239 lines
32 KiB
XML
239 lines
32 KiB
XML
<document id="0B89DA5EACF3E5B4EDCF858DF780693A" ID-CLB-Dataset="7045" ID-DOI="10.11646/zootaxa.4965.1.7" ID-GBIF-Dataset="0d69081f-0cf4-4be6-92a6-5a785e5ee98e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4723237" ID-ZooBank="1F5BE28F-C8B5-47FF-8A3F-68B5BC40BD6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619523548262" checkinUser="plazi" docAuthor="Alekseev, Vitalii Igorevich & Bukejs, Andris" docDate="2021" docId="03B587D0D9194D6406B0E2ECFB1FDE36" docLanguage="en" docName="zootaxa.4965.1.7.pdf" docOrigin="Zootaxa 4965 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Madelinia capillata Alekseev & Bukejs 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="154" masterDocId="FF8CFFA8D9104D680627E401FFB3DC0B" masterDocTitle="Fossil Melandryidae (Coleoptera: Tenebrionoidea) from Eocene Baltic amber of the Sambian peninsula: new genus, six new species, new records and key to described taxa" masterLastPageNumber="166" masterPageNumber="142" pageNumber="151" updateTime="1698937681551" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B08C5643A5C07F54B613963A9F9151E3">Fossil Melandryidae (Coleoptera: Tenebrionoidea) from Eocene Baltic amber of the Sambian peninsula: new genus, six new species, new records and key to described taxa</mods:title>
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<mods:namePart id="A784BD6CFCE6F2D4A2A42118CDE57281">Alekseev, Vitalii Igorevich</mods:namePart>
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<mods:nameIdentifier id="7BFC8A3699721D63082F5A3ECA15F845" type="ORCID">0000-0003-4390-5443</mods:nameIdentifier>
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<mods:affiliation id="4D7F73514B8EBEF29EC7556C036D93EB">Shirshov Institute of Oceanology, Russian Academy of Sciences, Nahimovskiy prospekt 36, 117997 Moscow, Russia. alekseew 0802 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4390 - 5443 & Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia</mods:affiliation>
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<mods:nameIdentifier id="26979670B32F519988A21D41DE5BE733" type="email">alekseew0802@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="1183951E7090EBEF19A6839B30434C4A">Bukejs, Andris</mods:namePart>
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<mods:affiliation id="63E284FB4B07020B87CA3665F6FBC792">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas 13, Daugavpils LV- 5401, Latvia. carabidae @ inbox. lv; https: // orcid. org / 0000 - 0001 - 7165 - 3023</mods:affiliation>
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<mods:nameIdentifier id="D9F897AA356F5D814E8179233BFAC122" type="email">carabidae@inbox.lv</mods:nameIdentifier>
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<mods:title id="DB73FCDB70119C78264B2E5C35424385">Zootaxa</mods:title>
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<mods:date id="7F33B888041E9D1139C5B13C5B91CFD6">2021</mods:date>
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<mods:number id="40EF5250C466AA27EEAD972B7B1F75CD">2021-04-27</mods:number>
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<mods:number id="1D8E910AA3E945894E2C96E63651FA60">4965</mods:number>
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<treatment id="03B587D0D9194D6406B0E2ECFB1FDE36" ID-DOI="http://doi.org/10.5281/zenodo.4723245" ID-GBIF-Taxon="181882189" ID-Zenodo-Dep="4723245" LSID="urn:lsid:plazi:treatment:03B587D0D9194D6406B0E2ECFB1FDE36" httpUri="http://treatment.plazi.org/id/03B587D0D9194D6406B0E2ECFB1FDE36" lastPageId="12" lastPageNumber="154" pageId="9" pageNumber="151">
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<subSubSection id="C306654DD9194D6106B0E2ECFE5BDB0C" box="[151,488,1773,1800]" pageId="9" pageNumber="151" type="nomenclature">
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<paragraph id="8BA336C6D9194D6106B0E2ECFE5BDB0C" blockId="9.[151,488,1773,1836]" box="[151,488,1773,1800]" pageId="9" pageNumber="151">
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<heading id="D0EB81AAD9194D6106B0E2ECFE5BDB0C" bold="true" box="[151,488,1773,1800]" fontSize="11" level="1" pageId="9" pageNumber="151" reason="1">
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<emphasis id="B968EAD4D9194D6106B0E2ECFE5BDB0C" bold="true" box="[151,488,1773,1800]" pageId="9" pageNumber="151">
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<taxonomicName id="4C1C4D45D9194D6106B0E2ECFE31DB03" authority="Alekseev & Bukejs, 2021" authorityName="Alekseev & Bukejs" authorityYear="2021" box="[151,386,1773,1800]" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="151" phylum="Arthropoda" rank="species" species="capillata" status="sp. nov.">
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<emphasis id="B968EAD4D9194D6106B0E2ECFE31DB03" bold="true" box="[151,386,1773,1800]" italics="true" pageId="9" pageNumber="151">Madelinia capillata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57AFD9194D6107ADE2ECFE5BDB0C" box="[394,488,1773,1799]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C306654DD9194D6106B0E310FEE6DB20" box="[151,341,1809,1836]" pageId="9" pageNumber="151" type="description">
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<paragraph id="8BA336C6D9194D6106B0E310FEE6DB20" blockId="9.[151,488,1773,1836]" box="[151,341,1809,1836]" pageId="9" pageNumber="151">
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(
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<figureCitation id="13272A43D9194D6106B8E310FF5CDB27" box="[159,239,1809,1836]" captionStart="FIGURE 5" captionStartId="11.[151,250,1889,1913]" captionTargetBox="[151,1436,181,1847]" captionTargetId="figure-20@11.[151,1436,181,1865]" captionTargetPageId="11" captionText="FIGURE 5. Madelinia capillata sp. nov., holotype, No 6708 [MAIG], habitus:A—dorsal view; B—left lateral view. Scale bars represent 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4723253" httpUri="https://zenodo.org/record/4723253/files/figure.png" pageId="9" pageNumber="151">Figs. 5</figureCitation>
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,
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<figureCitation id="13272A43D9194D6106DCE310FEAEDB20" box="[251,285,1809,1835]" captionStart="FIGURE 8" captionStartId="16.[151,250,1579,1603]" captionTargetBox="[153,1433,1137,1556]" captionTargetId="figure-408@16.[153,1434,1137,1556]" captionTargetPageId="16" captionText="FIGURE 8. Apical portion of aedeagus in ventral view: A—Electroxylita chronographica gen. et sp. nov.; B—Phloiotrya inmarinata sp. nov.; C—Madelinia capillata sp. nov.; D—Symphora pollocki sp. nov. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.4723265" httpUri="https://zenodo.org/record/4723265/files/figure.png" pageId="9" pageNumber="151">8C</figureCitation>
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,
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<figureCitation id="13272A43D9194D61070EE310FEFEDB20" box="[297,333,1809,1835]" captionStart="FIGURE 9" captionStartId="17.[151,250,1889,1913]" captionTargetBox="[178,1422,192,1833]" captionTargetId="figure-20@17.[151,1436,181,1865]" captionTargetPageId="17" captionText="FIGURE 9. Reconstructions of new fossil Melandryidae from Baltic amber:A—Electroxylita chronographica gen. et sp. nov.; B—Phloiotrya inmarinata sp. nov.; C—Microscapha kugelanni sp. nov.; D—Madelinia capillata sp. nov.; E—Symphora glaesonauta sp. nov. Not reproduced to the same scale, punctuation and setation are omitted." figureDoi="http://doi.org/10.5281/zenodo.4723267" httpUri="https://zenodo.org/record/4723267/files/figure.png" pageId="9" pageNumber="151">9D</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C306654DD9194D6206B0E358FF5ADD6E" lastPageId="10" lastPageNumber="152" pageId="9" pageNumber="151" type="materials_examined">
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<paragraph id="8BA336C6D9194D6206B0E358FE14DCDE" blockId="9.[151,1437,1881,2016]" lastBlockId="10.[151,1437,151,2014]" lastPageId="10" lastPageNumber="152" pageId="9" pageNumber="151">
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<emphasis id="B968EAD4D9194D6106B0E358FEF5DB78" bold="true" box="[151,326,1881,1907]" pageId="9" pageNumber="151">Type material.</emphasis>
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<materialsCitation id="3B743C9BD9194D62076AE358FE14DCDE" ID-GBIF-Occurrence="3107842308" collectionCode="MAIG" collectorName="Jonas Damzen" lastPageId="10" lastPageNumber="152" pageId="9" pageNumber="151" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54A78864D9194D61076AE358FE09DB78" box="[333,442,1881,1907]" pageId="9" pageNumber="151" type="holotype">Holotype</typeStatus>
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||
: No 6708 [
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<collectionCode id="ED0DAE03D9194D610412E358FD36DB7F" box="[565,645,1881,1908]" pageId="9" pageNumber="151">MAIG</collectionCode>
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||
] (ex. coll.
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||
<collectorName id="26E95310D9194D6104D9E35BFC12DB7F" box="[766,929,1882,1908]" pageId="9" pageNumber="151">Jonas Damzen</collectorName>
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||
JDC 8368), “
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<typeStatus id="54A78864D9194D61021BE358FB14DB78" box="[1084,1191,1881,1907]" pageId="9" pageNumber="151" type="holotype">Holotype</typeStatus>
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||
/
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||
<taxonomicName id="4C1C4D45D9194D610299E358FA2EDB78" authority="Alekseev & Bukejs, 2021" authorityName="Alekseev & Bukejs" authorityYear="2021" box="[1214,1437,1881,1907]" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="151" phylum="Arthropoda" rank="species" species="capillata" status="sp. nov.">
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<emphasis id="B968EAD4D9194D610299E358FA2EDB78" box="[1214,1437,1881,1907]" italics="true" pageId="9" pageNumber="151">Madelinia capillata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57AFD9194D6106B0E37FFF5CDB93" box="[151,239,1918,1944]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
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/ Alekseev et Bukejs des. 2021” [red printed label]; adult, male.
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<collectorName id="26E95310D9194D6105EEE37FFC58DB93" box="[969,1003,1918,1944]" pageId="9" pageNumber="151">An</collectorName>
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almost complete beetle (right antenna missing) with partially exposed metathoracic wings and apical portion of aedeagus; included in a transparent, yellow amber piece with approximate dimensions of
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<quantity id="4CE49B23D9194D6104EAE3C4FCAFDBEB" box="[717,796,1989,2016]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="151" unit="mm" value="21.0">21 mm</quantity>
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×
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<quantity id="4CE49B23D9194D61051CE3C4FC38DBEB" box="[827,907,1989,2016]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="151" unit="mm" value="12.0">12 mm</quantity>
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and a maximum thickness of
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<quantity id="4CE49B23D9194D6102F8E3C4FA90DBEB" box="[1247,1315,1989,2016]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="151" unit="mm" value="8.0">8 mm</quantity>
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; preserved without supplementary fixation.
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<collectorName id="26E95310D91A4D62042FE496FDD1DCBA" box="[520,610,151,177]" pageId="10" pageNumber="152">Organic</collectorName>
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syninclusions: one
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<collectorName id="26E95310D91A4D620564E496FC79DCBA" box="[835,970,151,177]" pageId="10" pageNumber="152">Nematocera</collectorName>
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(
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<taxonomicName id="4C1C4D45D91A4D6205FDE496FB81DCB9" box="[986,1074,151,178]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="152" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
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) specimen and some small stellate fagacean trichomes.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA336C6D91A4D6206E0E4DEFC42DD16" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">
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<emphasis id="B968EAD4D91A4D6206E0E4DEFEDDDCF2" bold="true" box="[199,366,223,249]" pageId="10" pageNumber="152">
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<typeStatus id="54A78864D91A4D6206E0E4DEFEB1DCF2" box="[199,258,223,249]" pageId="10" pageNumber="152">Type</typeStatus>
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stratum.
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</emphasis>
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Baltic amber from Eocene amber-bearing blue Earth layers, mostly Bartonian age is interpreted for the extinct central European resin-producing forests (
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<bibRefCitation id="EF8D4B37D91A4D620535E502FC56DD16" author="Bukejs, A. & Alekseev, V. I. & Pollock, D. A." box="[786,997,259,285]" pageId="10" pageNumber="152" pagination="261 - 273" refId="ref15937" refString="Bukejs, A., Alekseev, V. I. & Pollock, D. A. (2019) Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa, 4664 (2), 261 - 273. https: // doi. org / 10.11646 / zootaxa. 4664.2.8" type="journal article" year="2019">
|
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Bukejs
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<emphasis id="B968EAD4D91A4D62054EE502FC29DD16" box="[873,922,259,285]" italics="true" pageId="10" pageNumber="152">et al</emphasis>
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||
. 2019
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||
</bibRefCitation>
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).
|
||
</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E526FF5ADD6E" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">
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<materialsCitation id="3B743C9BD91A4D6206E0E526FF56DD6E" ID-GBIF-Occurrence="3107842305" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="10" pageNumber="152" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
|
||
<emphasis id="B968EAD4D91A4D6206E0E526FED6DD4B" bold="true" box="[199,357,294,321]" pageId="10" pageNumber="152">
|
||
<typeStatus id="54A78864D91A4D6206E0E526FEB1DD4A" box="[199,258,295,321]" pageId="10" pageNumber="152">Type</typeStatus>
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||
locality.
|
||
</emphasis>
|
||
<location id="8EC3601DD91A4D62074CE526FE63DD4A" LSID="urn:lsid:plazi:treatment:03B587D0D9194D6406B0E2ECFB1FDE36:8EC3601DD91A4D62074CE526FE63DD4A" box="[363,464,295,321]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="10" pageNumber="152" stateProvince="Kaliningrad">Yantarny</location>
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settlement (formerly Palmnicken),
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<collectingMunicipality id="6BC7ACBCD91A4D62057CE526FC0EDD4A" box="[859,957,295,321]" pageId="10" pageNumber="152">Sambian</collectingMunicipality>
|
||
(Samland)
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||
<collectingCounty id="62C24E4AD91A4D620219E526FB1DDD4A" box="[1086,1198,295,321]" pageId="10" pageNumber="152">Peninsula</collectingCounty>
|
||
,
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<collectingRegion id="49D8F824D91A4D62029FE526FA2BDD4A" box="[1208,1432,295,321]" country="Russia" name="Kaliningrad" pageId="10" pageNumber="152">Kaliningrad Region</collectingRegion>
|
||
,
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<collectingCountry id="F30B7656D91A4D6206B0E54AFF56DD6E" box="[151,229,331,357]" name="Russia" pageId="10" pageNumber="152">Russia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306654DD91A4D6406E0E56FFD20DCBA" lastPageId="12" lastPageNumber="154" pageId="10" pageNumber="152" type="description">
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<paragraph id="8BA336C6D91A4D6206E0E56FFB70DDA6" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">
|
||
<emphasis id="B968EAD4D91A4D6206E0E56FFEEADD83" bold="true" box="[199,345,366,392]" pageId="10" pageNumber="152">Description.</emphasis>
|
||
Measurements: body length about
|
||
<quantity id="4CE49B23D91A4D6204CFE56EFCF2DD82" box="[744,833,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="152" unit="mm" value="4.2">4.2 mm</quantity>
|
||
, body maximum width
|
||
<quantity id="4CE49B23D91A4D620277E56EFB19DD82" box="[1104,1194,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="152" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; pronotum length
|
||
<quantity id="4CE49B23D91A4D62035EE56EFF76DDA6" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="152" unit="mm" value="0.9">0.9 mm</quantity>
|
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, pronotum maximum width
|
||
<quantity id="4CE49B23D91A4D620423E592FDEEDDA6" box="[516,605,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="152" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; elytra length
|
||
<quantity id="4CE49B23D91A4D6204DBE592FCE7DDA6" box="[764,852,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="152" unit="mm" value="3.2">3.2 mm</quantity>
|
||
, elytra maximum width
|
||
<quantity id="4CE49B23D91A4D620241E592FB0CDDA6" box="[1126,1215,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="152" unit="mm" value="1.2">1.2 mm</quantity>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E5B6FC23DE12" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">Body oblong, flattened dorsally and slightly convex ventrally; brown with antennae (except scape) and legs paler. Body pubescence consists of: (1) fine, rather dense, almost recumbent setation dorsally and ventrally, and (2) slightly longer, sparse and erect setae on head, pronotum and elytra.</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E622FE88D81A" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">Head prognathous, well visible in dorsal view, densely covered with fine punctuation; frons flat; vertex slightly convex; genae short, about 0.3× as long as horizontal diameter of one eye. Fronto-clypeal suture apparently present but poorly discernible. Compound eyes rather large, entire, slightly prominent, oval, vertical diameter about 1.5× longer than horizontal diameter; facets distinct and fine, without intrafacetal setae; interocular distance at upper margin of eyes about 0.75× as vertical diameter of one eye. Maxillary palpi rather long, about 0.4× as long as antenna; with 4 palpomeres: palpomere 1 minute, triangular, nearly as long as wide, palpomere 2 elongate triangular, with slightly oblique apical margin, about 1.8× longer than maximum wide, palpomere 3 shorter than palpomere 2 (poorly visible in studied specimen), palpomere 4 large, cultriform, about 3.3× longer than wide; ratio of relative lengths of maxillary palpomeres 1–4 = to 5:14:?:30. Antennae with 11 antennomeres, filiform, inserted between and close to inner margin of eyes, pubescent, rather short, reaching posteriorly to humeri; scape cylindrical, 1.8× longer than wide; pedicle subcylindrical; antennomeres 3–10 conical, elongate, subequal in size, about 1.7–2.1× longer than maximum wide; antennomere 11 narrowly ovoid with pointed apex, 2.2× longer than maximum wide, and about 1.3× longer than antennomere 10; ratio of relative lengths of antennomeres 1–11 = 16:15:17:17:17:17:1 7:17:17:17:22.</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E01AFC24D95E" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">Pronotum slightly transverse, 1.2× wider than long, widest at middle, base closely associated with elytral base; densely covered with small punctuation, distance between punctures equal to 0.5–1.0× diameter of one puncture; disc convex, with shallow, rounded impression laterally. Anterior pronotal margin straight (in dorsal view); posterior margin slightly narrower than elytral anterior margin, weakly bisinuated, markedly protruding medially, distinctly margined; lateral margins rounded, slightly narrowed anteriad and more distinct posteriad. Lateral pronotal carina present, distinct in basal one-third of pronotal length. Anterior pronotal angles rounded; posterior angles acute, slightly protruding posteriad. Basal impressions triangular with widely rounded apex, deep, reaching about one-fourth of pronotal length. Prohypomera convex, densely covered with small punctuation. Scutellar shield rather large, subpentagonal, slightly elongate, about 1.1× longer than wide.</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E15EFB9CDA7E" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">Elytra narrowly ovate, elongate, about 2.7× longer than wide, widest in posterior one-third, almost parallelsided in anterior two-third, gradually narrowed in posterior one-third; humeri distinct, prominent; elytral apices rounded separately; elytral punctuation irregular, small and dense, distance between punctures lesser than diameter of one puncture, punctuation slightly sparser and smoother in posterior portion. Hind wings present, macropterous. Epipleura narrow, widest basally and gradually narrowed posteriad, reaching posterior margin of abdominal ventrite 1. Metepisternum rather wide, slightly dilated anteriorly, with subparallel lateral sides, about 4.7× longer than maximum wide, densely covered with small punctuation. Metaventrite with convex disc, densely covered with small punctures, distance between punctures lesser than diameter of one puncture.</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E27EFCBFDB7A" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">Legs slender, long, with dense, fine punctuation. Procoxae contiguous, conical; mesocoxae roundish, apparently very narrowly separated; metacoxae elongate oval, transverse, narrowly separated. Femora clavate, flattened. Tibiae subcylindrical, almost straight, slightly dilated apically, subequal in length to femora, with two short spurs of equal length and fringe of spinulae apically; metatibial spurs about 0.5× as long as metatibial width at apex. Tarsi long, metatarsus nearly as long as metatibia; tarsal formula 5-5-4; penultimate tarsomere of all legs deeply bilobed; metatarsomere 1 not shorter than one-half of metatibia, ratio of relative lengths of metatarsomeres 1–4 = 20:10:5:7. Pretarsal claws simple, strongly divergent, symmetrical.</paragraph>
|
||
<paragraph id="8BA336C6D91A4D6206E0E37AFE77DBD5" blockId="10.[151,1437,151,2014]" pageId="10" pageNumber="152">Abdomen with five visible ventrites, densely covered with small punctuation, distance between punctures lesser than diameter of one puncture; ventrite 5 with rounded apical margin; ratio of relative lengths of ventrites 1–5 = 20:17:15:11:12 (medially).</paragraph>
|
||
<caption id="DF63664ED91B4D6306B0E360FEFCDB97" ID-DOI="http://doi.org/10.5281/zenodo.4723253" ID-Zenodo-Dep="4723253" httpUri="https://zenodo.org/record/4723253/files/figure.png" pageId="11" pageNumber="153" startId="11.[151,250,1889,1913]" targetBox="[151,1436,181,1847]" targetPageId="11">
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||
<paragraph id="8BA336C6D91B4D6306B0E360FEFCDB97" blockId="11.[151,1436,1889,1949]" pageId="11" pageNumber="153">
|
||
<emphasis id="B968EAD4D91B4D6306B0E360FEA1DB72" bold="true" box="[151,274,1889,1913]" pageId="11" pageNumber="153">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C1C4D45D91B4D63073FE360FE6CDB72" authority="Alekseev & Bukejs, 2021" authorityName="Alekseev & Bukejs" authorityYear="2021" box="[280,479,1889,1913]" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="153" phylum="Arthropoda" rank="species" species="capillata" status="sp. nov.">
|
||
<emphasis id="B968EAD4D91B4D63073FE360FE6CDB72" box="[280,479,1889,1913]" italics="true" pageId="11" pageNumber="153">Madelinia capillata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EAD4D91B4D6307C2E360FD86DB72" bold="true" box="[485,565,1889,1913]" pageId="11" pageNumber="153">
|
||
<taxonomicNameLabel id="A25B57AFD91B4D6307C2E360FD86DB72" box="[485,565,1889,1913]" pageId="11" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No 6708 [MAIG], habitus: A—dorsal view; B—left lateral view. Scale bars represent 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA336C6D91C4D6406E0E496FD20DCBA" blockId="12.[151,1437,151,573]" box="[199,659,151,177]" pageId="12" pageNumber="154">
|
||
Apical portion of aedeagus as in
|
||
<figureCitation id="13272A43D91C4D64041FE496FD3CDCBA" box="[568,655,151,177]" captionStart="FIGURE 8" captionStartId="16.[151,250,1579,1603]" captionTargetBox="[153,1433,1137,1556]" captionTargetId="figure-408@16.[153,1434,1137,1556]" captionTargetPageId="16" captionText="FIGURE 8. Apical portion of aedeagus in ventral view: A—Electroxylita chronographica gen. et sp. nov.; B—Phloiotrya inmarinata sp. nov.; C—Madelinia capillata sp. nov.; D—Symphora pollocki sp. nov. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.4723265" httpUri="https://zenodo.org/record/4723265/files/figure.png" pageId="12" pageNumber="154">Fig. 8C</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306654DD91C4D6406E0E4BAFCAEDCF2" pageId="12" pageNumber="154" type="etymology">
|
||
<paragraph id="8BA336C6D91C4D6406E0E4BAFCAEDCF2" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="154">
|
||
<emphasis id="B968EAD4D91C4D6406E0E4BAFEFBDCDE" bold="true" box="[199,328,187,213]" pageId="12" pageNumber="154">Etymology</emphasis>
|
||
The epithet of this new species (Latin adjective
|
||
<emphasis id="B968EAD4D91C4D640548E4BAFC6EDCDE" box="[879,989,187,213]" italics="true" pageId="12" pageNumber="154">capillatus</emphasis>
|
||
, meaning “hairy, long-haired, old-fashioned”) refers to its long, conspicuous dorsal pubescence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306654DD91C4D6406E0E503FB1FDE36" pageId="12" pageNumber="154" type="diagnosis">
|
||
<paragraph id="8BA336C6D91C4D6406E0E503FE8DDDDA" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="154">
|
||
<emphasis id="B968EAD4D91C4D6406E0E503FE62DD17" bold="true" box="[199,465,258,284]" pageId="12" pageNumber="154">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C1C4D45D91C4D6407FCE502FD0EDD16" authority="Alekseev & Bukejs, 2021" authorityName="Alekseev & Bukejs" authorityYear="2021" box="[475,701,259,285]" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="154" phylum="Arthropoda" rank="species" species="capillata" status="sp. nov.">
|
||
<emphasis id="B968EAD4D91C4D6407FCE502FD0EDD16" box="[475,701,259,285]" italics="true" pageId="12" pageNumber="154">Madelinia capillata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EAD4D91C4D6404E1E502FC90DD16" bold="true" box="[710,803,259,285]" pageId="12" pageNumber="154">
|
||
<taxonomicNameLabel id="A25B57AFD91C4D6404E1E502FC90DD16" box="[710,803,259,285]" pageId="12" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C1C4D45D91C4D6405E6E502FF7CDD4A" authority="Alekseev et Pankowski, 2020" authorityName="Alekseev et Pankowski" authorityYear="2020" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="154" phylum="Arthropoda" rank="species" species="gedanoposita">
|
||
<emphasis id="B968EAD4D91C4D6405E6E502FB32DD16" box="[961,1153,259,285]" italics="true" pageId="12" pageNumber="154">M. gedanoposita</emphasis>
|
||
<bibRefCitation id="EF8D4B37D91C4D6402AEE502FF7CDD4A" author="Alekseev, V. I. & Pankowski, M." pageId="12" pageNumber="154" pagination="281 - 289" refId="ref15817" refString="Alekseev, V. I. & Pankowski, M. (2020) The first new genus of the tribe Hypulini Seidlitz (Coleoptera: Melandryidae) described from late Eocene Baltic amber. Zootaxa, 4869 (2), 281 - 289. https: // doi. org / 10.11646 / zootaxa. 4869.2.8" type="journal article" year="2020">
|
||
Alekseev et Pankowski,
|
||
<quantity id="4CE49B23D91C4D6406B0E526FF7CDD4A" box="[151,207,295,321]" metricMagnitude="1" metricUnit="m" metricValue="5.1308" pageId="12" pageNumber="154" unit="in" value="2020.0">2020</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in: (1) shape of pronotum, lateral pronotal margins rounded, narrowed anteriad and posteriad; (2) distinctly longer and conspicuous pubescence of the dorsum; (3) slightly elongate scutellar shield (transverse in
|
||
<taxonomicName id="4C1C4D45D91C4D640300E54AFF5ADD83" authorityName="Alekseev et Pankowski" authorityYear="2020" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="154" phylum="Arthropoda" rank="species" species="gedanoposita">
|
||
<emphasis id="B968EAD4D91C4D640300E54AFF5ADD83" italics="true" pageId="12" pageNumber="154">M. gedanoposita</emphasis>
|
||
</taxonomicName>
|
||
); (4) comparatively longer metatarsomere 2 (the ratio of metatarsomeres
|
||
<quantity id="4CE49B23D91C4D640212E56EFBCDDD82" box="[1077,1150,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="12" pageNumber="154" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">1–2 in</quantity>
|
||
the new species consists 2:1, whereas the ratio in
|
||
<taxonomicName id="4C1C4D45D91C4D640788E592FDDFDDA6" authorityName="Alekseev et Pankowski" authorityYear="2020" box="[431,620,403,429]" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="154" phylum="Arthropoda" rank="species" species="gedanoposita">
|
||
<emphasis id="B968EAD4D91C4D640788E592FDDFDDA6" box="[431,620,403,429]" italics="true" pageId="12" pageNumber="154">M. gedanoposita</emphasis>
|
||
</taxonomicName>
|
||
consists 4:1); and (5) smaller body (
|
||
<quantity id="4CE49B23D91C4D64022EE592FBD2DDA6" box="[1033,1121,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="154" unit="mm" value="4.2">4.2 mm</quantity>
|
||
in contrast to
|
||
<quantity id="4CE49B23D91C4D640324E593FAEADDA6" box="[1283,1369,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="154" unit="mm" value="5.8">5.8 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1C4D45D91C4D640359E592FE9DDDDA" authorityName="Alekseev et Pankowski" authorityYear="2020" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="154" phylum="Arthropoda" rank="species" species="gedanoposita">
|
||
<emphasis id="B968EAD4D91C4D640359E592FE9DDDDA" italics="true" pageId="12" pageNumber="154">M. gedanoposita</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336C6D91C4D6406E0E5DAFB1FDE36" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="154">
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||
<emphasis id="B968EAD4D91C4D6406E0E5DAFEB5DDFE" bold="true" box="[199,262,475,501]" pageId="12" pageNumber="154">Note.</emphasis>
|
||
The shape of pronotum and the body size can vary within certain limits in extant representatives of the tribe. The main diagnostic characters for species limitation within the extinct genus
|
||
<taxonomicName id="4C1C4D45D91C4D640278E5FEFB61DE12" authorityName="Alekseev et Pankowski" authorityYear="2020" box="[1119,1234,511,537]" class="Insecta" family="Melandryidae" genus="Madelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="154" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B968EAD4D91C4D640278E5FEFB61DE12" box="[1119,1234,511,537]" italics="true" pageId="12" pageNumber="154">Madelinia</emphasis>
|
||
</taxonomicName>
|
||
is considered the pubescence, the shape of scutellar shield and the comparative lenghts of metatarsomeres 1–2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |