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<document id="3636D924DBB818A6AF5074031246E143" ID-CLB-Dataset="3058" ID-DOI="10.5252/geodiversitas2021v43a24" ID-GBIF-Dataset="d9b664cc-3197-44d5-92ca-08f8ce0fcc4f" ID-ISSN="1638-9395" ID-Zenodo-Dep="5764710" ID-ZooBank="urn:lsid:zoobank.org:pub:D36D1E14-73BE-4176-8024-F3673A65B8C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638894616877" checkinUser="felipe" docAuthor="Psarras, Christos, Koskeridou, Efterpi &amp; Merle, Didier" docDate="2021" docId="03B58177FFA36C73FEE441D4A24FF8D6" docLanguage="en" docName="Geodiversitas.43.24.1309-1339.pdf" docOrigin="Geodiversitas 43 (24)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Conus (Kalloconus) helladicus Psarras &amp; Koskeridou &amp; Merle 2021, n. sp." docType="treatment" docUuid="796AC37E-AC9E-48FE-89B6-2C11F282F6B5" docUuidSource="ZooBank" docVersion="11" lastPageNumber="1328" masterDocId="FF8CF90FFFB66C66FFE64527A673FFDB" masterDocTitle="Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 1: genera Conilithes Swainson, 1840 and Conus (Kalloconus) da Motta, 1991" masterLastPageNumber="1339" masterPageNumber="1309" pageNumber="1328" updateTime="1699472060009" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods id="27A2A5E1952A3ACF8F559DDA80A7A563" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C37F564E895D98B2BD36B34F422E2956">
<mods:title id="0E3074076ECE2677700A3D3C1169C5D3">Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 1: genera Conilithes Swainson, 1840 and Conus (Kalloconus) da Motta, 1991</mods:title>
</mods:titleInfo>
<mods:name id="40EBEB71E360A67A133F903ED29AB9C6" type="personal">
<mods:role id="22669E2EE05B0800483C6933B9432BDE">
<mods:roleTerm id="987BD6AB7D5D6E208E99F0B53D8AC17C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="938A7CA77BC9B16A49A21B7644390D37">Psarras, Christos</mods:namePart>
<mods:affiliation id="B7B2E128746EC63F1F77937C054D0A62">National and Kapodistrian University of Athens, Faculty of Geology &amp; Geoenvironment, Department of Hist. Geology-Paleontology, Athens (Greece)</mods:affiliation>
<mods:nameIdentifier id="480C9DD828255B3549472259EF00EBB6" type="email">cpsarras@geol.uoa.gr</mods:nameIdentifier>
</mods:name>
<mods:name id="5ACC3B852C7699FEE28853A0C007FD4B" type="personal">
<mods:role id="68623823B14E7981814D841B6E55713B">
<mods:roleTerm id="AA480997A73569FC9E72FB3BD613F357">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="167A0A622F0E066D92A8490F2D489812">Koskeridou, Efterpi</mods:namePart>
<mods:affiliation id="602653AE98C3FF52EDFF10D2B7F2960C">National and Kapodistrian University of Athens, Faculty of Geology &amp; Geoenvironment, Department of Hist. Geology-Paleontology, Athens (Greece)</mods:affiliation>
<mods:nameIdentifier id="4EF18D10B4C1BF4012DB711E109FC91B" type="email">ekosker@geol.uoa.gr</mods:nameIdentifier>
</mods:name>
<mods:name id="FDAD7996B3ECD0F42D1569A8BCC67174" type="personal">
<mods:role id="9629758B4D060994812B6C12BE461495">
<mods:roleTerm id="0543AEF891168B23465615B06C37245B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BC3AC902B6E944259DC17C94EE8C4D5">Merle, Didier</mods:namePart>
<mods:affiliation id="B828DE760511F9DEC9C078102118C0FD">CR 2 P (CNRS, MNHN, UPMC, Sorbonne Université), Département Origines et Évolution, Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
<mods:nameIdentifier id="1CF2C94C51B87B4E1EC5B9F6323BFBD2" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="54E9C1322345936790A9ED499426B19F">text</mods:typeOfResource>
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<mods:title id="2397EC2934CADEB70C6A036E4E7884C8">Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part id="7FA3619499D326949FA23FA6968EA41A">
<mods:date id="2A5463C43E4F9126EF0CCA4645CFD4E5">2021</mods:date>
<mods:detail id="7F17CA45844A35B937E9BFEC33CE8F61" type="pubDate">
<mods:number id="5AE7B2CB1CBAAFFAD82CF2B02A6A7B6C">2021-12-02</mods:number>
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<mods:detail id="5BC6B448CEBD8EAA1060687B4E4546D7" type="volume">
<mods:number id="DDF7B6F38F75674948D0DB6D1868ABE1">43</mods:number>
</mods:detail>
<mods:detail id="7B0266DB516126111379A6D8F17C0A1E" type="issue">
<mods:number id="9724E06D40747A4B7D771BD7E7054655">24</mods:number>
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<mods:start id="AFC577141843E7671391B1A9D0AD85E0">1309</mods:start>
<mods:end id="B2206535F7F02C07B2257C1F84DEDD6C">1339</mods:end>
</mods:extent>
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<mods:classification id="64D11C6FDF19ACAEE5715946771DDE4C">journal article</mods:classification>
<mods:identifier id="BC234A4FC4137453F86DAB2D7BDA693E" type="CLB-Dataset">3058</mods:identifier>
<mods:identifier id="D5EE3F41CE99D67A1276301057285DC7" type="DOI">10.5252/geodiversitas2021v43a24</mods:identifier>
<mods:identifier id="085E9EA3C9512ED99F85769A98470E60" type="GBIF-Dataset">d9b664cc-3197-44d5-92ca-08f8ce0fcc4f</mods:identifier>
<mods:identifier id="AFB6EF7A723760481EB710866534FB10" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03B58177FFA36C73FEE441D4A24FF8D6" ID-DOI="http://doi.org/10.5281/zenodo.5768074" ID-GBIF-Taxon="191256706" ID-Zenodo-Dep="5768074" LSID="urn:lsid:zoobank.org:act:796AC37E-AC9E-48FE-89B6-2C11F282F6B5" httpUri="http://treatment.plazi.org/id/03B58177FFA36C73FEE441D4A24FF8D6" lastPageNumber="1328" pageId="21" pageNumber="1328">
<subSubSection id="C30663EAFFA36C73FEE441D4A4FAFAD5" box="[258,649,1267,1294]" pageId="21" pageNumber="1328" type="nomenclature">
<paragraph id="8BA33061FFA36C73FEE441D4A4FAFAD5" blockId="21.[258,649,1267,1326]" box="[258,649,1267,1294]" pageId="21" pageNumber="1328">
<heading id="D0EB870DFFA36C73FEE441D4A4FAFAD5" box="[258,649,1267,1294]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="1328" reason="2">
<taxonomicName id="4C1C4BE2FFA36C73FEE441D4A434FAD5" authority="Psarras &amp; Koskeridou &amp; Merle, 2021" authorityName="Psarras &amp; Koskeridou &amp; Merle" authorityYear="2021" box="[258,583,1267,1294]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="helladicus" status="sp. nov." subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FEE441D4A735FAD6" bold="true" box="[258,326,1267,1293]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FEB041D4A7BFFAD5" bold="true" box="[342,460,1267,1294]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FE3A41D4A434FAD5" bold="true" box="[476,583,1267,1294]" italics="true" pageId="21" pageNumber="1328">helladicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5108FFA36C73FDA841D3A4FAFAD5" box="[590,649,1268,1294]" pageId="21" pageNumber="1328" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30663EAFFA36C73FE8E4033A574FAB2" pageId="21" pageNumber="1328" type="description">
<paragraph id="8BA33061FFA36C73FE8E4033A450FAF5" blockId="21.[258,649,1267,1326]" box="[360,547,1299,1326]" pageId="21" pageNumber="1328">
(
<figureCitation id="13272CE4FFA36C73FE974034A7CEFAF5" box="[369,445,1299,1326]" captionStart="FIG" captionStartId="22.[132,143,1355,1372]" captionTargetBox="[132,1455,215,1313]" captionTargetId="figure-311@22.[132,1455,215,1313]" captionTargetPageId="22" captionText="FIG. 18. — Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece): A, Paratype MNHN.F.A72638: a solid club shaped shell with large relative diameter (form 1) and a distinct colour pattern; B, Holotype AMPG(IV) 2660, Psalidha: a low spired specimen with a relatively narrow diameter (form 2), with a distinct colour pattern; C, Paratype MNHN.F.A72636: a small-sized shell with large relative diameter (form 1); D, Paratype MNHN.F.A72637: a flat spired conid with slightly protruding early spire whorls (form 2). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5764746" httpUri="https://zenodo.org/record/5764746/files/figure.png" pageId="21" pageNumber="1328">Fig. 18</figureCitation>
;
<tableCitation id="C69E05DAFFA36C73FE2C4034A469FAF5" box="[458,538,1299,1326]" captionStart="TABLE" captionStartId="22.[132,143,1501,1518]" captionTargetPageId="22" captionText="TABLE 8. — Shell measurements and ratios of Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece). Mean and standard deviation are computed from nine specimens, the largest being the holotype (AMPG(IV) 2660)." httpUri="http://table.plazi.org/id/DF6360E9FFA06C70FF6240FAA573F9DE" pageId="21" pageNumber="1328" tableUuid="DF6360E9FFA06C70FF6240FAA573F9DE">Table 8</tableCitation>
)
</paragraph>
<paragraph id="8BA33061FFA36C73FF62407FA574FAB2" blockId="21.[132,775,1368,1385]" box="[132,775,1368,1385]" pageId="21" pageNumber="1328">
<uri id="FF8D3C63FFA36C73FF62407FA574FAB2" box="[132,775,1368,1385]" pageId="21" pageNumber="1328">
urn:lsid:zoobank.org:act:
<uuid id="FFBA0AB4FFA36C73FE86407FA574FAB2" box="[352,775,1368,1385]" pageId="21" pageNumber="1328">796AC37E-AC9E-48FE-89B6-2C11F282F6B5</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30663EAFFA36C73FF6240BBA6C8FA14" pageId="21" pageNumber="1328" type="reference_group">
<paragraph id="8BA33061FFA36C73FF6240BBA6C8FA14" blockId="21.[131,775,1436,1487]" pageId="21" pageNumber="1328">
<taxonomicName id="4C1C4BE2FFA36C73FF6240BBA727FA6F" ID-CoL="7RS7G" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1879" box="[132,340,1436,1460]" class="Gastropoda" family="Conidae" genus="Kalloconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="hungaricus">
<emphasis id="B968EC73FFA36C73FF6240BBA727FA6F" box="[132,340,1436,1460]" italics="true" pageId="21" pageNumber="1328">Kalloconus hungaricus</emphasis>
</taxonomicName>
–
<bibRefCitation id="EF8D4D90FFA36C73FE8E40BBA46EFA6E" author="LANDAU B. &amp; HARZHAUSER M. &amp; BUYUKMERIC Y. &amp; SILVA C." box="[360,541,1436,1461]" pageId="21" pageNumber="1328" pagination="1 - 13" refId="ref19741" refString="LANDAU B., HARZHAUSER M., BUYUKMERIC Y. &amp; SILVA C. 2013. - Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research 1: 1 - 13." type="journal article" year="2013">
Landau
<emphasis id="B968EC73FFA36C73FE5E40BAA795FA6F" box="[440,486,1436,1460]" italics="true" pageId="21" pageNumber="1328">et al.</emphasis>
2013
</bibRefCitation>
: pl. 37, figs 9-10, pl. 38, fig. 1.
</paragraph>
</subSubSection>
<subSubSection id="C30663EAFFA36C73FF6240CDA7A4F9ED" pageId="21" pageNumber="1328" type="diagnosis">
<paragraph id="8BA33061FFA36C73FF6240CDA7A4F9ED" blockId="21.[131,775,1513,1590]" pageId="21" pageNumber="1328">DIAGNOSIS. — A medium-sized shell, with almost flat spire whorls and a colour pattern bearing wide flammulae on spire whorls, with spiral rows of dashes on last whorl.</paragraph>
</subSubSection>
<subSubSection id="C30663EAFFA36C73FF624376A57BF830" pageId="21" pageNumber="1328" type="materials_examined">
<paragraph id="8BA33061FFA36C73FF624376A4A2F858" blockId="21.[130,776,1616,2027]" pageId="21" pageNumber="1328">
<typeStatus id="54A78EC3FFA36C73FF624376A6C6F9BD" box="[132,181,1617,1640]" pageId="21" pageNumber="1328">TYPE</typeStatus>
MATERIAL. —
<materialsCitation id="3B743A3CFFA36C73FEDD4377A57BF9B3" ID-GBIF-Occurrence="3776293314" box="[315,776,1616,1640]" collectionCode="AMPG" country="Greece" latitude="35.085583" location="Psalidha" longLatPrecision="1" longitude="24.962778" pageId="21" pageNumber="1328" specimenCode="AMPG(IV) 2660" specimenCount="1" stateProvince="Crete" typeStatus="holotype">
<typeStatus id="54A78EC3FFA36C73FEDD4377A7EBF9B3" box="[315,408,1616,1640]" pageId="21" pageNumber="1328" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBBA981AFFA36C73FE464377A434F9B3" box="[416,583,1616,1640]" pageId="21" pageNumber="1328">AMPG(IV) 2660</specimenCode>
,
<location id="8EC366BAFFA36C73FDA94377A4EEF9B3" LSID="urn:lsid:plazi:treatment:03B58177FFA36C73FEE441D4A24FF8D6:8EC366BAFFA36C73FDA94377A4EEF9B3" box="[591,669,1616,1640]" name="Psalidha" pageId="21" pageNumber="1328">Psalidha</location>
(
<figureCitation id="13272CE4FFA36C73FD4F4377A48FF9B3" box="[681,764,1616,1640]" captionStart="FIG" captionStartId="22.[132,143,1355,1372]" captionTargetBox="[132,1455,215,1313]" captionTargetId="figure-311@22.[132,1455,215,1313]" captionTargetPageId="22" captionText="FIG. 18. — Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece): A, Paratype MNHN.F.A72638: a solid club shaped shell with large relative diameter (form 1) and a distinct colour pattern; B, Holotype AMPG(IV) 2660, Psalidha: a low spired specimen with a relatively narrow diameter (form 2), with a distinct colour pattern; C, Paratype MNHN.F.A72636: a small-sized shell with large relative diameter (form 1); D, Paratype MNHN.F.A72637: a flat spired conid with slightly protruding early spire whorls (form 2). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5764746" httpUri="https://zenodo.org/record/5764746/files/figure.png" pageId="21" pageNumber="1328">Fig. 18B</figureCitation>
).
</materialsCitation>
<materialsCitation id="3B743A3CFFA36C73FF62434CA57BF8C7" ID-GBIF-Occurrence="3776293319" collectionCode="MNHN" country="Greece" latitude="35.085583" location="Psalidha" longLatPrecision="1" longitude="24.962778" pageId="21" pageNumber="1328" specimenCode="MNHN.F.A72636, MNHN.F.A72637, MNHN.F.A72638" specimenCount="3" stateProvince="Crete" typeStatus="paratype">
<specimenCount id="9D1AFBE8FFA36C73FF62434CA6C8F958" box="[132,187,1643,1667]" pageId="21" pageNumber="1328">Three</specimenCount>
<typeStatus id="54A78EC3FFA36C73FF25434CA752F958" box="[195,289,1643,1667]" pageId="21" pageNumber="1328" type="paratype">paratypes</typeStatus>
,
<specimenCode id="DBBA981AFFA36C73FEC9434CA794F958" box="[303,487,1643,1667]" pageId="21" pageNumber="1328">
MNHN.F.
<accessionNumber id="944FAD82FFA36C73FE71434CA794F958" box="[407,487,1643,1667]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A72636" pageId="21" pageNumber="1328" type="EnaNcbi">A72636</accessionNumber>
</specimenCode>
to
<specimenCode id="DBBA981AFFA36C73FDEA434CA4B7F958" box="[524,708,1643,1667]" pageId="21" pageNumber="1328">
MNHN.F.
<accessionNumber id="944FAD82FFA36C73FD92434CA4B7F958" box="[628,708,1643,1667]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A72638" pageId="21" pageNumber="1328" type="EnaNcbi">A72638</accessionNumber>
</specimenCode>
,
<collectingRegion id="49D8FE83FFA36C73FD37434CA575F958" box="[721,774,1643,1667]" country="Greece" name="Kriti" pageId="21" pageNumber="1328">Crete</collectingRegion>
(
<figureCitation id="13272CE4FFA36C73FF6A43A2A751F946" box="[140,290,1669,1693]" captionStart="FIG" captionStartId="22.[132,143,1355,1372]" captionTargetBox="[132,1455,215,1313]" captionTargetId="figure-311@22.[132,1455,215,1313]" captionTargetPageId="22" captionText="FIG. 18. — Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece): A, Paratype MNHN.F.A72638: a solid club shaped shell with large relative diameter (form 1) and a distinct colour pattern; B, Holotype AMPG(IV) 2660, Psalidha: a low spired specimen with a relatively narrow diameter (form 2), with a distinct colour pattern; C, Paratype MNHN.F.A72636: a small-sized shell with large relative diameter (form 1); D, Paratype MNHN.F.A72637: a flat spired conid with slightly protruding early spire whorls (form 2). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5764746" httpUri="https://zenodo.org/record/5764746/files/figure.png" pageId="21" pageNumber="1328">Fig. 18A, C, D</figureCitation>
). TYPE LOCALITY. —
<location id="8EC366BAFFA36C73FEDA4390A7FDF914" LSID="urn:lsid:plazi:treatment:03B58177FFA36C73FEE441D4A24FF8D6:8EC366BAFFA36C73FEDA4390A7FDF914" box="[316,398,1719,1743]" country="Greece" latitude="35.085583" longLatPrecision="1" longitude="24.962778" name="Psalidha" pageId="21" pageNumber="1328" stateProvince="Crete">Psalidha</location>
,
<geoCoordinate id="EE2856A6FFA36C73FE734390A452F914" box="[405,545,1719,1743]" degrees="35" direction="north" minutes="05" orientation="latitude" pageId="21" pageNumber="1328" precision="1" seconds="08.1" value="35.085583">35°05’08.1”N</geoCoordinate>
,
<geoCoordinate id="EE2856A6FFA36C73FDCF4390A4DDF914" box="[553,686,1719,1743]" degrees="24" direction="east" minutes="57" orientation="longitude" pageId="21" pageNumber="1328" precision="1" seconds="46.0" value="24.962778">24°57’46.0”E</geoCoordinate>
, Messara Basin, Tortonian, Crete,
<collectingCountry id="F30B70F1FFA36C73FE9243F5A7C9F931" box="[372,442,1746,1770]" name="Greece" pageId="21" pageNumber="1328">Greece</collectingCountry>
. STRATIGRAPHIC RANGE. —
<geologicalTimeScale id="F93CB0CAFFA36C73FE9D4223A577F8C7" box="[379,772,1796,1820]" pageId="21" pageNumber="1328">Tortonian of Greece (Messara Basin, Crete)</geologicalTimeScale>
.
</materialsCitation>
ETYMOLOGY. — As
<taxonomicName id="4C1C4BE2FFA36C73FEDC4210A6C6F8B2" authority="Hoernes &amp; Auinger, 1879" authorityName="Hoernes &amp; Auinger" authorityYear="1879" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="hungaricus" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FEDC4210A701F895" box="[314,370,1847,1870]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FE984211A7AEF895" box="[382,477,1846,1870]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FE0F4211A439F895" box="[489,586,1846,1870]" italics="true" pageId="21" pageNumber="1328">hungaricus</emphasis>
<bibRefCitation id="EF8D4D90FFA36C73FDAB4210A6C6F8B2" author="HOERNES R. &amp; AUINGER M." pageId="21" pageNumber="1328" pagination="1 - 52" refId="ref19161" refString="HOERNES R. &amp; AUINGER M. 1879. - Die Gasteropoden der Meeres- Ablagerungen der ersten und zweiten Miocanen Mediterran-Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt 12: 1 - 52." type="journal article" year="1879">Hoernes &amp; Auinger, 1879</bibRefCitation>
</taxonomicName>
was first found in
<collectingCountry id="F30B70F1FFA36C73FE954276A7B8F8B2" box="[371,459,1873,1897]" name="Hungary" pageId="21" pageNumber="1328">Hungary</collectingCountry>
, we propose
<taxonomicName id="4C1C4BE2FFA36C73FDB74276A6ACF858" authority="Psarras &amp; Koskeridou &amp; Merle, 2021" authorityName="Psarras &amp; Koskeridou &amp; Merle" authorityYear="2021" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="helladicus" status="sp. nov." subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FDB74276A4FFF8B3" box="[593,652,1873,1896]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FD7C4276A48CF8B2" box="[666,767,1873,1897]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FF62424CA6ACF858" box="[132,223,1899,1923]" italics="true" pageId="21" pageNumber="1328">helladicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5108FFA36C73FF00424BA76EF858" box="[230,285,1900,1923]" pageId="21" pageNumber="1328" rank="species">n. sp.</taxonomicNameLabel>
, a species found in
<collectingCountry id="F30B70F1FFA36C73FE3A424BA46CF85F" box="[476,543,1900,1924]" name="Greece" pageId="21" pageNumber="1328">Greece</collectingCountry>
(
<emphasis id="B968EC73FFA36C73FDCB424CA414F858" box="[557,615,1899,1923]" italics="true" pageId="21" pageNumber="1328">
<collectingCountry id="F30B70F1FFA36C73FDCB424CA414F858" box="[557,615,1899,1923]" name="Greece" pageId="21" pageNumber="1328">Hellas</collectingCountry>
</emphasis>
in Greek).
</paragraph>
<paragraph id="8BA33061FFA36C73FF6242B9A57BF830" blockId="21.[130,776,1616,2027]" pageId="21" pageNumber="1328">
OTHER MATERIAL EXAMINED. —
<materialsCitation id="3B743A3CFFA36C73FE0A42B8A45EF80B" ID-GBIF-Occurrence="3776293305" collectionCode="AMPG" country="Greece" location="Tefeli" pageId="21" pageNumber="1328" specimenCode="AMPG(IV) 2661, AMPG(IV) 2662, AMPG(IV) 2676, AMPG(IV) 2677" specimenCount="4" stateProvince="Tefeli">
<emphasis id="B968EC73FFA36C73FE0A42B8A447F86C" bold="true" box="[492,564,1951,1975]" pageId="21" pageNumber="1328">
<collectingCountry id="F30B70F1FFA36C73FE0A42B8A447F86C" box="[492,564,1951,1975]" name="Greece" pageId="21" pageNumber="1328">Greece</collectingCountry>
</emphasis>
.
<collectingRegion id="49D8FE83FFA36C73FDA142B9A4F6F86D" box="[583,645,1950,1974]" pageId="21" pageNumber="1328">Tefeli</collectingRegion>
:
<specimenCount id="9D1AFBE8FFA36C73FD7542B9A6CBF80B" pageId="21" pageNumber="1328" type="generic">four specimens</specimenCount>
<specimenCode id="DBBA981AFFA36C73FF27429FA7DCF80B" box="[193,431,1976,2000]" pageId="21" pageNumber="1328">AMPG(IV) 2661-2662</specimenCode>
,
<specimenCode id="DBBA981AFFA36C73FE5D429FA45EF80B" box="[443,557,1976,2000]" pageId="21" pageNumber="1328">2676-2677</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B743A3CFFA36C73FDDF429FA577F830" ID-GBIF-Occurrence="3776293315" collectionCode="MNHN" country="Greece" location="Crete" pageId="21" pageNumber="1328" specimenCode="MNHN.F.A72639" specimenCount="1" stateProvince="Crete">
<collectingRegion id="49D8FE83FFA36C73FDDF429FA400F80B" box="[569,627,1976,2000]" country="Greece" name="Kriti" pageId="21" pageNumber="1328">Crete</collectingRegion>
:
<specimenCount id="9D1AFBE8FFA36C73FD98429EA574F80B" box="[638,775,1976,2000]" pageId="21" pageNumber="1328" type="generic">one specimen</specimenCount>
(
<specimenCode id="DBBA981AFFA36C73FF6D42F4A746F831" box="[139,309,2002,2026]" pageId="21" pageNumber="1328">
MNHN.F.
<accessionNumber id="944FAD82FFA36C73FF0B42F5A746F831" box="[237,309,2002,2026]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A72639" pageId="21" pageNumber="1328" type="EnaNcbi">A72639</accessionNumber>
</specimenCode>
). All of them display a colour patterns under UV light
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30663EAFFA36C73FCCB45F0A5BAFC54" pageId="21" pageNumber="1328" type="description">
<paragraph id="8BA33061FFA36C73FCCB45F0A271FF2B" blockId="21.[811,1457,215,656]" box="[813,1026,215,241]" pageId="21" pageNumber="1328">SHELL DESCRIPTION</paragraph>
<paragraph id="8BA33061FFA36C73FCCB45D0A21EFD4B" blockId="21.[811,1457,215,656]" pageId="21" pageNumber="1328">Medium-sized, robust shells, with relatively low spired whorls. Early spire whorls coeloconoid. Last spire whorls, smooth, straight to concave, creating a low conical to flat outline. Suture impressed. Subsutural flexure shallow, weakly curved, moderately asymmetrical. Shoulder rounded, protruded, creating a bulky outline. Maximum diameter below shoulder. Last whorl straight. Aperture moderate, narrow near suture, straight. Apertural canal wide, fasciole twisted, demarcated from base and inner lip. There are two extreme forms. Form 1 consists of robust forms which are relatively wider in comparison to form 2 and have low angled spire whorls. Form 2 consists of relatively elongated forms with flat spire whorl. Intermediate forms also exist.</paragraph>
<paragraph id="8BA33061FFA36C73FCCB4791A2D4FD15" blockId="21.[813,1457,694,911]" box="[813,1191,694,720]" pageId="21" pageNumber="1328">DESCRIPTION OF COLOUR PATTERN</paragraph>
<paragraph id="8BA33061FFA36C73FCCB47F1A5BAFC54" blockId="21.[813,1457,694,911]" pageId="21" pageNumber="1328">The colour pattern consists of one layer of short and long, fluorescent, spiral dashes, arranged in evenly spaced spiral rows. The spire whorls display wide, fluorescent flammulae, with irregular boundaries on a non-fluorescent base colour. The flammulae do not connect with the colour pattern of the last whorl.</paragraph>
</subSubSection>
<subSubSection id="C30663EAFFA36C73FCCB4691A24FF8D6" pageId="21" pageNumber="1328" type="discussion">
<paragraph id="8BA33061FFA36C73FCCB4691A5E2FC15" blockId="21.[811,1457,950,1805]" box="[813,913,950,976]" pageId="21" pageNumber="1328">REMARKS</paragraph>
<paragraph id="8BA33061FFA36C73FCCB46F2A2A8FBB4" blockId="21.[811,1457,950,1805]" pageId="21" pageNumber="1328">
This species shows some variations in the relative diameter of its spire whorls (
<tableCitation id="C69E05DAFFA36C73FC3446D2A26DFBD4" box="[978,1054,1013,1039]" captionStart="TABLE" captionStartId="22.[132,143,1501,1518]" captionTargetPageId="22" captionText="TABLE 8. — Shell measurements and ratios of Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece). Mean and standard deviation are computed from nine specimens, the largest being the holotype (AMPG(IV) 2660)." httpUri="http://table.plazi.org/id/DF6360E9FFA06C70FF6240FAA573F9DE" pageId="21" pageNumber="1328" tableUuid="DF6360E9FFA06C70FF6240FAA573F9DE">Table 8</tableCitation>
). The difference between the elongated and robust forms is not very variable. However intermediate forms (
<figureCitation id="13272CE4FFA36C73FC9F4112A585FB94" box="[889,1014,1077,1103]" captionStart="FIG" captionStartId="22.[132,143,1355,1372]" captionTargetBox="[132,1455,215,1313]" captionTargetId="figure-311@22.[132,1455,215,1313]" captionTargetPageId="22" captionText="FIG. 18. — Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece): A, Paratype MNHN.F.A72638: a solid club shaped shell with large relative diameter (form 1) and a distinct colour pattern; B, Holotype AMPG(IV) 2660, Psalidha: a low spired specimen with a relatively narrow diameter (form 2), with a distinct colour pattern; C, Paratype MNHN.F.A72636: a small-sized shell with large relative diameter (form 1); D, Paratype MNHN.F.A72637: a flat spired conid with slightly protruding early spire whorls (form 2). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5764746" httpUri="https://zenodo.org/record/5764746/files/figure.png" pageId="21" pageNumber="1328">Fig. 18B, C</figureCitation>
) between both forms (
<figureCitation id="13272CE4FFA36C73FB0E4112A315FB94" box="[1256,1382,1077,1103]" captionStart="FIG" captionStartId="22.[132,143,1355,1372]" captionTargetBox="[132,1455,215,1313]" captionTargetId="figure-311@22.[132,1455,215,1313]" captionTargetPageId="22" captionText="FIG. 18. — Conus (Kalloconus) helladicus n. sp. from the Tortonian of Crete (Greece): A, Paratype MNHN.F.A72638: a solid club shaped shell with large relative diameter (form 1) and a distinct colour pattern; B, Holotype AMPG(IV) 2660, Psalidha: a low spired specimen with a relatively narrow diameter (form 2), with a distinct colour pattern; C, Paratype MNHN.F.A72636: a small-sized shell with large relative diameter (form 1); D, Paratype MNHN.F.A72637: a flat spired conid with slightly protruding early spire whorls (form 2). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5764746" httpUri="https://zenodo.org/record/5764746/files/figure.png" pageId="21" pageNumber="1328">Fig. 18A, D</figureCitation>
) point towards the existence of a single species.
</paragraph>
<paragraph id="8BA33061FFA36C73FCA54152A240FAD4" blockId="21.[811,1457,950,1805]" pageId="21" pageNumber="1328">
<taxonomicName id="4C1C4BE2FFA36C73FCA54152A2E1FB54" authorityName="Hoernes &amp; Auinger" authorityYear="1879" box="[835,1170,1141,1167]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="hungaricus" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FCA54152A5F5FB54" box="[835,902,1141,1167]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FC7F4152A27FFB54" box="[921,1036,1141,1167]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FBF84152A2E1FB54" box="[1054,1170,1141,1167]" italics="true" pageId="21" pageNumber="1328">hungaricus</emphasis>
</taxonomicName>
specimens sensu
<bibRefCitation id="EF8D4D90FFA36C73FABB4152A5DBFB74" author="LANDAU B. &amp; HARZHAUSER M. &amp; BUYUKMERIC Y. &amp; SILVA C." pageId="21" pageNumber="1328" pagination="1 - 13" refId="ref19741" refString="LANDAU B., HARZHAUSER M., BUYUKMERIC Y. &amp; SILVA C. 2013. - Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research 1: 1 - 13." type="journal article" year="2013">
Landau
<emphasis id="B968EC73FFA36C73FCCB41B2A512FB74" box="[813,865,1173,1199]" italics="true" pageId="21" pageNumber="1328">et al.</emphasis>
(2013
</bibRefCitation>
: pl. 37, figs 9, 10, pl. 38, fig. 1) from the
<collectingRegion id="49D8FE83FFA36C73FA9541B2A52EFB14" country="Turkey" name="Karaman" pageId="21" pageNumber="1328">Karaman</collectingRegion>
Basin (
<collectingCountry id="F30B70F1FFA36C73FC4D4193A589FB14" box="[939,1018,1204,1231]" name="Turkey" pageId="21" pageNumber="1328">Turkey</collectingCountry>
) are more likely to be
<taxonomicName id="4C1C4BE2FFA36C73FB004192A5E1FB35" authority="Psarras &amp; Koskeridou &amp; Merle, 2021" authorityName="Psarras &amp; Koskeridou &amp; Merle" authorityYear="2021" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="helladicus" status="sp. nov." subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FB004192A354FB14" box="[1254,1319,1205,1231]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FAD14193A3D5FB15" box="[1335,1446,1204,1230]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FCCB41F3A5E1FB35" box="[813,914,1236,1262]" italics="true" pageId="21" pageNumber="1328">helladicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5108FFA36C73FC7C41F2A5A4FB34" box="[922,983,1237,1263]" pageId="21" pageNumber="1328" rank="species">n. sp.</taxonomicNameLabel>
, because of their flat spire whorls and their identical colour pattern.
</paragraph>
<paragraph id="8BA33061FFA36C73FCA54032A24FF8D6" blockId="21.[811,1457,950,1805]" pageId="21" pageNumber="1328">
<taxonomicName id="4C1C4BE2FFA36C73FCA54032A3DCFAF4" authority="Hoernes &amp; Auinger, 1879" authorityName="Hoernes &amp; Auinger" authorityYear="1879" box="[835,1455,1300,1327]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="hungaricus" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FCA54032A5F7FAF4" box="[835,900,1301,1327]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FC704033A276FAF5" box="[918,1029,1300,1326]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FBF14033A2F4FAF5" box="[1047,1159,1300,1326]" italics="true" pageId="21" pageNumber="1328">hungaricus</emphasis>
<bibRefCitation id="EF8D4D90FFA36C73FB764032A3DCFAF4" author="HOERNES R. &amp; AUINGER M." box="[1168,1455,1300,1327]" pageId="21" pageNumber="1328" pagination="1 - 52" refId="ref19161" refString="HOERNES R. &amp; AUINGER M. 1879. - Die Gasteropoden der Meeres- Ablagerungen der ersten und zweiten Miocanen Mediterran-Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt 12: 1 - 52." type="journal article" year="1879">Hoernes &amp; Auinger, 1879</bibRefCitation>
</taxonomicName>
from Paratethys seems closely related to
<taxonomicName id="4C1C4BE2FFA36C73FB004012A5E1FAB5" authority="Psarras &amp; Koskeridou &amp; Merle, 2021" authorityName="Psarras &amp; Koskeridou &amp; Merle" authorityYear="2021" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="helladicus" status="sp. nov." subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FB004012A354FA94" box="[1254,1319,1333,1359]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FAD14013A3D5FA95" box="[1335,1446,1332,1358]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FCCB4073A5E1FAB5" box="[813,914,1364,1390]" italics="true" pageId="21" pageNumber="1328">helladicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5108FFA36C73FC714072A5A1FAB4" box="[919,978,1365,1391]" pageId="21" pageNumber="1328" rank="species">n. sp.</taxonomicNameLabel>
, but the medium height, conical spire whorls and the subsutural flexure of
<taxonomicName id="4C1C4BE2FFA36C73FB884053A3DCFA55" authorityName="Hoernes &amp; Auinger" authorityYear="1879" box="[1134,1455,1396,1423]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="hungaricus" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FB884053A2DCFA55" box="[1134,1199,1396,1422]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FB594053A35DFA55" box="[1215,1326,1396,1422]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FAD94053A3DCFA55" box="[1343,1455,1396,1422]" italics="true" pageId="21" pageNumber="1328">hungaricus</emphasis>
</taxonomicName>
(medium depth, moderately curved, seeHarzhauser &amp; Landau 2016) are characters separating both species.
<taxonomicName id="4C1C4BE2FFA36C73FAEC4093A2C8FA35" authority="Hoernes &amp; Auinger, 1879" authorityName="Hoernes &amp; Auinger" authorityYear="1879" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="tietzei" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FAEC4093A338FA15" box="[1290,1355,1460,1486]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FABC4093A523FA35" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FCB940F2A5EEFA35" box="[863,925,1493,1518]" italics="true" pageId="21" pageNumber="1328">tietzei</emphasis>
<bibRefCitation id="EF8D4D90FFA36C73FC4240F3A2C8FA35" author="HOERNES R. &amp; AUINGER M." box="[932,1211,1492,1518]" pageId="21" pageNumber="1328" pagination="1 - 52" refId="ref19161" refString="HOERNES R. &amp; AUINGER M. 1879. - Die Gasteropoden der Meeres- Ablagerungen der ersten und zweiten Miocanen Mediterran-Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt 12: 1 - 52." type="journal article" year="1879">Hoernes &amp; Auinger, 1879</bibRefCitation>
</taxonomicName>
differs in the relatively angulated shoulder and the medium depth of the subsutural flexure (
<bibRefCitation id="EF8D4D90FFA36C73FC6F4334A2B5F9F6" author="HARZHAUSER M. &amp; LANDAU B." box="[905,1222,1555,1582]" pageId="21" pageNumber="1328" pagination="1" refId="ref18832" refString="HARZHAUSER M. &amp; LANDAU B. 2016. - A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 001 - 178. https: // doi. org / 10.11646 / zootaxa. 4210.1.1" type="journal article" year="2016">Harzhauser &amp; Landau 2016</bibRefCitation>
).
<taxonomicName id="4C1C4BE2FFA36C73FB064333A2F2F995" authority="Erunal-Erentoz, 1958" authorityName="Erunal-Erentoz" authorityYear="1958" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="gulemani" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FB064333A351F9F5" box="[1248,1314,1556,1582]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FAD24334A3D5F9F6" box="[1332,1446,1555,1581]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FCCB4314A5FEF996" box="[813,909,1587,1613]" italics="true" pageId="21" pageNumber="1328">gulemani</emphasis>
Erünal-Erentöz, 1958
</taxonomicName>
bears a similar morphology and a colour pattern. The differences between both species exist on the spiral whorl height and the smoother shoulder of
<taxonomicName id="4C1C4BE2FFA36C73FCAC43B4A209F976" authorityName="Erunal-Erentoz" authorityYear="1958" box="[842,1146,1683,1710]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="gulemani" subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FCAC43B4A5F8F976" box="[842,907,1683,1709]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FC7D43B4A279F976" box="[923,1034,1683,1709]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FBFC43B4A209F976" box="[1050,1146,1683,1709]" italics="true" pageId="21" pageNumber="1328">gulemani</emphasis>
</taxonomicName>
. All these species seem to be very closely related, but the differential characters of
<taxonomicName id="4C1C4BE2FFA36C73FA964394A26AF936" authority="Psarras &amp; Koskeridou &amp; Merle, 2021" authorityName="Psarras &amp; Koskeridou &amp; Merle" authorityYear="2021" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="1328" phylum="Mollusca" rank="species" species="helladicus" status="sp. nov." subGenus="Kalloconus">
<emphasis id="B968EC73FFA36C73FA964394A3C2F916" box="[1392,1457,1715,1741]" italics="true" pageId="21" pageNumber="1328">Conus</emphasis>
(
<emphasis id="B968EC73FFA36C73FCD043F4A5D6F936" box="[822,933,1747,1773]" italics="true" pageId="21" pageNumber="1328">Kalloconus</emphasis>
)
<emphasis id="B968EC73FFA36C73FC5243F4A26AF936" box="[948,1049,1747,1773]" italics="true" pageId="21" pageNumber="1328">helladicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5108FFA36C73FBF943F4A22AF936" box="[1055,1113,1747,1773]" pageId="21" pageNumber="1328" rank="species">n. sp.</taxonomicNameLabel>
caused us to consider the Greek material as a new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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