245 lines
34 KiB
XML
245 lines
34 KiB
XML
<document id="844D87F8300BBABDD2408330523EC95F" ID-CLB-Dataset="46146" ID-DOI="10.5281/zenodo.278958" ID-GBIF-Dataset="56fb15f0-b55b-4ce9-a9ad-d8a2f5a95029" ID-ISSN="1175-5326" ID-Zenodo-Dep="278958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460401525037" checkinUser="plazi" docAuthor="Williams, Kevin A., Brothers, Denis J. & Pitts, James P." docDate="2011" docId="03BF6217D83D214C06B64EEAFD99FBAC" docLanguage="en" docName="zt03064p068.pdf" docOrigin="Zootaxa 3064" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tobantilla frigidula Cresson 1902, comb. nov." docType="treatment" docVersion="10" lastPageNumber="56" masterDocId="FF861A6FD833214306214C5EFF9CFFBC" masterDocTitle="New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae)" masterLastPageNumber="68" masterPageNumber="41" pageNumber="55" updateTime="1698299261469" updateUser="ExternalLinkService">
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<mods:title id="41923D9F20FCAACE33AB7A700DF99545">New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae)</mods:title>
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<mods:name id="124CD3484250284A01F52E7DCE6D562A" type="personal">
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<mods:namePart id="A508BAD74B45BC804052AC7687CA8292">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="7ACF5AB7C986A6232F919ADBF7608492">Brothers, Denis J.</mods:namePart>
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<mods:name id="3BF3BD01C31BE197FE71044DEACB9BF9" type="personal">
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<mods:roleTerm id="934AAB41472A8C3F99E767EE2C7E646E">Author</mods:roleTerm>
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<mods:namePart id="9490F594C4B7BF526E88125577DBA6E4">Pitts, James P.</mods:namePart>
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<mods:typeOfResource id="24D2D6077502637047771AAC3456EE52">text</mods:typeOfResource>
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<mods:title id="4DBB3FEA39A7C1714010374660FF2AAE">Zootaxa</mods:title>
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<mods:part id="B7FC4BB315D43390CE1806510764D0A1">
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<mods:date id="09A77F1E5C73F03852F0EC62389DDEE9">2011</mods:date>
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<mods:detail id="5B25BF18D6D0AF623029947501A86BE6" type="volume">
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<mods:number id="95233CF3ED29798985EE4F711402349B">3064</mods:number>
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<mods:classification id="CB976231BD3C1330AF134CA6F557A3DE">journal article</mods:classification>
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<mods:identifier id="7D6457BF0A761A46209054F6ECE161C2" type="CLB-Dataset">46146</mods:identifier>
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<mods:identifier id="D346DADECD01E352B16DC13E44D54D7B" type="DOI">10.5281/zenodo.278958</mods:identifier>
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<mods:identifier id="F77BEC0ACF347C654AD1987B13749235" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BF6217D83D214C06B64EEAFD99FBAC" ID-DOI="http://doi.org/10.5281/zenodo.6191531" ID-GBIF-Taxon="119419953" ID-Zenodo-Dep="6191531" LSID="urn:lsid:plazi:treatment:03BF6217D83D214C06B64EEAFD99FBAC" httpUri="http://treatment.plazi.org/id/03BF6217D83D214C06B64EEAFD99FBAC" lastPageId="15" lastPageNumber="56" pageId="14" pageNumber="55">
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<subSubSection id="C30C808AD83D214D06B64EEAFEC2FD52" pageId="14" pageNumber="55" type="nomenclature">
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<paragraph id="8BA9D301D83D214D06B64EEAFD7CFD71" blockId="14.[151,736,691,750]" box="[151,736,691,718]" pageId="14" pageNumber="55">
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<heading id="D0E1646DD83D214D06B64EEAFD7CFD71" bold="true" box="[151,736,691,718]" fontSize="11" level="1" pageId="14" pageNumber="55" reason="1">
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<emphasis id="B9620F13D83D214D06B64EEAFD7CFD71" bold="true" box="[151,736,691,718]" pageId="14" pageNumber="55">
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<taxonomicName id="4C16A882D83D214D06B64EEAFDD0FD71" authority="Cresson, 1902" authorityName="Cresson" authorityYear="1902" box="[151,588,691,718]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="frigidula" status="comb. nov.">
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<emphasis id="B9620F13D83D214D06B64EEAFE1FFD72" bold="true" box="[151,387,692,718]" italics="true" pageId="14" pageNumber="55">Tobantilla frigidula</emphasis>
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||
(
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<bibRefCitation id="EF87AEF0D83D214D07B34EEDFDD8FD71" author="Cresson" box="[402,580,691,717]" pageId="14" pageNumber="55" refString="Cresson, E. T. (1902) Descriptions of some Brazilian Mutilla. Transactions of the American Entomological Society, 28, 1 - 82." type="journal article" year="1902">Cresson, 1902</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="A251B268D83D214D04784EEDFD7CFD71" box="[601,736,691,717]" pageId="14" pageNumber="55">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9D301D83D214D06B64E88FEC2FD52" blockId="14.[151,736,691,750]" box="[151,350,726,750]" pageId="14" pageNumber="55">
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||
(
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||
<figureCitation id="132DCF84D83D214D06BE4E88FF7BFD52" box="[159,231,726,750]" captionStart="FIGURES 1 – 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="14" pageNumber="55">Figs 3</figureCitation>
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,
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<figureCitation id="132DCF84D83D214D06D24E88FEB1FD52" box="[243,301,726,750]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="14" pageNumber="55">9, 13</figureCitation>
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,
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<figureCitation id="132DCF84D83D214D07184E88FECAFD52" box="[313,342,726,750]" captionStart="FIGURES 19 – 38" captionStartId="10.[151,264,1672,1695]" captionTargetBox="[176,1419,215,1644]" captionTargetId="figure@10.[157,1429,193,1651]" captionTargetPageId="10" captionText="FIGURES 19 – 38. Tobantilla species. 19 – 34. Male genitalia: dorsal view (left), ventral view (middle), lateral view of penis valve (above), and lateral view with penis valve removed (right). 19 – 22. T. andrikos sp. nov.; 23 – 26. T. drosos sp. nov.; 27 – 30. T. ephemeros sp. nov.; 31 – 34. T. xouthos sp. nov. 35 – 36. Male clypeus. 28. T. andrikos sp. nov.; 29. T. ephemeros sp. nov. 37, 38. Female pygidium (T 6). 37. T. frigidula (Cresson); 38. T. kolasma sp. nov." httpUri="https://zenodo.org/record/278962/files/figure.png" pageId="14" pageNumber="55">37</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C808AD83D214D06B64F41FD70FCEF" pageId="14" pageNumber="55" type="reference_group">
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<paragraph id="8BA9D301D83D214D06B64F41FD70FCEF" blockId="14.[151,1206,798,851]" pageId="14" pageNumber="55">
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<treatmentCitationGroup id="AB06F42FD83D214D06B64F41FD70FCEF" pageId="14" pageNumber="55">
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<taxonomicName id="4C16A882D83D214D06B64F41FE47FC88" authority="Cresson, 1902" authorityName="Cresson" authorityYear="1902" box="[151,475,799,821]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="frigidula">
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<emphasis id="B9620F13D83D214D06B64F41FEDDFC89" box="[151,321,799,821]" italics="true" pageId="14" pageNumber="55">Mutilla frigidula</emphasis>
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<bibRefCitation id="EF87AEF0D83D214D07664F41FE47FC88" author="Cresson" box="[327,475,799,820]" pageId="14" pageNumber="55" refString="Cresson, E. T. (1902) Descriptions of some Brazilian Mutilla. Transactions of the American Entomological Society, 28, 1 - 82." type="journal article" year="1902">Cresson, 1902</bibRefCitation>
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</taxonomicName>
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. Trans. Amer. Ent. Soc. 28: 40.
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<typeStatus id="54AD6DA3D83D214D053C4F40FCE7FC88" box="[797,891,798,820]" pageId="14" pageNumber="55" type="holotype">Holotype</typeStatus>
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Ƥ,
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<collectingCountry id="F3019391D83D214D05834F40FC7EFC88" box="[930,994,798,820]" name="Brazil" pageId="14" pageNumber="55">Brazil</collectingCountry>
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, Santarem (CMNH).
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<taxonomicName id="4C16A882D83D214D06B64F63FDBBFCEE" authority="Nonveiller 1990: 113" authorityName="Nonveiller" authorityPageNumber="113" authorityYear="1990" box="[151,551,829,851]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="frigidula">
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<emphasis id="B9620F13D83D214D06B64F63FEDDFCEF" box="[151,321,829,851]" italics="true" pageId="14" pageNumber="55">Mutilla frigidula</emphasis>
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:
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<treatmentCitation id="0AB7F510D83D214D076F4F63FDBBFCEE" author="Nonveiller" box="[334,551,829,851]" page="113" pageId="14" pageNumber="55" year="1990">
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<bibRefCitation id="EF87AEF0D83D214D076F4F63FE6AFCEE" author="Nonveiller" box="[334,502,829,851]" pageId="14" pageNumber="55" refString="Nonveiller, G. (1990) Catalogue of the Mutillidae, Myrmosidae and Bradynobaenidae of the Neotropical Region including Mexico (Insecta, Hymenoptera). Hymenopterorum Catalogus (Nova Editio), 18. SPB Academic Publishing, Den Haag, pp. 1 - 150." type="journal article" year="1990">Nonveiller 1990</bibRefCitation>
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: 113
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</treatmentCitation>
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</taxonomicName>
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, Ƥ (incertae sedis).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C808AD83D214D06B64F21FBE1FBB8" pageId="14" pageNumber="55" type="diagnosis">
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<paragraph id="8BA9D301D83D214D06B64F21FBE1FBB8" blockId="14.[151,1437,895,2037]" pageId="14" pageNumber="55">
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<emphasis id="B9620F13D83D214D06B64F21FE8DFC24" bold="true" box="[151,273,895,920]" pageId="14" pageNumber="55">Diagnosis.</emphasis>
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FEMALE. The pygidial plate is small, distant from the apical margin and connected to it by a single median longitudinal carina (
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<figureCitation id="132DCF84D83D214D07F24FFAFDB7FC00" box="[467,555,932,956]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="14" pageNumber="55">Figs 13</figureCitation>
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,
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<figureCitation id="132DCF84D83D214D04164FFAFDCAFC00" box="[567,598,932,956]" captionStart="FIGURES 19 – 38" captionStartId="10.[151,264,1672,1695]" captionTargetBox="[176,1419,215,1644]" captionTargetId="figure@10.[157,1429,193,1651]" captionTargetPageId="10" captionText="FIGURES 19 – 38. Tobantilla species. 19 – 34. Male genitalia: dorsal view (left), ventral view (middle), lateral view of penis valve (above), and lateral view with penis valve removed (right). 19 – 22. T. andrikos sp. nov.; 23 – 26. T. drosos sp. nov.; 27 – 30. T. ephemeros sp. nov.; 31 – 34. T. xouthos sp. nov. 35 – 36. Male clypeus. 28. T. andrikos sp. nov.; 29. T. ephemeros sp. nov. 37, 38. Female pygidium (T 6). 37. T. frigidula (Cresson); 38. T. kolasma sp. nov." httpUri="https://zenodo.org/record/278962/files/figure.png" pageId="14" pageNumber="55">37</figureCitation>
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); the head and mesosoma are dark red to black (
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<figureCitation id="132DCF84D83D214D02594FFAFB20FC00" box="[1144,1212,932,956]" captionStart="FIGURES 1 – 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="14" pageNumber="55">Fig. 3</figureCitation>
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); the scutellar scale is broadly arcuate, but not reaching the lateral propodeal margins; the posterior propodeal face is clothed with silver setae; and T2 has two large, nearly confluent yellowish-orange integumental patches.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C808AD83D214C06E64851FE04FEA0" lastPageId="15" lastPageNumber="56" pageId="14" pageNumber="55" type="description">
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<paragraph id="8BA9D301D83D214C06E64851FEB9FF4B" blockId="14.[151,1437,895,2037]" lastBlockId="15.[151,1437,151,1040]" lastPageId="15" lastPageNumber="56" pageId="14" pageNumber="55">
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<emphasis id="B9620F13D83D214D06E64851FEC4FB9B" bold="true" box="[199,344,1039,1064]" pageId="14" pageNumber="55">Description.</emphasis>
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FEMALE. Body length
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<quantity id="4CEE7EE4D83D214D04514851FD7DFB94" box="[624,737,1039,1064]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="8.0" metricValueMin="5.5" pageId="14" pageNumber="55" unit="mm" value="6.75" valueMax="8.0" valueMin="5.5">5.5-8 mm</quantity>
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.
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<emphasis id="B9620F13D83D214D04CD484EFCF3FB94" box="[748,879,1040,1064]" italics="true" pageId="14" pageNumber="55">Coloration.</emphasis>
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Body and appendages dark red; T2 with one pair of large trapezoidal yellow spots. Tibial spurs white. Head and mesosoma clothed with fairly sparse decumbent pale silvery setae, except anteromedial portion of mesosomal dorsum with patch of dark brown setae, anterolateral portions of mesonotum clothed with denser patch of silver setae, area directly anterior to scutellar scale with cluster of dark brown setae, and dorsum of head and mesosoma with scattered erect long brachyplumose brown setae; posterior face of propodeum with moderately dense decumbent silver setae and a few dark brown setae dorsally. T2 clothed with dark brown erect and appressed setae between spots and along apical margin, lateral thirds and integumental spots with sparse silver erect setae. T3–6 entirely clothed with decumbent silver setae and interspersed erect dark brown and silver setae.
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<emphasis id="B9620F13D83D214D0408496EFDFAFAF4" box="[553,614,1328,1352]" italics="true" pageId="14" pageNumber="55">Head</emphasis>
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. Angulate posterolaterally, with occipital carina stronger laterally from base of weak narrowly triangular glabrous tubercle on posterolateral margin. Head width 1.1 × pronotal width. Eye almost circular. Front, vertex and gena reticulate. Genal carina produced, extending anteriorly to hypostomal carina. Mandible oblique, tapering, bidentate apically but preapical inner tooth minute and usually obliterated, unarmed ventrally. Antennal scrobe lacking carina. Antennal tubercle finely and sparsely punctate basally. Scape simply punctate. Flagellomere 1 1.6 × pedicel length; flagellomere 2 1.1 × pedicel length.
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<emphasis id="B9620F13D83D214D02B549BBFA90FA40" box="[1172,1292,1509,1532]" italics="true" pageId="14" pageNumber="55">Mesosoma</emphasis>
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. Mesosomal length 1.25 × width; pronotum slightly wider than mesothorax. Mesosomal dorsum reticulate. Humeral carina well developed. Lateral face of pronotum and mesopleuron micropunctate, dull, mesopleural ridge coarsely reticulate; clothed with coarse silver setae. Metapleuron dorsal to endophragmal pit and dorsal two-thirds of lateral face of propodeum clothed with fine appressed silver setae; ventral regions of metapleuron and lateral face of propodeum micropunctate and clothed with coarse silver setae. In dorsal view, mesosoma gradually narrowed posterior to pronotum, lateral margin of mesothorax weakly divergent anterior to propodeal spiracle. Scutellar scale broad and highly raised, with obscure w-shaped transverse carina anterior to it. Dorsal and posterior propodeal faces not separated by transverse carina. Dorsal propodeal face convergent posterior to spiracle, posterolateral angle rounded, posterior face virtually flat, vertical, reticulate and clothed with moderately dense decumbent lanceolate setae on dorsal half.
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<emphasis id="B9620F13D83D214D073F4B13FE0FF8D8" box="[286,403,1869,1892]" italics="true" pageId="14" pageNumber="55">Metasoma</emphasis>
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. T1 narrow and petiolate, somewhat cylindrical, 0.4 × as wide as T2. T2 1.2 × longer than wide, dorsomedially flattened, with maximum width situated posteriorly. Disc of T2 deeply and narrowly reticulate anteriorly and coarsely punctate posteriorly; T3–5 densely punctate. S1 with longitudinal carina, S2 with coarse moderately-spaced punctures, punctures small and dense on S3–5. Pygidium lacking lateral carinae, medially having two posteriorly convergent carinae with scattered longitudinal rugae and shagreened medially, anteriorly this “pygidial plate” 0.2 × the pygidial width; areas lateral to these carinae moderately punctate and clothed with erect setae; posterior 0.2 × pygidial area asetose and obscurely punctured, posterior margin with contiguous low transverse carina.
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</paragraph>
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<paragraph id="8BA9D301D83C214C06E44D5AFE04FEA0" blockId="15.[151,1437,151,1040]" box="[197,408,260,284]" pageId="15" pageNumber="56">MALE. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C30C808AD83C214C06E44D79FB3FFE10" pageId="15" pageNumber="56" type="materials_examined">
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<paragraph id="8BA9D301D83C214C06E44D79FB3FFE10" blockId="15.[151,1437,151,1040]" pageId="15" pageNumber="56">
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<emphasis id="B9620F13D83C214C06E44D79FE2AFEFC" bold="true" box="[197,438,295,320]" pageId="15" pageNumber="56">Material examined.</emphasis>
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<typeStatus id="54AD6DA3D83C214C07E44D76FE67FEFC" box="[453,507,296,320]" pageId="15" pageNumber="56">Type</typeStatus>
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material.
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<emphasis id="B9620F13D83C214C045F4D76FD79FEFC" box="[638,741,296,320]" italics="true" pageId="15" pageNumber="56">
|
||
<typeStatus id="54AD6DA3D83C214C045F4D76FD79FEFC" box="[638,741,296,320]" pageId="15" pageNumber="56" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
, Ƥ,
|
||
<collectingCountry id="F3019391D83C214C05074D76FC0CFEFC" box="[806,912,296,320]" name="Brazil" pageId="15" pageNumber="56">BRAZIL</collectingCountry>
|
||
: Pará, Santarem, H.H. Smith (identified as
|
||
<taxonomicName id="4C16A882D83C214C06B64D12FD9DFED8" authority="Cresson" authorityName="Cresson" box="[151,513,332,356]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="frigidula">
|
||
<emphasis id="B9620F13D83C214C06B64D12FE10FED8" box="[151,396,332,356]" italics="true" pageId="15" pageNumber="56">Lomachaeta frigidula</emphasis>
|
||
(Cresson)
|
||
</taxonomicName>
|
||
by D. Quintero in 1996,
|
||
<collectionCode id="ED074BC4D83C214C05024D12FCE0FED8" box="[803,892,332,356]" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="15" pageNumber="56">CMNH</collectionCode>
|
||
). Other material.
|
||
<collectingCountry id="F3019391D83C214C02634D12FB30FED8" box="[1090,1196,332,356]" name="Brazil" pageId="15" pageNumber="56">BRAZIL</collectingCountry>
|
||
: Amazonas, Amazon River,
|
||
<quantity id="4CEE7EE4D83C214C06C74D31FEADFE3B" box="[230,305,367,391]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="56" unit="km" value="50.0">50 km</quantity>
|
||
S Manaus, 1Ƥ,
|
||
<date id="FFA8F5C1D83C214C07D74D31FDCFFE3B" box="[502,595,367,391]" pageId="15" pageNumber="56" value="2004-01-05">5.I.2004</date>
|
||
(
|
||
<collectionCode id="ED074BC4D83C214C04474D31FD26FE3B" box="[614,698,367,391]" pageId="15" pageNumber="56">EMUS</collectionCode>
|
||
); Pará, Taperinha, 1Ƥ,
|
||
<date id="FFA8F5C1D83C214C05EA4D31FBD5FE3B" box="[971,1097,367,391]" pageId="15" pageNumber="56" value="1968-12-08">8.XII.1968</date>
|
||
, on mango leaves, coll. R.L. Jeanne (
|
||
<collectionCode id="ED074BC4D83C214C06D34DCAFED4FE10" box="[242,328,404,428]" pageId="15" pageNumber="56">MCZC</collectionCode>
|
||
); Rondônia, P.E. Guajara Mirim, 1Ƥ,
|
||
<date id="FFA8F5C1D83C214C04CF4DCAFCCBFE10" box="[750,855,404,428]" pageId="15" pageNumber="56" value="1998-02-06">6.II.1998</date>
|
||
, coll. J.R.M. Santos (
|
||
<collectionCode id="ED074BC4D83C214C02664DCAFB0BFE10" box="[1095,1175,404,428]" httpUri="http://grbio.org/cool/95yr-zg8p" name="Centro de Pesquisas do Cacau" pageId="15" pageNumber="56">CPDC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C808AD83C214C06E44DE9FE1DFE48" pageId="15" pageNumber="56" type="distribution">
|
||
<paragraph id="8BA9D301D83C214C06E44DE9FD67FE73" blockId="15.[151,1437,151,1040]" box="[197,763,439,464]" pageId="15" pageNumber="56">
|
||
<emphasis id="B9620F13D83C214C06E44DE9FEC2FE73" bold="true" box="[197,350,439,464]" pageId="15" pageNumber="56">Distribution.</emphasis>
|
||
<collectingCountry id="F3019391D83C214C07444DE9FE35FE73" box="[357,425,439,463]" name="Brazil" pageId="15" pageNumber="56">Brazil</collectingCountry>
|
||
(Amazonas, Pará, Rondônia).
|
||
</paragraph>
|
||
<paragraph id="8BA9D301D83C214C06E44D82FE1DFE48" blockId="15.[151,1437,151,1040]" box="[197,385,476,500]" pageId="15" pageNumber="56">
|
||
<emphasis id="B9620F13D83C214C06E44D82FE98FE48" bold="true" box="[197,260,476,500]" pageId="15" pageNumber="56">Host.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C808AD83C214C06E44E5EFD99FBAC" pageId="15" pageNumber="56" type="discussion">
|
||
<paragraph id="8BA9D301D83C214C06E44E5EFDD2FC8B" blockId="15.[151,1437,151,1040]" pageId="15" pageNumber="56">
|
||
<emphasis id="B9620F13D83C214C06E44E5EFEA5FDAB" bold="true" box="[197,313,512,536]" pageId="15" pageNumber="56">Remarks.</emphasis>
|
||
This species has not been treated since its description (
|
||
<bibRefCitation id="EF87AEF0D83C214C05E04DA1FBFCFDAB" author="Cresson" box="[961,1120,511,535]" pageId="15" pageNumber="56" refString="Cresson, E. T. (1902) Descriptions of some Brazilian Mutilla. Transactions of the American Entomological Society, 28, 1 - 82." type="journal article" year="1902">Cresson 1902</bibRefCitation>
|
||
) and was still in the genus
|
||
<taxonomicName id="4C16A882D83C214C06B64E7AFE01FD80" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,413,548,572]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C06B64E7AFF76FD80" box="[151,234,548,572]" italics="true" pageId="15" pageNumber="56">Mutilla</emphasis>
|
||
<bibRefCitation id="EF87AEF0D83C214C06D14E7AFE01FD80" author="Linnaeus" box="[240,413,548,572]" pageId="15" pageNumber="56" refString="Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Editio Decima Reformata. Laurentii Salvii, Holmiae [= Stockholm]. [4] + 1 - 823 + [5]." type="journal article" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
</taxonomicName>
|
||
until this paper (
|
||
<bibRefCitation id="EF87AEF0D83C214C047A4E7AFC84FD80" author="Nonveiller" box="[603,792,548,572]" pageId="15" pageNumber="56" refString="Nonveiller, G. (1990) Catalogue of the Mutillidae, Myrmosidae and Bradynobaenidae of the Neotropical Region including Mexico (Insecta, Hymenoptera). Hymenopterorum Catalogus (Nova Editio), 18. SPB Academic Publishing, Den Haag, pp. 1 - 150." type="journal article" year="1990">Nonveiller 1990</bibRefCitation>
|
||
). The pygidial sculpture (
|
||
<figureCitation id="132DCF84D83C214C02164E7AFB11FD80" box="[1079,1165,548,572]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="15" pageNumber="56">Figs 13</figureCitation>
|
||
,
|
||
<figureCitation id="132DCF84D83C214C02BB4E7AFB25FD80" box="[1178,1209,548,572]" captionStart="FIGURES 19 – 38" captionStartId="10.[151,264,1672,1695]" captionTargetBox="[176,1419,215,1644]" captionTargetId="figure@10.[157,1429,193,1651]" captionTargetPageId="10" captionText="FIGURES 19 – 38. Tobantilla species. 19 – 34. Male genitalia: dorsal view (left), ventral view (middle), lateral view of penis valve (above), and lateral view with penis valve removed (right). 19 – 22. T. andrikos sp. nov.; 23 – 26. T. drosos sp. nov.; 27 – 30. T. ephemeros sp. nov.; 31 – 34. T. xouthos sp. nov. 35 – 36. Male clypeus. 28. T. andrikos sp. nov.; 29. T. ephemeros sp. nov. 37, 38. Female pygidium (T 6). 37. T. frigidula (Cresson); 38. T. kolasma sp. nov." httpUri="https://zenodo.org/record/278962/files/figure.png" pageId="15" pageNumber="56">37</figureCitation>
|
||
), coloration (
|
||
<figureCitation id="132DCF84D83C214C036F4E7AFA0BFD80" box="[1358,1431,548,572]" captionStart="FIGURES 1 – 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="15" pageNumber="56">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="132DCF84D83C214C06B64E19FF3AFDE3" box="[151,166,583,607]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="15" pageNumber="56">9</figureCitation>
|
||
) and distribution in northern
|
||
<collectingCountry id="F3019391D83C214C07D24E19FDABFDE3" box="[499,567,583,607]" name="Brazil" pageId="15" pageNumber="56">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="132DCF84D83C214C04694E19FD06FDE3" box="[584,666,583,607]" captionStart="FIGURE 39" captionStartId="12.[151,250,1315,1338]" captionTargetBox="[195,1400,384,1293]" captionTargetId="figure@12.[180,1407,370,1294]" captionTargetPageId="12" captionText="FIGURE 39. Distribution of Tobantilla species." httpUri="https://zenodo.org/record/278963/files/figure.png" pageId="15" pageNumber="56">Fig. 39</figureCitation>
|
||
) likely contributed to the difficulty in placing this species in
|
||
<taxonomicName id="4C16A882D83C214C036C4E16FF58FD38" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C036C4E16FF58FD38" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
. The two previously recognized
|
||
<taxonomicName id="4C16A882D83C214C041E4E32FD2DFD38" box="[575,689,620,644]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C041E4E32FD2DFD38" box="[575,689,620,644]" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
species have a well defined pygidial plate and were only known from
|
||
<collectingCountry id="F3019391D83C214C06F54ED1FED8FD1B" box="[212,324,655,679]" name="Argentina" pageId="15" pageNumber="56">Argentina</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF87AEF0D83C214C07734ED1FE7CFD1B" author="Casal" box="[338,480,655,679]" pageId="15" pageNumber="56" refString="Casal, O. H. (1965 a) Tobantilla montonera genero y especie nuevos de Sphaeropthalminae (Hymenoptera: Mutillidae). Revista de la Sociedad Mexicana de Historia Natural, 25, 209 - 215." type="journal article" year="1965" yearSuffix="a">Casal 1965a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF87AEF0D83C214C07CA4ED1FDBBFD1B" author="Casal" box="[491,551,655,679]" pageId="15" pageNumber="56" refString="Casal, O. H. (1969) La segunda especie de Tobantilla Casal, 1964 (Hymenoptera: Mutillidae). Physis, 29, 101 - 102." type="journal article" year="1969">1969</bibRefCitation>
|
||
). Like the other
|
||
<taxonomicName id="4C16A882D83C214C04FC4ECEFCD3FD14" box="[733,847,656,680]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C04FC4ECEFCD3FD14" box="[733,847,656,680]" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
females, the mesosoma is flattened dorsally, appearing virtually concave anterior to the scutellar scale. The scutellar and propodeal armature of this species is nearly identical to that of
|
||
<taxonomicName id="4C16A882D83C214C074D4E87FE7FFD4C" box="[364,483,728,752]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="kolasma">
|
||
<emphasis id="B9620F13D83C214C074D4E87FE7FFD4C" box="[364,483,728,752]" italics="true" pageId="15" pageNumber="56">T. kolasma</emphasis>
|
||
</taxonomicName>
|
||
, a species that matches the traditional diagnostic features for
|
||
<taxonomicName id="4C16A882D83C214C02BD4E86FA92FD4C" box="[1180,1294,728,752]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C02BD4E86FA92FD4C" box="[1180,1294,728,752]" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
. The petiole is apparently more disciform than other
|
||
<taxonomicName id="4C16A882D83C214C04434EA2FD48FCA8" box="[610,724,764,788]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C04434EA2FD48FCA8" box="[610,724,764,788]" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
, but is weakly constricted apically and is therefore considered petiolate, like other
|
||
<taxonomicName id="4C16A882D83C214C07564F7EFE75FC84" box="[375,489,800,824]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C07564F7EFE75FC84" box="[375,489,800,824]" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
females.
|
||
</paragraph>
|
||
<paragraph id="8BA9D301D83C214C06E44F1AFD99FBAC" blockId="15.[151,1437,151,1040]" pageId="15" pageNumber="56">
|
||
<bibRefCitation id="EF87AEF0D83C214C06E44F1AFEECFCE0" author="Cresson" box="[197,368,836,860]" pageId="15" pageNumber="56" refString="Cresson, E. T. (1902) Descriptions of some Brazilian Mutilla. Transactions of the American Entomological Society, 28, 1 - 82." type="journal article" year="1902">Cresson (1902)</bibRefCitation>
|
||
noted the presence of a “rather long longitudinal groove” on each side of the propodeum. These are simply large reticulations situated immediately posteromesal to each propodeal spiracle. In the
|
||
<typeStatus id="54AD6DA3D83C214C03234F39FAFBFCC3" box="[1282,1383,871,895]" pageId="15" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
, this reticulation is apparently undivided and elongate, but there is significant variation in the punctation of this region. In the specimen from Rondônia, it bears three contiguous setigerous reticulations that are slightly less differentiated from each other than from the surrounding reticulations. This trait does not seem much different from what is seen in other
|
||
<taxonomicName id="4C16A882D83C214C070C4FA6FE03FBAC" box="[301,415,1016,1040]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9620F13D83C214C070C4FA6FE03FBAC" box="[301,415,1016,1040]" italics="true" pageId="15" pageNumber="56">Tobantilla</emphasis>
|
||
</taxonomicName>
|
||
females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |