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<document id="F2BB66CAC6D357D842FAE8CC1129758D" ID-DOI="10.1643/ci-19-304" ID-ISSN="2766-1520" ID-Zenodo-Dep="13628562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1722898778496" checkinUser="felipe" docAuthor="Ciccotto, Patrick J. &amp; Page, Lawrence M." docDate="2020" docId="03B587CBFFDB0C581E7785B688EC23E5" docLanguage="en" docName="Copeia.108.3.485-502.pdf" docOrigin="Copeia 108 (3)" docSource="http://dx.doi.org/10.1643/ci-19-304" docStyle="DocumentStyle:FE116AF270F49333EC2F8F2E31B37DD9.1:Copeia.2011-2019.journal_article.1cover.type1" docStyleId="FE116AF270F49333EC2F8F2E31B37DD9" docStyleName="Copeia.2011-2019.journal_article.1cover.type1" docStyleVersion="1" docTitle="Henicorhynchus caudimaculatus" docType="treatment" docVersion="4" lastPageNumber="502" masterDocId="FF8CFFB3FFDE0C511D2C831B8D252366" masterDocTitle="Revision of the Genus Henicorhynchus, with a Revised Diagnosis of Gymnostomus (Cyprinidae: Labeoninae)" masterLastPageNumber="502" masterPageNumber="485" pageNumber="485" updateTime="1725282116278" updateUser="ExternalLinkService">
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<mods:title id="6AF2E024562D97B5B104CC0ED4E723DD">Revision of the Genus Henicorhynchus, with a Revised Diagnosis of Gymnostomus (Cyprinidae: Labeoninae)</mods:title>
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<treatment id="03B587CBFFDB0C581E7785B688EC23E5" ID-DOI="http://doi.org/10.5281/zenodo.13291184" ID-Zenodo-Dep="13291184" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5" httpUri="http://treatment.plazi.org/id/03B587CBFFDB0C581E7785B688EC23E5" lastPageId="9" pageId="5">
<subSubSection id="C3066556FFDB0C541E7785B689392585" pageId="5" type="nomenclature">
<paragraph id="8BA336DDFFDB0C541E7785B6885A25A2" blockId="5.[859,1407,1709,1732]" box="[859,1407,1709,1732]" pageId="5">
<heading id="D0EB81B1FFDB0C541E7785B6885A25A2" bold="true" box="[859,1407,1709,1732]" fontSize="10" level="2" pageId="5" reason="2">
<taxonomicName id="4C1C4D5EFFDB0C541E7785B6885A25A2" ID-CoL="6M28Z" authority="(Fowler, 1934)" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[859,1407,1709,1732]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFDB0C541E7785B6885A25A2" bold="true" box="[859,1407,1709,1732]" pageId="5">
<emphasis id="B968EACFFFDB0C541E7785B689E025A2" bold="true" box="[859,1221,1709,1732]" italics="true" pageId="5">Henicorhynchus caudimaculatus</emphasis>
(
<bibRefCitation id="EF8D4B2CFFDB0C5419F885B6885225A2" author="Fowler, H. W." box="[1236,1399,1709,1732]" pageId="5" pagination="67 - 163" refId="ref17302" refString="Fowler, H. W. 1934. Zoological results of the Third de Schauensee Siamese Expedition, Part I. - Fishes. Proceedings of the Academy of Natural Sciences of Philadelphia 86: 67 - 163." type="journal article" year="1934">Fowler, 1934</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA336DDFFDB0C541E7785D789392585" blockId="5.[859,1052,1740,1793]" box="[859,1052,1740,1763]" pageId="5">Striped Mud Carp</paragraph>
</subSubSection>
<subSubSection id="C3066556FFDB0C541E7785F18E962467" box="[859,947,1770,1793]" pageId="5" type="description">
<paragraph id="8BA336DDFFDB0C541E7785F18E962467" blockId="5.[859,1052,1740,1793]" box="[859,947,1770,1793]" pageId="5">
<figureCitation id="13272A58FFDB0C541E7785F18E962467" box="[859,947,1770,1793]" captionStart="FIG" captionStartId="4.[103,131,1976,1996]" captionTargetBox="[103,1489,1118,1946]" captionTargetId="figure-371@4.[103,1489,1118,1946]" captionTargetPageId="4" captionText="FIG. 4. Henicorhynchus caudimaculatus. (A) ANSP 58332, holotype, 41.4 mm SL, (B) USNM 107960, holotype of Cirrhinus lineatus, 108.8 mm SL. Images not to scale. Photo A by K. Luckenbill (ANSP); photo B by S. Raredon (USNM)." figureDoi="http://doi.org/10.5281/zenodo.13628573" httpUri="https://zenodo.org/record/13628573/files/figure.png" pageId="5">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3066556FFDB0C541E77840288382481" pageId="5" type="reference_group">
<paragraph id="8BA336DDFFDB0C541E7784028E9A240A" blockId="5.[859,1530,1817,2024]" pageId="5">
<treatmentCitationGroup id="AB0C11F3FFDB0C541E7784028E9A240A" pageId="5">
<treatmentCitation id="0ABD10CCFFDB0C541E77840288452449" author="Fowler, H. W." box="[859,1376,1817,1840]" page="133" pageId="5" year="1934">
<taxonomicName id="4C1C4D5EFFDB0C541E77840288452449" ID-CoL="59TDB" authority="Fowler, 1934: 133" authorityName="Fowler" authorityPageNumber="133" authorityYear="1934" box="[859,1376,1817,1840]" family="Cyprinidae" genus="Tylognathus" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFDB0C541E77840289AE2449" box="[859,1163,1817,1839]" italics="true" pageId="5">Tylognathus caudimaculatus</emphasis>
<materialsCitation id="3B743C80FFDB0C5419B6840188452449" box="[1178,1376,1818,1840]" collectionCode="ANSP" country="Thailand" location="Chiang Mai" pageId="5" specimenCode="ANSP 58332" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
<bibRefCitation id="EF8D4B2CFFDB0C5419B6840188452449" author="Fowler, H. W." box="[1178,1376,1818,1840]" pageId="5" pagination="67 - 163" refId="ref17302" refString="Fowler, H. W. 1934. Zoological results of the Third de Schauensee Siamese Expedition, Part I. - Fishes. Proceedings of the Academy of Natural Sciences of Philadelphia 86: 67 - 163." type="journal article" year="1934">Fowler, 1934:133</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, figs. 8990. Type locality:
<collectingCountry id="F30B764DFFDB0C54190D842389A22428" box="[1057,1159,1848,1870]" name="Thailand" pageId="5">Thailand</collectingCountry>
,
<collectingRegion id="49D8F83FFFDB0C5419B78423880E2428" box="[1179,1323,1848,1870]" country="Thailand" name="Chiang Mai" pageId="5">Chiang Mai</collectingRegion>
.
<typeStatus id="54A7887FFFDB0C5418128423888D2428" box="[1342,1448,1848,1870]" pageId="5" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBBA9EA6FFDB0C54189084238E9E240A" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="5">ANSP 58332</specimenCode>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA336DDFFDB0C541E77846E89B724AF" blockId="5.[859,1530,1817,2024]" pageId="5">
<treatmentCitationGroup id="AB0C11F3FFDB0C541E77846E89B724AF" pageId="5">
<treatmentCitation id="0ABD10CCFFDB0C541E77846E89CC24EA" author="Smith, H. M." box="[859,1257,1909,1932]" page="163" pageId="5" year="1945">
<taxonomicName id="4C1C4D5EFFDB0C541E77846E89CC24EA" ID-CoL="VGT8" authority="Smith, 1945: 163" authorityName="Smith" authorityPageNumber="163" authorityYear="1945" box="[859,1257,1909,1932]" family="Cyprinidae" genus="Cirrhinus" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B968EACFFFDB0C541E77846E893924ED" box="[859,1052,1909,1931]" italics="true" pageId="5">Cirrhinus lineatus</emphasis>
<materialsCitation id="3B743C80FFDB0C541906846D89CC24EA" box="[1066,1257,1910,1932]" collectionCode="USNM" country="Thailand" location="Lam Ton Lang" pageId="5" specimenCode="USNM 107960" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EF8D4B2CFFDB0C541906846D89CC24EA" author="Smith, H. M." box="[1066,1257,1910,1932]" pageId="5" pagination="1 - 662" refId="ref18173" refString="Smith, H. M. 1945. The fresh-water fishes of Siam, or Thailand. Bulletin of the United States National Museum 188: 1 - 662." type="journal article" year="1945">Smith, 1945:163</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, fig. 25. Type locality:
<collectingCountry id="F30B764DFFDB0C541E5F848F8EFC24CC" box="[883,985,1940,1962]" name="Thailand" pageId="5">Thailand</collectingCountry>
,
<location id="8EC36006FFDB0C541EC1848E898424CC" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFDB0C541EC1848E898424CC" box="[1005,1185,1941,1962]" country="Thailand" name="Lam Ton Lang" pageId="5">Lam Ton Lang</location>
, a tributary of Menam Sak.
<typeStatus id="54A7887FFFDB0C541E5F84A88EF824AF" box="[883,989,1971,1993]" pageId="5" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBBA9EA6FFDB0C541EC484A889A824AF" box="[1000,1165,1971,1993]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" type="Museum">USNM 107960</specimenCode>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA336DDFFDB0C541E7784CA88382481" blockId="5.[859,1530,1817,2024]" box="[859,1309,2001,2024]" pageId="5">
<emphasis id="B968EACFFFDB0C541E7784CA895A2481" box="[859,1151,2001,2023]" italics="true" pageId="5">
<taxonomicName id="4C1C4D5EFFDB0C541E7784CA89412481" ID-CoL="VGSP" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[859,1124,2001,2023]" family="Cyprinidae" genus="Cirrhinus" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="caudimaculatus">Cirrhinus caudimaculatus</taxonomicName>
</emphasis>
<bibRefCitation id="EF8D4B2CFFDB0C54195384C9883C2481" author="Roberts, T. R." box="[1151,1305,2002,2024]" pageId="5" pagination="171 - 203" refId="ref18024" refString="Roberts, T. R. 1997. Systematic revision of the tropical Asian labeonin cyprinid fish genus Cirrhinus, with descriptions of new species and biological observations on C. lobatus. Natural History Bulletin of the Siam Society 45: 171 - 203." type="journal article" year="1997">Roberts, 1997</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF636655FFD80C571D4B875E8E372712" ID-DOI="http://doi.org/10.5281/zenodo.13628575" ID-Zenodo-Dep="13628575" httpUri="https://zenodo.org/record/13628575/files/figure.png" pageId="6" startId="6.[103,131,1093,1113]" targetBox="[103,1490,178,1063]" targetPageId="6" targetType="figure">
<paragraph id="8BA336DDFFD80C571D4B875E8E372712" blockId="6.[103,1490,1092,1140]" pageId="6">
FIG. 5.
<taxonomicName id="4C1C4D5EFFD80C571D9D875F8C8F273E" box="[177,426,1092,1112]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="entmema">
<emphasis id="B968EACFFFD80C571D9D875F8C8F273E" bold="true" box="[177,426,1092,1112]" italics="true" pageId="6">Henicorhynchus entmema</emphasis>
</taxonomicName>
. (A) ANSP 59092, holotype, 46.4 mm SL, (B) USNM 119490, holotype of
<taxonomicName id="4C1C4D5EFFD80C57194A875F89E3273E" authorityName="Smith" authorityYear="1945" box="[1126,1222,1092,1112]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="lobatus">
<emphasis id="B968EACFFFD80C57194A875F89E3273E" bold="true" box="[1126,1222,1092,1112]" italics="true" pageId="6">H. lobatus</emphasis>
</taxonomicName>
, 96.7 mm SL. Images not to scale. Photo A by K. Luckenbill (ANSP); photo B by S. Raredon (USNM).
</paragraph>
</caption>
<subSubSection id="C3066556FFD80C571D4B87848F2A2794" pageId="6" type="description">
<paragraph id="8BA336DDFFD80C571D4B87848CFD27D3" blockId="6.[103,608,1183,1267]" box="[103,472,1183,1206]" pageId="6">
<emphasis id="B968EACFFFD80C571D4B87848C1C27D3" box="[103,313,1183,1205]" italics="true" pageId="6">
<taxonomicName id="4C1C4D5EFFD80C571D4B87848C3B27D3" authorityName="Smith" authorityYear="1945" box="[103,286,1183,1205]" family="Cyprinidae" genus="Cirrhinus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="lineatus">Cirrhinus lineatus</taxonomicName>
</emphasis>
<bibRefCitation id="EF8D4B2CFFD80C571C1587BB8CF127D3" author="Roberts, T. R." box="[313,468,1184,1206]" pageId="6" pagination="171 - 203" refId="ref18024" refString="Roberts, T. R. 1997. Systematic revision of the tropical Asian labeonin cyprinid fish genus Cirrhinus, with descriptions of new species and biological observations on C. lobatus. Natural History Bulletin of the Siam Society 45: 171 - 203." type="journal article" year="1997">Roberts, 1997</bibRefCitation>
.
</paragraph>
<paragraph id="8BA336DDFFD80C571D4B87A48F4527B5" blockId="6.[103,608,1183,1267]" box="[103,608,1214,1236]" pageId="6">
<taxonomicName id="4C1C4D5EFFD80C571D4B87A48C8027B2" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[103,421,1214,1236]" family="Cyprinidae" genus="Gymnostomus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFD80C571D4B87A48C8027B2" box="[103,421,1214,1236]" italics="true" pageId="6">Gymnostomus caudimaculatus</emphasis>
</taxonomicName>
<bibRefCitation id="EF8D4B2CFFD80C571CEC87A58F7927B5" author="Kottelat, M." box="[448,604,1214,1236]" pageId="6" pagination="1 - 663" refId="ref17745" refString="Kottelat, M. 2013. The fishes of the inland waters of Southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves, and estuaries. Raffles Bulletin of Zoology Supplement 27: 1 - 663." type="journal article" year="2013">Kottelat, 2013</bibRefCitation>
.
</paragraph>
<paragraph id="8BA336DDFFD80C571D4B87C68F2A2794" blockId="6.[103,608,1183,1267]" box="[103,527,1244,1267]" pageId="6">
<taxonomicName id="4C1C4D5EFFD80C571D4B87C68C712794" baseAuthorityName="Smith" baseAuthorityYear="1945" box="[103,340,1244,1266]" family="Cyprinidae" genus="Gymnostomus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B968EACFFFD80C571D4B87C68C712794" box="[103,340,1244,1266]" italics="true" pageId="6">Gymnostomus lineatus</emphasis>
</taxonomicName>
<bibRefCitation id="EF8D4B2CFFD80C571C4387C68F2E2794" author="Kottelat, M." box="[367,523,1245,1267]" pageId="6" pagination="1 - 663" refId="ref17745" refString="Kottelat, M. 2013. The fishes of the inland waters of Southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves, and estuaries. Raffles Bulletin of Zoology Supplement 27: 1 - 663." type="journal article" year="2013">Kottelat, 2013</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066556FFD80C581D4B86088F7E23E2" lastPageId="9" pageId="6" type="diagnosis">
<paragraph id="8BA336DDFFD80C571D4B86088E212621" blockId="6.[103,773,1299,1351]" pageId="6">
<emphasis id="B968EACFFFD80C571D4B86088DC9264C" bold="true" box="[103,236,1299,1322]" italics="true" pageId="6">Diagnosis.—</emphasis>
A member of
<taxonomicName id="4C1C4D5EFFD80C571CAA86088F0F264F" authorityName="Smith" authorityYear="1945" box="[390,554,1299,1321]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="genus">
<emphasis id="B968EACFFFD80C571CAA86088F0F264F" box="[390,554,1299,1321]" italics="true" pageId="6">Henicorhynchus</emphasis>
</taxonomicName>
distinguished from other members of the genus (
<tableCitation id="C69E0366FFD80C571CFD862A8F022620" box="[465,551,1329,1351]" captionStart="Table 1" captionStartId="4.[103,151,178,198]" captionText="Table 1. Characteristics that distinguish species of Henicorhynchus." pageId="6">Table 1</tableCitation>
) by the following
</paragraph>
<paragraph id="8BA336DDFFD80C571E1887BB88F72621" blockId="6.[820,1490,1184,1351]" pageId="6">combination of characters: mouth subterminal, weakly oblique to nearly horizontal (less than or equal to 208 to midline of body); maxillary barbels present; edge of rostral cap straight, without medial indent; longitudinal stripes on flanks in most specimens, particularly larger individuals;</paragraph>
<paragraph id="8BA336DDFFD80C571D4B84C98DB82480" blockId="6.[103,157,2002,2022]" box="[103,157,2002,2022]" pageId="6">
<emphasis id="B968EACFFFD80C571D4B84C98DB82480" bold="true" box="[103,157,2002,2022]" pageId="6">FIG. 6.</emphasis>
</paragraph>
<paragraph id="8BA336DDFFD80C571D9E84C989272480" blockId="6.[178,1026,2002,2022]" box="[178,1026,2002,2022]" pageId="6">
<emphasis id="B968EACFFFD80C571D9E84C989272480" bold="true" box="[178,1026,2002,2022]" pageId="6">
<taxonomicName id="4C1C4D5EFFD80C571D9E84C98CD62480" authority=", CAS" authorityName="CAS" box="[178,499,2002,2022]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="ornatipinnis">
<emphasis id="B968EACFFFD80C571D9E84C98CE72480" bold="true" box="[178,450,2002,2022]" italics="true" pageId="6">Henicorhynchus ornatipinnis</emphasis>
, CAS
</taxonomicName>
91756,
<materialsCitation id="3B743C80FFD80C571F6F84C98EDB2480" box="[579,1022,2002,2022]" collectionCode="CAS" collectorName="Photo &amp; J. Fong" pageId="6" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A7887FFFD80C571F6F84C98FBD2480" box="[579,664,2002,2022]" pageId="6" type="holotype">holotype</typeStatus>
,
<quantity id="4CE49B38FFD80C571F8E84C98FDC2480" box="[674,761,2002,2022]" metricMagnitude="-2" metricUnit="m" metricValue="6.94" pageId="6" unit="mm" value="69.4">69.4 mm</quantity>
SL.
<collectorName id="26E9530BFFD80C571E0E84C98E7F2480" box="[802,858,2002,2022]" pageId="6">Photo</collectorName>
by
<collectorName id="26E9530BFFD80C571E5284C98E9A2480" box="[894,959,2002,2022]" pageId="6">J. Fong</collectorName>
(
<collectionCode id="ED0DAE18FFD80C571EE084C98EDD2480" box="[972,1016,2002,2022]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</emphasis>
</paragraph>
<paragraph id="8BA336DDFFD80C571D378B178F782B78" blockId="6.[27,605,2060,2096]" box="[27,605,2060,2078]" pageId="6">
Downloaded From: https://bioone.org/journals/Copeia on
<date id="FFA2101DFFD80C571CD88B178F782B78" box="[500,605,2060,2078]" pageId="6" value="2024-08-05">05 Aug 2024</date>
</paragraph>
<paragraph id="8BA336DDFFD80C571D378B3B8CB42B56" blockId="6.[27,605,2060,2096]" box="[27,401,2079,2096]" pageId="6">Terms of Use: https://bioone.org/terms-of-use</paragraph>
<caption id="DF636655FFD90C561DA2819C88DC2600" ID-DOI="http://doi.org/10.5281/zenodo.13628579" ID-Zenodo-Dep="13628579" httpUri="https://zenodo.org/record/13628579/files/figure.png" pageId="7" startId="7.[142,170,647,667]" targetBox="[142,1530,178,618]" targetPageId="7" targetType="figure">
<paragraph id="8BA336DDFFD90C561DA2819C88DC2600" blockId="7.[142,1530,647,1382]" pageId="7">
<emphasis id="B968EACFFFD90C561DA2819C89D921DB" bold="true" pageId="7">
FIG. 7.
<taxonomicName id="4C1C4D5EFFD90C561DF5819C8F3021FD" authority=", ANSP" authorityName="ANSP" box="[217,533,647,667]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="B968EACFFFD90C561DF5819C8CF021FD" bold="true" box="[217,469,647,667]" italics="true" pageId="7">Henicorhynchus siamensis</emphasis>
, ANSP
</taxonomicName>
68069, holotype of
<taxonomicName id="4C1C4D5EFFD90C561FF5819C8E9D21FD" authorityName="Fowler" authorityYear="1937" box="[729,952,647,667]" family="Cyprinidae" genus="Cirrhinus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="marginipinnis">
<emphasis id="B968EACFFFD90C561FF5819C8E9D21FD" bold="true" box="[729,952,647,667]" italics="true" pageId="7">Cirrhinus marginipinnis</emphasis>
</taxonomicName>
, 113.7 mm SL. Photo by K. Luckenbill (ANSP).
<emphasis id="B968EACFFFD90C5619DC81B289D921DB" bold="true" box="[1264,1276,681,701]" italics="true" pageId="7">=</emphasis>
</emphasis>
caudal peduncle with small brown spot at medial insertion of opercula occasionally silver. Dorsal
<superScript id="7C699B95FFD90C5619C181D289D121B5" attach="left" box="[1261,1268,713,723]" fontSize="4" pageId="7">1</superScript>
to of flank light to
<subScript id="17983498FFD90C5619DB81CA89DB21BD" attach="right" box="[1271,1278,721,731]" fontSize="4" pageId="7">3</subScript>
caudal fin in smaller specimens; caudal fin mostly clear, with dark brown, with scattered darker
<emphasis id="B968EACFFFD90C5619FC81FF89F9219E" bold="true" box="[1232,1244,740,760]" italics="true" pageId="7">=</emphasis>
brown spots on scales in scattered melanophores; pectoral, pelvic, and anal fins some specimens; ventral to
<superScript id="7C699B95FFD90C5619E0801E89F62069" attach="left" box="[1228,1235,773,783]" fontSize="4" pageId="7">2</superScript>
cream, yellow, or light
<subScript id="17983498FFD90C5619FB801689FB2071" attach="right" box="[1239,1246,781,791]" fontSize="4" pageId="7">3</subScript>
hyaline in life. brown. Brown to black spot on medial portion of caudal peduncle near caudal-fin insertion in smaller specimens
<emphasis id="B968EACFFFD90C561DA2804C8FA7200B" box="[142,642,855,878]" italics="true" pageId="7">
<emphasis id="B968EACFFFD90C561DA2804C8C102008" bold="true" box="[142,309,855,878]" italics="true" pageId="7">Comparisons.—</emphasis>
<taxonomicName id="4C1C4D5EFFD90C561C19804C8FA7200B" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[309,642,855,877]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="caudimaculatus">Henicorhynchus caudimaculatus</taxonomicName>
</emphasis>
is most similar (,100 mm SL). Up to seven brown longitudinal stripes on to
<taxonomicName id="4C1C4D5EFFD90C561D80806C8C0F20EA" box="[172,298,887,908]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="entmema">
<emphasis id="B968EACFFFD90C561D80806C8C0F20EA" box="[172,298,887,908]" italics="true" pageId="7">H. entmema</emphasis>
</taxonomicName>
in having a subterminal mouth and maxillary flanks in most specimens; usually distinct in specimens barbels, but differs in having a straight edge on the rostral cap greater than approximately 40 mm SL. Venter cream, yellow, (vs. a medial indent on the rostral cap of
<taxonomicName id="4C1C4D5EFFD90C561F7A80A98FF020A1" box="[598,725,946,967]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="entmema">
<emphasis id="B968EACFFFD90C561F7A80A98FF020A1" box="[598,725,946,967]" italics="true" pageId="7">H. entmema</emphasis>
</taxonomicName>
) and in or light brown. Dorsal fin with dark brown clusters of usually having a series of longitudinal stripes along the flank melanophores forming blotches in middle portions of (vs. flank always immaculate in
<taxonomicName id="4C1C4D5EFFD90C561CD980F58F502765" box="[501,629,1006,1027]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="entmema">
<emphasis id="B968EACFFFD90C561CD980F58F502765" box="[501,629,1006,1027]" italics="true" pageId="7">H. entmema</emphasis>
</taxonomicName>
). interradial membranes, distal edge brown; pectoral, pelvic, and anal fins hyaline; caudal fin generally clear, with
<emphasis id="B968EACFFFD90C561DA2873F8C06275D" bold="true" box="[142,291,1060,1083]" italics="true" pageId="7">Description.—</emphasis>
Morphometrics and meristics in Table 2. Mouth scattered melanophores. subterminal, weakly oblique to nearly horizontal. Maxillary barbels present. Edge of rostral cap straight, without medial
<emphasis id="B968EACFFFD90C561E7787798EF7271F" bold="true" box="[859,978,1122,1145]" italics="true" pageId="7">Remarks.—</emphasis>
In the original description of
<taxonomicName id="4C1C4D5EFFD90C56181C877888D7271E" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[1328,1522,1122,1144]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFD90C56181C877888D7271E" box="[1328,1522,1122,1144]" italics="true" pageId="7">H. caudimaculatus</emphasis>
</taxonomicName>
, indent; rostral cap occasionally with well-developed tuber-
<bibRefCitation id="EF8D4B2CFFD90C561E77879B8ECA27F3" author="Fowler, H. W." box="[859,1007,1152,1174]" pageId="7" pagination="67 - 163" refId="ref17302" refString="Fowler, H. W. 1934. Zoological results of the Third de Schauensee Siamese Expedition, Part I. - Fishes. Proceedings of the Academy of Natural Sciences of Philadelphia 86: 67 - 163." type="journal article" year="1934">Fowler (1934)</bibRefCitation>
noted a grayish axial streak along the flank of cles. Pectoral fin with 1416 branched rays. Lateral-line scales the holotype (p. 126, fig. 90) in addition to a brown spot on and pored scales on caudal fin 3134
<emphasis id="B968EACFFFD90C561F3287A28F0B27B6" bold="true" box="[542,558,1209,1232]" pageId="7">þ</emphasis>
23; predorsal scales 9 the caudal peduncle near the insertion of the caudal-fin rays. 11; scale rows above lateral line 5; scale rows below lateral In similarly sized specimens previously identified as
<taxonomicName id="4C1C4D5EFFD90C5618AA87C188D72789" baseAuthorityName="Smith" baseAuthorityYear="1945" box="[1414,1522,1241,1263]" family="Cyprinidae" genus="Gymnostomus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B968EACFFFD90C5618AA87C188D72789" box="[1414,1522,1241,1263]" italics="true" pageId="7">H. lineatus</emphasis>
</taxonomicName>
, line 4 5; scale rows between lateral line and pelvic-fin a similar spot on the caudal peduncle is present and the lateral origin 4. Maximum length
<emphasis id="B968EACFFFD90C561CE286098CC5264F" bold="true" box="[462,480,1298,1321]" pageId="7">¼</emphasis>
150 mm SL (
<bibRefCitation id="EF8D4B2CFFD90C561F59860F8E33264F" author="Kottelat, M." box="[629,790,1300,1322]" pageId="7" refId="ref17691" refString="Kottelat, M. 2001. Fishes of Laos. Wildlife Heritage Trust, Colombo, Sri Lanka." type="book" year="2001">Kottelat, 2001</bibRefCitation>
). stripes vary from very faint to bold (also noted by Roberts Dorsum of head and body brown. Dorsal of side of head [1997]), with some small specimens also possessing one light to dark brown; ventral
<emphasis id="B968EACFFFD90C561CFD86498CC62600" bold="true" box="[465,483,1362,1382]" pageId="7">ł</emphasis>
cream to yellow; cheeks and distinct stripe. The types of
<taxonomicName id="4C1C4D5EFFD90C56198C865488E62603" authorityName="Fowler" authorityYear="1934" box="[1184,1475,1359,1381]" family="Cyprinidae" genus="Tylognathus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFD90C56198C865488E62603" box="[1184,1475,1359,1381]" italics="true" pageId="7">Tylognathus caudimaculatus</emphasis>
</taxonomicName>
and
</paragraph>
</caption>
<caption id="DF636655FFD90C561DA284C9893B2480" ID-DOI="http://doi.org/10.5281/zenodo.13628581" ID-Zenodo-Dep="13628581" box="[142,1054,2002,2022]" httpUri="https://zenodo.org/record/13628581/files/figure.png" pageId="7" startId="7.[142,170,2002,2022]" targetBox="[142,1530,1424,1973]" targetPageId="7">
<paragraph id="8BA336DDFFD90C561DA284C9893B2480" blockId="7.[142,1054,2002,2022]" box="[142,1054,2002,2022]" pageId="7">
<emphasis id="B968EACFFFD90C561DA284C98DE12480" bold="true" box="[142,196,2002,2022]" pageId="7">FIG. 8.</emphasis>
<emphasis id="B968EACFFFD90C561DF584C9893B2480" bold="true" box="[217,1054,2002,2022]" pageId="7">
<taxonomicName id="4C1C4D5EFFD90C561DF584C98CDF2480" authority=", UF" authorityName="UF" box="[217,506,2002,2022]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="B968EACFFFD90C561DF584C98CF02480" bold="true" box="[217,469,2002,2022]" italics="true" pageId="7">Henicorhynchus siamensis</emphasis>
, UF
</taxonomicName>
191421, live,
<quantity id="4CE49B38FFD90C561FAF84C98FFC2480" box="[643,729,2002,2022]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" pageId="7" unit="mm" value="51.5">51.5 mm</quantity>
SL. Photo by Z. Randall (FLMNH).
</emphasis>
</paragraph>
</caption>
<caption id="DF636655FFD60C591D4B85CD8CB82462" ID-DOI="http://doi.org/10.5281/zenodo.13628583" ID-Zenodo-Dep="13628583" httpUri="https://zenodo.org/record/13628583/files/figure.png" pageId="8" startId="8.[103,131,1750,1770]" targetBox="[104,1490,178,1718]" targetPageId="8" targetType="figure">
<paragraph id="8BA336DDFFD60C591D4B85CD8CB82462" blockId="8.[103,1490,1749,1796]" pageId="8">
FIG. 9. Distributions of georeferenced specimens examined of: (A)
<taxonomicName id="4C1C4D5EFFD60C591FF185CE8ED9258F" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[733,1020,1749,1769]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="caudiguttatus">
<emphasis id="B968EACFFFD60C591FF185CE8ED9258F" bold="true" box="[733,1020,1749,1769]" italics="true" pageId="8">Henicorhynchus caudiguttatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D5EFFD60C59190685CE8996258F" box="[1066,1203,1749,1769]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="ornatipinnis">
<emphasis id="B968EACFFFD60C59190685CE8996258F" bold="true" box="[1066,1203,1749,1769]" italics="true" pageId="8">H. ornatipinnis</emphasis>
</taxonomicName>
, (B)
<taxonomicName id="4C1C4D5EFFD60C5919CD85CE88B4258F" authorityName="Fowler" authorityYear="1934" box="[1249,1425,1749,1769]" family="Cyprinidae" genus="Tylognathus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFD60C5919CD85CE88B4258F" bold="true" box="[1249,1425,1749,1769]" italics="true" pageId="8">H. caudimaculatus</emphasis>
</taxonomicName>
, (C)
<taxonomicName id="4C1C4D5EFFD60C59189285CE8DE72462" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="entmema">
<emphasis id="B968EACFFFD60C59189285CE8DE72462" bold="true" italics="true" pageId="8">H. entmema</emphasis>
</taxonomicName>
, and (D)
<taxonomicName id="4C1C4D5EFFD60C591C0C85EB8CBD2462" box="[288,408,1776,1796]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="B968EACFFFD60C591C0C85EB8CBD2462" bold="true" box="[288,408,1776,1796]" italics="true" pageId="8">H. siamensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA336DDFFD60C591D4B842B8E202481" blockId="8.[103,773,1840,2024]" pageId="8">
<taxonomicName id="4C1C4D5EFFD60C591D4B842B8C9F2420" authority="Smith, 1945" authorityName="Smith" authorityYear="1945" box="[103,442,1840,1862]" family="Cyprinidae" genus="Cirrhinus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B968EACFFFD60C591D4B842B8C042420" box="[103,289,1840,1862]" italics="true" pageId="8">Cirrhinus lineatus</emphasis>
<bibRefCitation id="EF8D4B2CFFD60C591C03842B8C9F2420" author="Smith, H. M." box="[303,442,1840,1862]" pageId="8" pagination="1 - 662" refId="ref18173" refString="Smith, H. M. 1945. The fresh-water fishes of Siam, or Thailand. Bulletin of the United States National Museum 188: 1 - 662." type="journal article" year="1945">Smith, 1945</bibRefCitation>
</taxonomicName>
share similar oromandibular morphology, specifically a subterminal, weakly oblique mouth, maxillary barbels, and a smooth rostral cap without a medial indent. Based on similarities in oromandibular structures and color patterns when comparing specimens of variable sizes, we recognize
<taxonomicName id="4C1C4D5EFFD60C591DCD84CA8C812481" authorityName="Smith" authorityYear="1945" box="[225,420,2001,2023]" family="Cyprinidae" genus="Cirrhinus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B968EACFFFD60C591DCD84CA8C812481" box="[225,420,2001,2023]" italics="true" pageId="8">Cirrhinus lineatus</emphasis>
</taxonomicName>
as a junior synonym of
<emphasis id="B968EACFFFD60C591FC084C98E202481" box="[748,773,2002,2023]" italics="true" pageId="8">H.</emphasis>
</paragraph>
<paragraph id="8BA336DDFFD60C591D378B178F782B78" blockId="8.[27,605,2060,2096]" box="[27,605,2060,2078]" pageId="8">
Downloaded From: https://bioone.org/journals/Copeia on
<date id="FFA2101DFFD60C591CD88B178F782B78" box="[500,605,2060,2078]" pageId="8" value="2024-08-05">05 Aug 2024</date>
</paragraph>
<paragraph id="8BA336DDFFD60C591D378B3B8CB42B56" blockId="8.[27,605,2060,2096]" box="[27,401,2079,2096]" pageId="8">Terms of Use: https://bioone.org/terms-of-use</paragraph>
<paragraph id="8BA336DDFFD60C591E18842B8981248E" blockId="8.[820,1490,1840,2024]" pageId="8">
<taxonomicName id="4C1C4D5EFFD60C591E18842B8EF42420" authorityName="Fowler" authorityYear="1934" box="[820,977,1840,1862]" family="Cyprinidae" genus="Tylognathus" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFD60C591E18842B8EF42420" box="[820,977,1840,1862]" italics="true" pageId="8">caudimaculatus</emphasis>
</taxonomicName>
. There is ontogenetic variation in color pattern, with small individuals possessing a brown to black spot at the base of the caudal peduncle, with longitudinal stripes developing with age and the spot eventually disappearing. A similar pattern is observed in
<taxonomicName id="4C1C4D5EFFD60C5919A084AA89BA248E" authority="(Fowler, 1934) (Ciccotto and Page, 2016 a)" authorityName="Ciccotto and Page" authorityYear="2016" baseAuthorityName="Fowler" baseAuthorityYear="1934" family="Cyprinidae" genus="Lobocheilos" kingdom="Animalia" order="Cypriniformes" pageId="8" phylum="Chordata" rank="species" species="rhabdoura">
<emphasis id="B968EACFFFD60C5919A084AA885724A1" box="[1164,1394,1969,1991]" italics="true" pageId="8">Lobocheilos rhabdoura</emphasis>
(
<bibRefCitation id="EF8D4B2CFFD60C5918A884AA8E4F2481" author="Fowler, H. W." pageId="8" pagination="67 - 163" refId="ref17302" refString="Fowler, H. W. 1934. Zoological results of the Third de Schauensee Siamese Expedition, Part I. - Fishes. Proceedings of the Academy of Natural Sciences of Philadelphia 86: 67 - 163." type="journal article" year="1934">Fowler, 1934</bibRefCitation>
) (
<bibRefCitation id="EF8D4B2CFFD60C591EAD84C989BD248E" author="Ciccotto, P. J. &amp; L. M. Page" box="[897,1176,2002,2024]" pageId="8" pagination="879 - 889" refId="ref17154" refString="Ciccotto, P. J., and L. M. Page. 2016 a. From 12 to one species: variation in Lobocheilos rhabdoura (Fowler, 1934) (Cyprinidae: Labeonini). Copeia 104: 879 - 889." type="journal article" year="2016">Ciccotto and Page, 2016a</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA336DDFFD70C581DA2836B8F7E23E2" blockId="9.[142,603,112,132]" box="[142,603,112,132]" pageId="9">
Ciccotto and Page—Revision of
<taxonomicName id="4C1C4D5EFFD70C581CE6836B8F7E23E2" authorityName="Smith" authorityYear="1945" box="[458,603,112,132]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="9" phylum="Chordata" rank="genus">
<emphasis id="B968EACFFFD70C581CE6836B8F7E23E2" box="[458,603,112,132]" italics="true" pageId="9">Henicorhynchus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3066556FFD70C581DA283A98F5E2265" pageId="9" type="distribution">
<paragraph id="8BA336DDFFD70C581DA283A98F5E2265" blockId="9.[142,812,178,260]" pageId="9">
<emphasis id="B968EACFFFD70C581DA283A98F5223AE" box="[142,631,178,201]" italics="true" pageId="9">
<emphasis id="B968EACFFFD70C581DA283A98C0C23AF" bold="true" box="[142,297,178,201]" italics="true" pageId="9">Distribution.—</emphasis>
<taxonomicName id="4C1C4D5EFFD70C581C0583A98F5223AE" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[297,631,178,200]" family="Cyprinidae" genus="Henicorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="9" phylum="Chordata" rank="species" species="caudimaculatus">Henicorhynchus caudimaculatus</taxonomicName>
</emphasis>
is distributed in the Chao Phraya basin in
<collectingCountry id="F30B764DFFD70C581C8183CB8F2A2380" box="[429,527,208,230]" name="Thailand" pageId="9">Thailand</collectingCountry>
and the Mekong basin in
<collectingCountry id="F30B764DFFD70C581DA283F58C272262" box="[142,258,238,260]" name="Cambodia" pageId="9">Cambodia</collectingCountry>
, Lao PDR, and
<collectingCountry id="F30B764DFFD70C581C8183F58F2A2262" box="[429,527,238,260]" name="Thailand" pageId="9">Thailand</collectingCountry>
(
<figureCitation id="13272A58FFD70C581F3383F58F4B2265" box="[543,622,238,260]" captionStart="FIG" captionStartId="8.[103,131,1750,1770]" captionTargetBox="[104,1490,178,1718]" captionTargetId="figure-101@8.[103,1491,178,1720]" captionTargetPageId="8" captionText="FIG. 9. Distributions of georeferenced specimens examined of: (A) Henicorhynchus caudiguttatus and H. ornatipinnis, (B) H. caudimaculatus, (C) H. entmema, and (D) H. siamensis." figureDoi="http://doi.org/10.5281/zenodo.13628583" httpUri="https://zenodo.org/record/13628583/files/figure.png" pageId="9">Fig. 9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066556FFD70C581DA2823888EC23E5" pageId="9" type="materials_examined">
<paragraph id="8BA336DDFFD70C581DA282388F41248E" blockId="9.[142,812,291,2024]" pageId="9">
<emphasis id="B968EACFFFD70C581DA282388C53225C" bold="true" box="[142,374,291,314]" italics="true" pageId="9">Material examined.—</emphasis>
(Identified, but not measured if no SL given).
<collectingCountry id="F30B764DFFD70C581DC8825A8C7C2231" box="[228,345,321,343]" name="Cambodia" pageId="9">Cambodia</collectingCountry>
: Mekong basin:
<materialsCitation id="3B743C80FFD70C581F3A825A8DE222C9" collectingDate="1994-02-14" collectionCode="CAS" collectorName="O Champha &amp; Te Veng" latitude="14.10068" location="Tonle San" longLatPrecision="1" longitude="107.13528" pageId="9" specimenCode="CAS 81552, 5" specimenCount="1" stateProvince="Ratanakiri">
<specimenCode id="DBBA9EA6FFD70C581F3A825A8FB02231" box="[534,661,321,343]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 81552</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F88825A8F962231" box="[676,691,321,343]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">5</specimenCode>
,
<collectorName id="26E9530BFFD70C581FEE825A8D9F2212" pageId="9">O Champha</collectorName>
,
<quantity id="4CE49B38FFD70C581DE982448C312212" box="[197,276,350,372]" metricMagnitude="3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="9" unit="km" value="6.0" valueMax="8.0" valueMin="4.0">48 km</quantity>
upstream from its mouth into
<location id="8EC36006FFD70C581F4982458FF52212" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F4982458FF52212" box="[613,720,350,372]" latitude="14.10068" longLatPrecision="1" longitude="107.13528" name="Tonle San" pageId="9" stateProvince="Ratanakiri">Tonle San</location>
(near
<collectorName id="26E9530BFFD70C581E3F82448DEC22F7" pageId="9">Te Veng</collectorName>
,
<collectingRegion id="49D8F83FFFD70C581DF982678CA222F7" box="[213,391,380,402]" country="Cambodia" name="Ratanakiri" pageId="9">Ratanakiri prov.</collectingRegion>
),
<geoCoordinate id="EE28501AFFD70C581CB782678F3422F7" box="[411,529,380,402]" degrees="14.10068" direction="north" orientation="latitude" pageId="9" precision="1" value="14.10068">14.10068N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581F3282678FB922F7" box="[542,668,380,402]" degrees="107.13528" direction="east" orientation="longitude" pageId="9" precision="1" value="107.13528">107.13528E</geoCoordinate>
,
<date id="FFA2101DFFD70C581F8582678DE222C9" pageId="9" value="1994-02-14">
<collectingDate id="EFE6E9F5FFD70C581F8582678DE222C9" pageId="9" value="1994-02-14">14 February 1994</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581DFA82818FBB22AA" collectingDate="1994-02-06" collectionCode="CAS" collectorName="Se Khone" latitude="13.51678" location="Stung Treng" longLatPrecision="1" longitude="105.96678" pageId="9" specimenCode="CAS 91763, 4" specimenCount="1" stateProvince="Stung Treng">
<specimenCode id="DBBA9EA6FFD70C581DFA82818C7022C9" box="[214,341,409,431]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 91763</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581C4F82818C5622C9" box="[355,371,410,431]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">4</specimenCode>
,
<quantity id="4CE49B38FFD70C581CAE82818F0122C9" box="[386,548,410,431]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="4.92" metricValueMin="3.9799999999999995" pageId="9" unit="mm" value="44.5" valueMax="49.2" valueMin="39.8">39.849.2 mm</quantity>
SL,
<collectorName id="26E9530BFFD70C581F7482818FE622C9" box="[600,707,409,431]" pageId="9">Se Khone</collectorName>
at
<collectingRegion id="49D8F83FFFD70C581FC082818DEA22AA" country="Cambodia" name="Stung Treng" pageId="9">Stung Treng</collectingRegion>
,
<geoCoordinate id="EE28501AFFD70C581DF782AC8C7422AA" box="[219,337,439,461]" degrees="13.51678" direction="north" orientation="latitude" pageId="9" precision="1" value="13.51678">13.51678N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581C7182AC8CF922AA" box="[349,476,439,461]" degrees="105.96678" direction="east" orientation="longitude" pageId="9" precision="1" value="105.96678">105.96678E</geoCoordinate>
,
<date id="FFA2101DFFD70C581CC482AC8FBB22AA" box="[488,670,439,461]" pageId="9" value="1994-02-06">
<collectingDate id="EFE6E9F5FFD70C581CC482AC8FBB22AA" box="[488,670,439,461]" pageId="9" value="1994-02-06">6 February 1994</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581F8782AC8CCC2142" collectingDate="1994-02-12" collectionCode="CAS" collectorName="Rotanah Kiri &amp; O Changni" latitude="13.73948" location="Bang Lung" longLatPrecision="1" longitude="106.98738" municipality="Ann Long Mea" pageId="9" specimenCode="CAS 91765" specimenCount="1" stateProvince="Stung Treng">
<specimenCode id="DBBA9EA6FFD70C581F8782AC8E0222AB" box="[683,807,439,461]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 91765</specimenCode>
,
<quantity id="4CE49B38FFD70C581DA282CF8C3B228C" box="[142,286,468,490]" metricMagnitude="0" metricUnit="m" metricValue="1.1000999999999999" pageId="9" unit="mm" value="1100.1">1, 100.1 mm</quantity>
SL,
<collectorName id="26E9530BFFD70C581C7C82CF8CC5228C" box="[336,480,468,490]" pageId="9">Rotanah Kiri</collectorName>
,
<collectorName id="26E9530BFFD70C581CC082CF8F49228C" box="[492,620,468,490]" pageId="9">O Changni</collectorName>
, small stream on road from
<collectingMunicipality id="6BC7ACA7FFD70C581C3682E98CF22161" box="[282,471,498,519]" pageId="9">Ann Long Mea</collectingMunicipality>
to
<location id="8EC36006FFD70C581F3C82E98FB22161" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F3C82E98FB22161" box="[528,663,498,519]" latitude="13.73948" longLatPrecision="1" longitude="106.98738" municipality="Ann Long Mea" name="Bang Lung" pageId="9" stateProvince="Stung Treng">Bang Lung</location>
,
<geoCoordinate id="EE28501AFFD70C581F8182E98E022161" box="[685,807,498,520]" degrees="13.73948" direction="north" orientation="latitude" pageId="9" precision="1" value="13.73948">13.73948N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581DA281148C282143" box="[142,269,527,549]" degrees="106.98738" direction="east" orientation="longitude" pageId="9" precision="1" value="106.98738">106.98738E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C3181148CCC2142" box="[285,489,527,549]" pageId="9" value="1994-02-12">
<collectingDate id="EFE6E9F5FFD70C581C3181148CCC2142" box="[285,489,527,549]" pageId="9" value="1994-02-12">12 February 1994</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581CD5810B8CD52106" collectingDate="1996-01-25" collectionCode="UMMZ" location="Phnom Penh" municipality="Mekong River" pageId="9" specimenCode="UMMZ 234464, 2" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581CD5810B8F8D2143" box="[505,680,527,549]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 234464</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F9481148FED2142" box="[696,712,527,548]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">2</specimenCode>
,
<collectingRegion id="49D8F83FFFD70C581FF481148E0D2143" box="[728,808,527,549]" country="Cambodia" name="Kandal" pageId="9">Kandal</collectingRegion>
,
<collectingMunicipality id="6BC7ACA7FFD70C581DA281368C152125" box="[142,304,557,579]" pageId="9">Mekong River</collectingMunicipality>
at northern tip of island upstream from
<location id="8EC36006FFD70C581DA281518C002106" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581DA281518C002106" box="[142,293,586,608]" municipality="Mekong River" name="Phnom Penh" pageId="9" stateProvince="Kandal">Phnom Penh</location>
,
<date id="FFA2101DFFD70C581C1F81518CD52106" box="[307,496,586,608]" pageId="9" value="1996-01-25">
<collectingDate id="EFE6E9F5FFD70C581C1F81518CD52106" box="[307,496,586,608]" pageId="9" value="1996-01-25">25 January 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581CD381508C3C21FD" collectingDate="1996-03-01" collectionCode="UMMZ" county="Prek Muk" location="Phnom Penh" municipality="Tonle Sap" pageId="9" specimenCode="UMMZ 234727, 1" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581CD381508F892139" box="[511,684,586,608]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 234727</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F9681518FED2139" box="[698,712,586,607]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">1</specimenCode>
,
<collectingRegion id="49D8F83FFFD70C581FF481518E0D2106" box="[728,808,586,608]" country="Cambodia" name="Kandal" pageId="9">Kandal</collectingRegion>
,
<collectingCounty id="62C24E51FFD70C581DA281738DD92118" box="[142,252,616,638]" pageId="9">Prek Muk</collectingCounty>
<collectingRegion id="49D8F83FFFD70C581C2481738C7D2118" box="[264,344,616,638]" country="Cambodia" name="Kandal" pageId="9">Kandal</collectingRegion>
, across
<collectingMunicipality id="6BC7ACA7FFD70C581C9A81738F03211B" box="[438,550,616,638]" pageId="9">Tonle Sap</collectingMunicipality>
from
<location id="8EC36006FFD70C581F5881738E282118" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F5881738E282118" box="[628,781,616,638]" county="Prek Muk" municipality="Tonle Sap" name="Phnom Penh" pageId="9" stateProvince="Kandal">Phnom Penh</location>
,
<date id="FFA2101DFFD70C581E3281738C3C21FD" pageId="9" value="1996-03-01">
<collectingDate id="EFE6E9F5FFD70C581E3281738C3C21FD" pageId="9" value="1996-03-01">1 March 1996</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B743C80FFD70C581C0A819D8F1D21FD" box="[294,568,645,667]" collectorName="Lao" location="Mekong" pageId="9" specimenCount="1" stateProvince="Kandal">
<collectorName id="26E9530BFFD70C581C0A819D8C6B21FD" box="[294,334,646,667]" pageId="9">Lao</collectorName>
PDR:
<location id="8EC36006FFD70C581CBB819E8CD421FD" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581CBB819E8CD421FD" box="[407,497,645,667]" name="Mekong" pageId="9" stateProvince="Kandal">Mekong</location>
basin
</materialsCitation>
:
<materialsCitation id="3B743C80FFD70C581F69819E8C9A2192" collectingDate="1995-04-10" collectionCode="CAS" collectorName="Attapu" county="Houay" latitude="18.41558" location="Ban Hin Lat" longLatPrecision="1" longitude="103.60088" municipality="Xe Pian" pageId="9" specimenCode="CAS 92453, 2" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581F69819E8FE121FD" box="[581,708,645,667]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 92453</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581FFE819E8FC721FC" box="[722,738,645,666]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">2</specimenCode>
,
<quantity id="4CE49B38FFD70C581FDC819E8C2521DE" metricMagnitude="-2" metricUnit="m" metricValue="9.065" metricValueMax="10.45" metricValueMin="7.68" pageId="9" unit="mm" value="90.65" valueMax="104.5" valueMin="76.8">76.8 104.5 mm</quantity>
SL,
<collectorName id="26E9530BFFD70C581C0381B88C5921DE" box="[303,380,675,696]" pageId="9">Attapu</collectorName>
,
<collectingCounty id="62C24E51FFD70C581CA481B88CF521DE" box="[392,464,675,696]" pageId="9">Houay</collectingCounty>
Samong, about
<quantity id="4CE49B38FFD70C581FA981B88F9821DF" box="[645,701,675,697]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" unit="km" value="1.0">1 km</quantity>
upstream from its mouth into
<collectingMunicipality id="6BC7ACA7FFD70C581C5081DA8CF121B0" box="[380,468,704,726]" pageId="9">Xe Pian</collectingMunicipality>
near
<location id="8EC36006FFD70C581F3B81DA8F8621B0" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F3B81DA8F8621B0" box="[535,675,704,726]" county="Houay" latitude="18.41558" longLatPrecision="1" longitude="103.60088" municipality="Xe Pian" name="Ban Hin Lat" pageId="9" stateProvince="Kandal">Ban Hin Lat</location>
,
<geoCoordinate id="EE28501AFFD70C581F9D81DB8E0221B0" box="[689,807,704,726]" degrees="18.41558" direction="north" orientation="latitude" pageId="9" precision="1" value="18.41558">18.41558N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581DA281C58C282195" box="[142,269,733,756]" degrees="103.60088" direction="east" orientation="longitude" pageId="9" precision="1" value="103.60088">103.60088E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C3181C58C9A2192" box="[285,447,734,756]" pageId="9" value="1995-04-10">
<collectingDate id="EFE6E9F5FFD70C581C3181C58C9A2192" box="[285,447,734,756]" pageId="9" value="1995-04-10">10 April 1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581CE381C58FA42048" collectingDate="1995-04-14" collectionCode="CAS" county="Sekong watershed" location="Se Kaman" municipality="Se Kaman" pageId="9" specimenCode="CAS 92480, 10" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581CE381C58F6A2195" box="[463,591,734,755]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 92480</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F7381C58F592195" box="[607,636,734,755]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">10</specimenCode>
,
<collectingCounty id="62C24E51FFD70C581FA181C58DE12077" pageId="9">Sekong watershed</collectingCounty>
, small mountain stream entering right side of
<collectingMunicipality id="6BC7ACA7FFD70C581FEC81E08E092077" box="[704,812,763,785]" pageId="9">Se Kaman</collectingMunicipality>
just upstream from
<location id="8EC36006FFD70C581C4780028CFE2048" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581C4780028CFE2048" box="[363,475,793,814]" county="Sekong watershed" municipality="Se Kaman" name="Se Kaman" pageId="9" stateProvince="Kandal">Se Kaman</location>
,
<date id="FFA2101DFFD70C581CCB80028FA42048" box="[487,641,792,815]" pageId="9" value="1995-04-14">
<collectingDate id="EFE6E9F5FFD70C581CCB80028FA42048" box="[487,641,792,815]" pageId="9" value="1995-04-14">14 April 1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581FA280028F03200F" collectingDate="1993-06" collectionCode="CAS" collectorName="Bang Hang Khone" latitude="13.96758" location="Khone Falls" longLatPrecision="1" longitude="105.92988" municipality="Mekong River" pageId="9" specimenCode="CAS 94791, 8" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581FA280028E2E2048" box="[654,779,793,814]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 94791</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581E3B80028E032048" box="[791,806,793,814]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">8</specimenCode>
,
<collectingMunicipality id="6BC7ACA7FFD70C581DA2802D8C0F202A" box="[142,298,822,844]" pageId="9">Mekong River</collectingMunicipality>
, at
<collectorName id="26E9530BFFD70C581C7D802C8F3A202A" box="[337,543,822,844]" pageId="9">Bang Hang Khone</collectorName>
, just below
<location id="8EC36006FFD70C581F88802D8E0D202A" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F88802D8E0D202A" box="[676,808,822,844]" latitude="13.96758" longLatPrecision="1" longitude="105.92988" municipality="Mekong River" name="Khone Falls" pageId="9" stateProvince="Kandal">Khone Falls</location>
,
<geoCoordinate id="EE28501AFFD70C581DA2804F8C21200F" box="[142,260,851,874]" degrees="13.96758" direction="north" orientation="latitude" pageId="9" precision="1" value="13.96758">13.96758N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581C34804F8CB2200F" box="[280,407,851,874]" degrees="105.92988" direction="east" orientation="longitude" pageId="9" precision="1" value="105.92988">105.92988E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C87804F8F03200F" box="[427,550,852,873]" pageId="9" value="1993-06">
<collectingDate id="EFE6E9F5FFD70C581C87804F8F03200F" box="[427,550,852,873]" pageId="9" value="1993-06">June 1993</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581F16804F8C4F20E1" collectionCode="CAS" location="Nam Hinboun watershed" pageId="9" specimenCode="CAS 96192, 3" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581F16804F8F9B200C" box="[570,702,852,874]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 96192</specimenCode>
,
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,
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,
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at
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, just downstream from
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<specimenCode id="DBBA9EA6FFD70C581C8C80B68F6B20A4" box="[416,590,940,962]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 238636</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F7080B78F4F20A7" box="[604,618,940,961]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">1</specimenCode>
,
<collectorName id="26E9530BFFD70C581F5780B78FCD20A4" box="[635,744,940,962]" pageId="9">Vientiane</collectorName>
,
<collectorName id="26E9530BFFD70C581FDA80B68DED20B9" pageId="9">Nam Ngao</collectorName>
at south end of
<location id="8EC36006FFD70C581C8E80D18FBA2086" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581C8E80D18FBA2086" box="[418,671,970,992]" latitude="18.56488" longLatPrecision="1" longitude="102.63888" name="Nam Ngum Reservoir" pageId="9" stateProvince="Kandal">Nam Ngum Reservoir</location>
,
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,
<date id="FFA2101DFFD70C581C3280FC8CFB209A" box="[286,478,999,1021]" pageId="9" value="1997-02-05">
<collectingDate id="EFE6E9F5FFD70C581C3280FC8CFB209A" box="[286,478,999,1021]" pageId="9" value="1997-02-05">5 February 1997</collectingDate>
</date>
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;
<materialsCitation id="3B743C80FFD70C581CC380F38E09209B" box="[495,812,999,1021]" collectionCode="UMMZ" pageId="9" specimenCode="UMMZ 238669, 1" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581CC380F38FBA209B" box="[495,671,999,1021]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 238669</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F9C80FC8F9B209A" box="[688,702,999,1020]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">1</specimenCode>
, same as
</materialsCitation>
<materialsCitation id="3B743C80FFD70C581DA2871E8C1C277D" box="[142,313,1029,1051]" collectionCode="UMMZ" pageId="9" specimenCode="UMMZ 238636" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581DA2871E8C1C277D" box="[142,313,1029,1051]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 238636</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581C69871E8CF22733" collectingDate="1997-04-25" collectionCode="UMMZ" latitude="20.13338" location="Mekong River" longLatPrecision="1" longitude="100.38338" municipality="Bokeo Province" pageId="9" specimenCode="UMMZ 240585, 6" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581C69871E8CD5277C" box="[325,496,1028,1050]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 240585</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581CD0871E8F2E277D" box="[508,523,1029,1051]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">6</specimenCode>
,
<collectingMunicipality id="6BC7ACA7FFD70C581F34871E8FE3277D" box="[536,710,1029,1051]" pageId="9">Bokeo Province</collectingMunicipality>
,
<location id="8EC36006FFD70C581FFE871E8DEB275E" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581FFE871E8DEB275E" latitude="20.13338" longLatPrecision="1" longitude="100.38338" municipality="Bokeo Province" name="Mekong River" pageId="9" stateProvince="Kandal">Mekong River</location>
,
<locationDeviation id="9A112FFFFFD70C581DC887398FB4275E" box="[228,657,1058,1080]" location="Houay Xai" pageId="9">
<quantity id="4CE49B38FFD70C581DC887398C0C275E" box="[228,297,1058,1080]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" unit="km" value="5.0">5 km</quantity>
upstream from Houay Xai
</locationDeviation>
,
<geoCoordinate id="EE28501AFFD70C581F8487398E02275E" box="[680,807,1058,1080]" degrees="20.13338" direction="north" orientation="latitude" pageId="9" precision="1" value="20.13338">20.13338N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581DA2875B8C372733" box="[142,274,1087,1109]" degrees="100.38338" direction="east" orientation="longitude" pageId="9" precision="1" value="100.38338">100.38338E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C0A87248CF22733" box="[294,471,1087,1110]" pageId="9" value="1997-04-25">
<collectingDate id="EFE6E9F5FFD70C581C0A87248CF22733" box="[294,471,1087,1110]" pageId="9" value="1997-04-25">25 April 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581CC0875B8C9F27F6" collectingDate="1997-04-22" collectionCode="UMMZ" collectorName="Bokeo" latitude="20.36678" location="Mekong River" longLatPrecision="1" longitude="100.36678" pageId="9" specimenCode="UMMZ 241035, 1" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581CC0875B8F802733" box="[492,677,1087,1109]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 241035</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F95875B8FE22733" box="[697,711,1088,1109]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">1</specimenCode>
,
<collectorName id="26E9530BFFD70C581FF3875B8E022730" box="[735,807,1088,1110]" pageId="9">Bokeo</collectorName>
,
<location id="8EC36006FFD70C581DA287468C0E2715" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581DA287468C0E2715" box="[142,299,1117,1139]" latitude="20.36678" longLatPrecision="1" longitude="100.36678" name="Mekong River" pageId="9" stateProvince="Kandal">Mekong River</location>
,
<locationDeviation id="9A112FFFFFD70C581C1487468F812715" box="[312,676,1117,1139]" location="Houay Xai" pageId="9">
<quantity id="4CE49B38FFD70C581C1487468C572715" box="[312,370,1117,1139]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="9" unit="km" value="9.0">9 km</quantity>
upstream from Houay Xai
</locationDeviation>
,
<geoCoordinate id="EE28501AFFD70C581F9D87468E022715" box="[689,807,1117,1139]" degrees="20.36678" direction="north" orientation="latitude" pageId="9" precision="1" value="20.36678">20.36678N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581DA287618C2827F6" box="[142,269,1146,1168]" degrees="100.36678" direction="east" orientation="longitude" pageId="9" precision="1" value="100.36678">100.36678E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C3787608C9F27F6" box="[283,442,1147,1169]" pageId="9" value="1997-04-22">
<collectingDate id="EFE6E9F5FFD70C581C3787608C9F27F6" box="[283,442,1147,1169]" pageId="9" value="1997-04-22">22 April 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581CE487608DEF27AD" collectingDate="1997-04-26" collectionCode="UMMZ" collectorName="Bokeo &amp; Nam Tin &amp; Ban Paktin" latitude="20.18338" location="Nam Tin" longLatPrecision="52" longitude="100.558" pageId="9" specimenCode="UMMZ 241136, 2" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581CE487608F5327F6" box="[456,630,1146,1168]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 241136</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581FA887608FB127F6" box="[644,660,1147,1168]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">2</specimenCode>
,
<collectorName id="26E9530BFFD70C581F8E87608FCD27F7" box="[674,744,1147,1169]" pageId="9">Bokeo</collectorName>
,
<collectorName id="26E9530BFFD70C581FDA87608D9627C8" pageId="9">Nam Tin</collectorName>
at
<collectorName id="26E9530BFFD70C581DF787828C7C27C8" box="[219,345,1176,1198]" pageId="9">Ban Paktin</collectorName>
,
<location id="8EC36006FFD70C581C4A87828CE827C8" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581C4A87828CE827C8" box="[358,461,1176,1198]" latitude="20.18338" longLatPrecision="52" longitude="100.558" name="Nam Tin" pageId="9" stateProvince="Kandal">Nam Tin</location>
,
<geoCoordinate id="EE28501AFFD70C581CF787838F7427C8" box="[475,593,1176,1198]" degrees="20.18338" direction="north" orientation="latitude" pageId="9" precision="1" value="20.18338">20.18338N</geoCoordinate>
,
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,
<date id="FFA2101DFFD70C581FFC87838DEF27AD" pageId="9" value="1997-04-26">
<collectingDate id="EFE6E9F5FFD70C581FFC87838DEF27AD" pageId="9" value="1997-04-26">26 April 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581DCD87AD8F9D278E" collectingDate="1997-04-24" collectionCode="UMMZ" collectorName="Bokeo &amp; Nam Ngam" latitude="20.38338" location="Ban Namngion" longLatPrecision="1" longitude="100.36678" pageId="9" specimenCode="UMMZ 241148, 3" specimenCount="1" stateProvince="Kandal">
<specimenCode id="DBBA9EA6FFD70C581DCD87AD8CB827AD" box="[225,413,1206,1227]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 241148</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581C9F87AD8CE627AD" box="[435,451,1206,1227]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">3</specimenCode>
,
<collectorName id="26E9530BFFD70C581CF687AE8F0127AD" box="[474,548,1205,1227]" pageId="9">Bokeo</collectorName>
,
<collectorName id="26E9530BFFD70C581F1587AD8FE227AD" box="[569,711,1206,1227]" pageId="9">Nam Ngam</collectorName>
at
<location id="8EC36006FFD70C581E2C87AD8C22278F" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581E2C87AD8C22278F" latitude="20.38338" longLatPrecision="1" longitude="100.36678" name="Ban Namngion" pageId="9" stateProvince="Kandal">Ban Namngion</location>
,
<geoCoordinate id="EE28501AFFD70C581C3F87C88CAD278E" box="[275,392,1235,1257]" degrees="20.38338" direction="north" orientation="latitude" pageId="9" precision="1" value="20.38338">20.38338N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581CB887C88F36278E" box="[404,531,1235,1257]" degrees="100.36678" direction="east" orientation="longitude" pageId="9" precision="1" value="100.36678">100.36678E</geoCoordinate>
,
<date id="FFA2101DFFD70C581F3287C88F9D278E" box="[542,696,1235,1257]" pageId="9" value="1997-04-24">
<collectingDate id="EFE6E9F5FFD70C581F3287C88F9D278E" box="[542,696,1235,1257]" pageId="9" value="1997-04-24">24 April 1997</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B743C80FFD70C581FEE87C88C4E2660" country="Thailand" location="Chao Phraya" pageId="9" specimenCount="1">
<collectingCountry id="F30B764DFFD70C581FEE87C88E0D278F" box="[706,808,1235,1257]" name="Thailand" pageId="9">Thailand</collectingCountry>
:
<location id="8EC36006FFD70C581DA287EB8C042660" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581DA287EB8C042660" box="[142,289,1264,1286]" country="Thailand" name="Chao Phraya" pageId="9">Chao Phraya</location>
basin
</materialsCitation>
:
<materialsCitation id="3B743C80FFD70C581C5787EA8E022627" collectingDate="1933-01-01" collectionCode="ANSP" collectorName="Chieng Mai &amp; North" country="Thailand" latitude="19.21128" location="720 km N of Bangkok" longLatPrecision="1" longitude="98.97088" pageId="9" specimenCode="ANSP 58332" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581C5787EA8F2E2660" box="[379,523,1264,1286]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9">ANSP 58332</specimenCode>
,
<typeStatus id="54A7887FFFD70C581F3087EB8F5A2660" box="[540,639,1264,1286]" pageId="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1C4D5EFFD70C581F8187EB8C152642" authorityName="Fowler" authorityYear="1934" family="Cyprinidae" genus="Tylognathus" kingdom="Animalia" order="Cypriniformes" pageId="9" phylum="Chordata" rank="species" species="caudimaculatus">
<emphasis id="B968EACFFFD70C581F8187EB8C152642" italics="true" pageId="9">Tylognathus caudimaculatus</emphasis>
</taxonomicName>
,
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SL,
<collectorName id="26E9530BFFD70C581CF786158F462642" box="[475,611,1294,1316]" pageId="9">Chieng Mai</collectorName>
,
<collectorName id="26E9530BFFD70C581F5C86158F972642" box="[624,690,1294,1316]" pageId="9">North</collectorName>
<collectingCountry id="F30B764DFFD70C581F9086158FD12642" box="[700,756,1294,1316]" name="Thailand" pageId="9">Siam</collectingCountry>
,
<location id="8EC36006FFD70C581E2F86158C7A2627" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581E2F86158C7A2627" country="Thailand" latitude="19.21128" longLatPrecision="1" longitude="98.97088" name="720 km N of Bangkok" pageId="9" stateProvince="Krung Thep Mahanakhon">
<quantity id="4CE49B38FFD70C581E2F86158D942627" metricMagnitude="5" metricUnit="m" metricValue="7.2" pageId="9" unit="km" value="720.0">720 km</quantity>
N of
<collectingRegion id="49D8F83FFFD70C581DD186308C7A2627" box="[253,351,1323,1345]" country="Thailand" name="Krung Thep Mahanakhon" pageId="9">Bangkok</collectingRegion>
</location>
,
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,
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,
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<collectingDate id="EFE6E9F5FFD70C581F5986378E022627" box="[629,807,1324,1345]" pageId="9" value="1933-01-01">1 January 1933</collectingDate>
</date>
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;
<materialsCitation id="3B743C80FFD70C581DA286528F2E2638" box="[142,523,1353,1375]" collectionCode="ANSP" country="Thailand" pageId="9" specimenCode="ANSP 58333, 1" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="holotype">
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,
<specimenCode id="DBBA9EA6FFD70C581C0A86528C112638" box="[294,308,1353,1374]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9">1</specimenCode>
,
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, same as
</materialsCitation>
<materialsCitation id="3B743C80FFD70C581F3F86528FBA2639" box="[531,671,1353,1375]" collectionCode="ANSP" country="Thailand" pageId="9" specimenCode="ANSP 58332" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581F3F86528FBA2639" box="[531,671,1353,1375]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9">ANSP 58332</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581F8786528CEB26D1" collectingDate="1991-02-28" collectionCode="CAS" collectorName="Menam Wang" country="Thailand" latitude="18.93218" location="Lampang" longLatPrecision="1" longitude="99.64958" pageId="9" specimenCode="CAS 91781, 1" specimenCount="1" stateProvince="Lampang" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581F8786528E022638" box="[683,807,1353,1374]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 91781</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581DA2867C8DB9261A" box="[142,156,1383,1404]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">1</specimenCode>
,
<quantity id="4CE49B38FFD70C581D9C867D8C3F261A" box="[176,282,1382,1404]" metricMagnitude="-2" metricUnit="m" metricValue="5.33" pageId="9" unit="mm" value="53.3">53.3 mm</quantity>
SL,
<collectorName id="26E9530BFFD70C581C78867C8CDD261A" box="[340,504,1383,1404]" pageId="9">Menam Wang</collectorName>
,
<quantity id="4CE49B38FFD70C581F26867C8F70261A" box="[522,597,1382,1404]" metricMagnitude="4" metricUnit="m" metricValue="7.9" pageId="9" unit="km" value="79.0">79 km</quantity>
by road north of
<collectingRegion id="49D8F83FFFD70C581DA2869E8DD026FC" box="[142,245,1413,1434]" country="Thailand" name="Lampang" pageId="9">Lampang</collectingRegion>
and
<quantity id="4CE49B38FFD70C581C6C869F8CA426FC" box="[320,385,1412,1434]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="9" unit="km" value="6.0">6 km</quantity>
east of highway 1035,
<geoCoordinate id="EE28501AFFD70C581F9C869F8E0226FC" box="[688,807,1412,1434]" degrees="18.93218" direction="north" orientation="latitude" pageId="9" precision="1" value="18.93218">18.93218N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581DA286BA8DDA26D1" box="[142,255,1441,1463]" degrees="99.64958" direction="east" orientation="longitude" pageId="9" precision="1" value="99.64958">99.64958E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C2786B98CEB26D1" box="[267,462,1441,1463]" pageId="9" value="1991-02-28">
<collectingDate id="EFE6E9F5FFD70C581C2786B98CEB26D1" box="[267,462,1441,1463]" pageId="9" value="1991-02-28">28 February 1991</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581CF786B98C3C2694" collectingDate="1965-05-08" collectionCode="UMMZ" collectorName="Blaumipol Dam" country="Thailand" latitude="17.24068" location="Ping River" longLatPrecision="1" longitude="98.98158" pageId="9" specimenCode="UMMZ 236742, 19" specimenCount="1" stateProvince="Lampang" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581CF786B98FA026D1" box="[475,645,1441,1463]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 236742</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581FBD86B98F8B26D1" box="[657,686,1442,1463]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">19</specimenCode>
,
<location id="8EC36006FFD70C581F9786BA8E0926D1" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F9786BA8E0926D1" box="[699,812,1441,1463]" country="Thailand" latitude="17.24068" longLatPrecision="1" longitude="98.98158" name="Ping River" pageId="9" stateProvince="Lampang">Ping River</location>
below
<collectorName id="26E9530BFFD70C581DF486A48CA326B2" box="[216,390,1471,1493]" pageId="9">Blaumipol Dam</collectorName>
at guesthouse,
<geoCoordinate id="EE28501AFFD70C581F1886A48F8F26B2" box="[564,682,1471,1493]" degrees="17.24068" direction="north" orientation="latitude" pageId="9" precision="1" value="17.24068">17.24068N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581F9A86A48E0226B2" box="[694,807,1471,1493]" degrees="98.98158" direction="east" orientation="longitude" pageId="9" precision="1" value="98.98158">98.98158E</geoCoordinate>
,
<date id="FFA2101DFFD70C581DA286C68C3C2694" box="[142,281,1500,1522]" pageId="9" value="1965-05-08">
<collectingDate id="EFE6E9F5FFD70C581DA286C68C3C2694" box="[142,281,1500,1522]" pageId="9" value="1965-05-08">8 May 1965</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581C0486C68C43254B" collectingDate="1964-11-11" collectionCode="UMMZ" collectorName="Bung Borapet" country="Thailand" latitude="15.70918" location="Nau River" longLatPrecision="1" longitude="100.13548" pageId="9" specimenCode="UMMZ 236915, 1" specimenCount="1" stateProvince="Nakhon Sawan" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581C0486C68CF02694" box="[296,469,1500,1522]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">UMMZ 236915</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581CCF86C68CD42694" box="[483,497,1501,1522]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="9" type="University or college">1</specimenCode>
,
<collectorName id="26E9530BFFD70C581F2E86C68FB92694" box="[514,668,1501,1522]" pageId="9">Bung Borapet</collectorName>
, tributary to
<location id="8EC36006FFD70C581DA286E18DDB2576" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581DA286E18DDB2576" box="[142,254,1530,1552]" country="Thailand" latitude="15.70918" longLatPrecision="1" longitude="100.13548" name="Nau River" pageId="9" stateProvince="Nakhon Sawan">Nau River</location>
,
<quantity id="4CE49B38FFD70C581C2686E18C662576" box="[266,323,1530,1552]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" unit="km" value="4.0">4 km</quantity>
N.
<collectingRegion id="49D8F83FFFD70C581C4286E18F3C2569" box="[366,537,1530,1552]" country="Thailand" name="Nakhon Sawan" pageId="9">Nakhon Sawan</collectingRegion>
,
<geoCoordinate id="EE28501AFFD70C581F0A86E18FBE2569" box="[550,667,1530,1552]" degrees="15.70918" direction="north" orientation="latitude" pageId="9" precision="1" value="15.70918">15.70918N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581F8586E18E022569" box="[681,807,1530,1552]" degrees="100.13548" direction="east" orientation="longitude" pageId="9" precision="1" value="100.13548">100.13548E</geoCoordinate>
,
<date id="FFA2101DFFD70C581DA285038C43254B" box="[142,358,1559,1581]" pageId="9" value="1964-11-11">
<collectingDate id="EFE6E9F5FFD70C581DA285038C43254B" box="[142,358,1559,1581]" pageId="9" value="1964-11-11">11 November 1964</collectingDate>
</date>
</materialsCitation>
. Mekong basin:
<materialsCitation id="3B743C80FFD70C581F3185038F0B25E3" collectingDate="1990-05-12" collectionCode="CAS" collectorName="Huay Ngao &amp; Mekong" country="Thailand" latitude="20.1428" location="Chiang Khong" longLatPrecision="5" longitude="100.52698" municipality="Ban Chaem Pong" pageId="9" specimenCode="CAS 79169, 5" specimenCount="1" stateProvince="Nakhon Sawan" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581F3185038FBF254B" box="[541,666,1559,1581]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 79169</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F8A850C8F90254B" box="[678,693,1559,1581]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">5</specimenCode>
, mouth of
<collectorName id="26E9530BFFD70C581DA2852E8C2E252C" box="[142,267,1589,1610]" pageId="9">Huay Ngao</collectorName>
where it flows into
<collectorName id="26E9530BFFD70C581CDC852E8F68252D" box="[496,589,1589,1611]" pageId="9">Mekong</collectorName>
,
<quantity id="4CE49B38FFD70C581F75852E8FB6252D" box="[601,659,1589,1611]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" unit="km" value="1.0">1 km</quantity>
south of
<collectingMunicipality id="6BC7ACA7FFD70C581E2D852E8C0D250E" pageId="9">Ban Chaem Pong</collectingMunicipality>
(ca.
<quantity id="4CE49B38FFD70C581C4385488C9A250E" box="[367,447,1618,1640]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="9" unit="km" value="30.0">30 km</quantity>
southeast of
<location id="8EC36006FFD70C581F4385498E05250E" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F4385498E05250E" box="[623,800,1618,1640]" country="Thailand" latitude="20.1428" longLatPrecision="5" longitude="100.52698" municipality="Ban Chaem Pong" name="Chiang Khong" pageId="9" stateProvince="Nakhon Sawan">Chiang Khong</location>
),
<geoCoordinate id="EE28501AFFD70C581DA2856B8DD325E3" box="[142,246,1648,1670]" degrees="20.1428" direction="north" orientation="latitude" pageId="9" precision="5" value="20.1428">20.1428N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581C2A856B8CA025E3" box="[262,389,1648,1670]" degrees="100.52698" direction="east" orientation="longitude" pageId="9" precision="1" value="100.52698">100.52698E</geoCoordinate>
,
<date id="FFA2101DFFD70C581CB8856B8F0B25E3" box="[404,558,1648,1669]" pageId="9" value="1990-05-12">
<collectingDate id="EFE6E9F5FFD70C581CB8856B8F0B25E3" box="[404,558,1648,1669]" pageId="9" value="1990-05-12">12 May 1990</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581F12856B8F8725A6" collectingDate="1990-05-15" collectionCode="CAS" collectorName="Menam Kok" country="Thailand" latitude="20.0498" location="Tha Ton" longLatPrecision="5" longitude="99.38468" pageId="9" specimenCode="CAS 91764, 5" specimenCount="1" stateProvince="Nakhon Sawan" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581F12856B8F9B25E0" box="[574,702,1648,1670]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 91764</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581FE1856B8FFE25E0" box="[717,731,1648,1670]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">5</specimenCode>
(of 7),
<quantity id="4CE49B38FFD70C581DA285968C1525C5" box="[142,304,1677,1699]" metricMagnitude="-2" metricUnit="m" metricValue="5.42" metricValueMax="5.9799999999999995" metricValueMin="4.86" pageId="9" unit="mm" value="54.2" valueMax="59.8" valueMin="48.6">48.659.8 mm</quantity>
SL,
<collectorName id="26E9530BFFD70C581C4E85958CCE25C5" box="[354,491,1677,1699]" pageId="9">Menam Kok</collectorName>
at
<location id="8EC36006FFD70C581F3E85968F5525C5" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581F3E85968F5525C5" box="[530,624,1677,1699]" country="Thailand" latitude="20.0498" longLatPrecision="5" longitude="99.38468" name="Tha Ton" pageId="9" stateProvince="Nakhon Sawan">Tha Ton</location>
and up to
<quantity id="4CE49B38FFD70C581FDD85968E0E25C5" box="[753,811,1677,1699]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" unit="km" value="5.0">5 km</quantity>
downstream,
<geoCoordinate id="EE28501AFFD70C581C0F85B08CAE25A6" box="[291,395,1707,1729]" degrees="20.0498" direction="north" orientation="latitude" pageId="9" precision="5" value="20.0498">20.0498N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581CBA85B08F2225A6" box="[406,519,1707,1729]" degrees="99.38468" direction="east" orientation="longitude" pageId="9" precision="1" value="99.38468">99.38468E</geoCoordinate>
,
<date id="FFA2101DFFD70C581F3D85B08F8725A6" box="[529,674,1707,1729]" pageId="9" value="1990-05-15">
<collectingDate id="EFE6E9F5FFD70C581F3D85B08F8725A6" box="[529,674,1707,1729]" pageId="9" value="1990-05-15">15 May 1990</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581F8185B08C0D247E" collectingDate="1989-01-16" collectionCode="CAS" collectorName="Mekong" country="Thailand" latitude="20.17438" location="between Pak Ing and Jom Paeng" longLatPrecision="1" longitude="100.49018" pageId="9" specimenCode="CAS 91766, 9" specimenCount="1" stateProvince="Nakhon Sawan" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581F8185B08E0225A7" box="[685,807,1707,1729]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 91766</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581DA285D38DB825BB" box="[142,157,1736,1757]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">9</specimenCode>
,
<collectorName id="26E9530BFFD70C581D9C85D38C2F25B8" box="[176,266,1736,1758]" pageId="9">Mekong</collectorName>
mainstream
<location id="8EC36006FFD70C581C8485D38E0925B8" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581C8485D38E0925B8" box="[424,812,1736,1758]" country="Thailand" latitude="20.17438" longLatPrecision="1" longitude="100.49018" name="between Pak Ing and Jom Paeng" pageId="9" stateProvince="Nakhon Sawan">between Pak Ing and Jom Paeng</location>
(about
<quantity id="4CE49B38FFD70C581DC985FD8C1B259A" box="[229,318,1766,1788]" metricMagnitude="3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="9" unit="km" value="4.5" valueMax="5.0" valueMin="4.0">45 km</quantity>
downstream),
<geoCoordinate id="EE28501AFFD70C581CD885FD8F4F259D" box="[500,618,1765,1787]" degrees="20.17438" direction="north" orientation="latitude" pageId="9" precision="1" value="20.17438">20.17438N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581F5285FD8FD9259D" box="[638,764,1765,1787]" degrees="100.49018" direction="east" orientation="longitude" pageId="9" precision="1" value="100.49018">100.49018E</geoCoordinate>
,
<date id="FFA2101DFFD70C581E3C85FD8C0D247E" pageId="9" value="1989-01-16">
<collectingDate id="EFE6E9F5FFD70C581E3C85FD8C0D247E" pageId="9" value="1989-01-16">16 January 1989</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581C1584188C862435" collectingDate="1991-04-06" collectionCode="CAS" country="Thailand" latitude="16.82748" location="That Phanom" longLatPrecision="1" longitude="104.74998" municipality="Mekong River" pageId="9" specimenCode="CAS 92855, 1" specimenCount="1" stateProvince="Nakhon Sawan" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581C1584188C9F247F" box="[313,442,1795,1817]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 92855</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581CE784188CFC247E" box="[459,473,1795,1816]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">1</specimenCode>
,
<quantity id="4CE49B38FFD70C581CC7841F8F71247F" box="[491,596,1796,1817]" metricMagnitude="-2" metricUnit="m" metricValue="7.77" pageId="9" unit="mm" value="77.7">77.7 mm</quantity>
SL,
<collectingMunicipality id="6BC7ACA7FFD70C581FA184188E09247F" box="[653,812,1795,1817]" pageId="9">Mekong River</collectingMunicipality>
mainstream,
<quantity id="4CE49B38FFD70C581C0B843A8C532451" box="[295,374,1825,1847]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="9" unit="km" value="14.0">14 km</quantity>
south of
<location id="8EC36006FFD70C581CD6843A8FB82451" LSID="urn:lsid:plazi:treatment:03B587CBFFDB0C581E7785B688EC23E5:8EC36006FFD70C581CD6843A8FB82451" box="[506,669,1825,1847]" country="Thailand" latitude="16.82748" longLatPrecision="1" longitude="104.74998" municipality="Mekong River" name="That Phanom" pageId="9" stateProvince="Nakhon Sawan">That Phanom</location>
,
<geoCoordinate id="EE28501AFFD70C581F9D843A8E022450" box="[689,807,1824,1847]" degrees="16.82748" direction="north" orientation="latitude" pageId="9" precision="1" value="16.82748">16.82748N</geoCoordinate>
,
<geoCoordinate id="EE28501AFFD70C581DA284258C282432" box="[142,269,1854,1876]" degrees="104.74998" direction="east" orientation="longitude" pageId="9" precision="1" value="104.74998">104.74998E</geoCoordinate>
,
<date id="FFA2101DFFD70C581C3484258C862435" box="[280,419,1854,1876]" pageId="9" value="1991-04-06">
<collectingDate id="EFE6E9F5FFD70C581C3484258C862435" box="[280,419,1854,1876]" pageId="9" value="1991-04-06">6 April 1991</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B743C80FFD70C581C8284258F802432" box="[430,677,1854,1876]" collectionCode="CAS" country="Thailand" pageId="9" specimenCode="CAS 96196, 1" specimenCount="1" stateProvince="Nakhon Sawan" typeStatus="holotype">
<specimenCode id="DBBA9EA6FFD70C581C8284258F0F2432" box="[430,554,1854,1876]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">CAS 96196</specimenCode>
,
<specimenCode id="DBBA9EA6FFD70C581F1984258F662435" box="[565,579,1854,1875]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" type="Museum">1</specimenCode>
, same as
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