396FF2FFEF5259D" bold="true" box="[199,300,151,177]" pageId="13" pageNumber="74">Ecology.</emphasis>
We found this species to co-occur in syntopy with
<taxonomicName id="4C689AA5FFF4FFC94028FF2FFA87259D" authority="Gorochov, 2013" authorityName="Gorochov" authorityYear="2013" box="[889,1374,151,177]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="kinabalu" subGenus="Maxitara">
<emphasis id="B91C3D34FFF4FFC94028FF2FFC68259C" box="[889,945,151,176]" italics="true" pageId="13" pageNumber="74">Itara</emphasis>
(
<emphasis id="B91C3D34FFF4FFC94092FF2FFBF2259C" box="[963,1067,151,176]" italics="true" pageId="13" pageNumber="74">Maxitara</emphasis>
)
<emphasis id="B91C3D34FFF4FFC9476CFF2FFB46259D" box="[1085,1183,151,177]" italics="true" pageId="13" pageNumber="74">kinabalu</emphasis>
<bibRefCitation id="EFF99CD7FFF4FFC947F9FF2FFA87259D" author="Gorochov, A. V." box="[1192,1374,151,177]" pageId="13" pageNumber="74" pagination="224 - 229" refId="ref7250" refString="Gorochov, A. V. (2013) Eighth addition to the revision of Itarinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 22 (2), 224 - 229. https: // doi. org / 10.31610 / zsr / 2013.22.2.224" type="journal article" year="2013">Gorochov, 2013</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1353FDA3FFF4FFC9463EFF2FFF6C25F9" captionStart="FIGURE 14" captionStartId="16.[151,250,1924,1948]" captionTargetBox="[151,1436,181,1900]" captionTargetId="figure-21@16.[151,1436,181,1900]" captionTargetPageId="16" captionText="FIGURE 14. Itara (Maxitara) kinabalu in their natural environment at Tenompok Forest Reserve, Sabah: male singing (A), mating pair (B)." figureDoi="http://doi.org/10.5281/zenodo.10808036" httpUri="https://zenodo.org/record/10808036/files/figure.png" pageId="13" pageNumber="74">Fig. 14</figureCitation>
), although it appears that this new species is lower in abundance (as more specimens of
<taxonomicName id="4C689AA5FFF4FFC9472DFF03FA4525F9" authorityName="Gorochov" authorityYear="2013" box="[1148,1436,187,213]" class="Insecta" family="Gryllidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="kinabalu" subGenus="Maxitara">
<emphasis id="B91C3D34FFF4FFC9472DFF03FB6D25F8" box="[1148,1204,187,212]" italics="true" pageId="13" pageNumber="74">Itara</emphasis>
(
<emphasis id="B91C3D34FFF4FFC94792FF03FAF225F8" box="[1219,1323,187,212]" italics="true" pageId="13" pageNumber="74">Maxitara</emphasis>
)
<emphasis id="B91C3D34FFF4FFC9466BFF03FA4525F9" box="[1338,1436,187,213]" italics="true" pageId="13" pageNumber="74">kinabalu</emphasis>
</taxonomicName>
were collected).
<materialsCitation id="3B00EB7BFFF4FFC94396FEBBFE052449" country="Malaysia" county="Tenompok Forest Reserve" location="Calling" municipality="Tenompok Forest Reserve" pageId="13" pageNumber="74" specimenCount="1" stateProvince="Sabah">
<emphasis id="B91C3D34FFF4FFC94396FEBBFEBF2430" bold="true" box="[199,358,258,285]" pageId="13" pageNumber="74">
<typeStatus id="54D35F84FFF4FFC94396FEBBFEDB2431" box="[199,258,259,285]" pageId="13" pageNumber="74">Type</typeStatus>
locality.
</emphasis>
EAST
<collectingCountry id="F37FA1B6FFF4FFC942E8FEBBFD962431" box="[441,591,259,285]" name="Malaysia" pageId="13" pageNumber="74">MALAYSIA</collectingCountry>
:
<collectingRegion id="49AC2FC4FFF4FFC94108FEBBFD7B2431" box="[601,674,259,285]" country="Malaysia" name="Sabah" pageId="13" pageNumber="74">Sabah</collectingRegion>
:
<collectingCounty id="62B699AAFFF4FFC941FDFEBBFC082431" box="[684,977,259,285]" pageId="13" pageNumber="74">Tenompok Forest Reserve</collectingCounty>
<emphasis id="B91C3D34FFF4FFC94396FE9EFEB8246C" bold="true" box="[199,353,294,320]" pageId="13" pageNumber="74">Distribution.</emphasis>
Borneo (
<collectingRegion id="49AC2FC4FFF4FFC94298FE9FFDCA246D" box="[457,531,295,321]" country="Malaysia" name="Sabah" pageId="13" pageNumber="74">Sabah</collectingRegion>
:
<collectingMunicipality id="6BB37B5CFFF4FFC9414CFE9FFC9D246D" box="[541,836,295,321]" pageId="13" pageNumber="74">Tenompok Forest Reserve</collectingMunicipality>
)
<emphasis id="B91C3D34FFF4FFC94396FEF2FEBA2449" bold="true" box="[199,355,330,357]" pageId="13" pageNumber="74">
<location id="8EB7B7FDFFF4FFC94396FEF2FEC72448" LSID="urn:lsid:plazi:treatment:03C15030FFF1FFC943C6FA7EFE392449:8EB7B7FDFFF4FFC94396FEF2FEC72448" box="[199,286,330,356]" country="Malaysia" county="Tenompok Forest Reserve" municipality="Tenompok Forest Reserve" name="Calling" pageId="13" pageNumber="74" stateProvince="Sabah">Calling</location>
song.
</emphasis>
<location id="8EB7B7FDFFF4FFC9423BFEF3FE052449" LSID="urn:lsid:plazi:treatment:03C15030FFF1FFC943C6FA7EFE392449:8EB7B7FDFFF4FFC9423BFEF3FE052449" box="[362,476,331,357]" country="Malaysia" county="Tenompok Forest Reserve" municipality="Tenompok Forest Reserve" name="Unknown" pageId="13" pageNumber="74" stateProvince="Sabah">Unknown</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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