228 lines
21 KiB
XML
228 lines
21 KiB
XML
<document ID-DOI="10.5281/zenodo.272951" ID-GBIF-Dataset="e2c88f4c-3ac2-45f9-95e4-99b75561a081" ID-ISSN="1175-5326" ID-Zenodo-Dep="272951" ID-ZooBank="6D68A019-6F63-4AA4-A8B3-92D351F1F69B" checkinTime="1484409208030" checkinUser="plazi" docAuthor="Van, Rob W. M." docDate="2017" docId="03A80010776AFF92FF14A5789110F85E" docLanguage="en" docName="zootaxa.4217.1.1.pdf.imf" docOrigin="Zootaxa 1" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5ce3343f" docStyleName="zootaxa.2013.monograph" docTitle="Raspailia (Parasyringella) thamnopilosa Van, 2017, sp. nov." docType="treatment" docVersion="19" lastPageNumber="63" masterDocId="FF9178687757FFACFF83A36B907DFF9A" masterDocTitle="Sponges of the Guyana Shelf" masterLastPageNumber="225" masterPageNumber="1" pageNumber="62" updateTime="1645784257244" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Sponges of the Guyana Shelf</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Van, Rob W. M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>225</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.272951</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e2c88f4c-3ac2-45f9-95e4-99b75561a081</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">272951</mods:identifier>
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<mods:identifier type="ZooBank">6D68A019-6F63-4AA4-A8B3-92D351F1F69B</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5698612" ID-GBIF-Taxon="127419896" ID-Zenodo-Dep="5698612" LSID="urn:lsid:plazi:treatment:03A80010776AFF92FF14A5789110F85E" httpUri="http://treatment.plazi.org/id/03A80010776AFF92FF14A5789110F85E" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
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<subSubSection pageId="61" pageNumber="62" type="nomenclature">
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<paragraph blockId="61.[151,742,1555,1614]" box="[151,742,1555,1581]" pageId="61" pageNumber="62">
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<heading bold="true" box="[151,742,1555,1581]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
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<emphasis bold="true" box="[151,742,1555,1581]" pageId="61" pageNumber="62">
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<taxonomicName box="[151,642,1555,1581]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="61" pageNumber="62" phylum="Porifera" rank="species" species="thamnopilosa" status="sp. nov." subGenus="Parasyringella">
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<emphasis bold="true" box="[151,642,1555,1581]" italics="true" pageId="61" pageNumber="62">Raspailia (Parasyringella) thamnopilosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[648,742,1555,1581]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="61.[151,742,1555,1614]" box="[151,311,1589,1614]" pageId="61" pageNumber="62">
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<figureCitation box="[151,270,1589,1614]" captionStart="FIGURE 40" captionStartId="62.[151,250,1115,1137]" captionTargetBox="[151,1436,193,1093]" captionTargetId="figure@62.[151,1436,193,1093]" captionTargetPageId="62" captionText="FIGURE 40. Raspailia (Parasyringella) thamnopilosa sp. nov., a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b – d, SEM images of spicules, b, style, b 1, details of b, c, oxea, c 1, details of c, d, rhabdostyle, d 1, details of d." httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="61" pageNumber="62">Figures 40</figureCitation>
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a–d
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</paragraph>
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</subSubSection>
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<subSubSection pageId="61" pageNumber="62" type="materials_examined">
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<paragraph blockId="61.[151,1437,1661,2010]" pageId="61" pageNumber="62">
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<materialsCitation ID-GBIF-Occurrence="1424726426" collectingDate="1969-04-17" collectionCode="RMNH" collectorName="Luymes O. C. P. S." country="Suriname" elevation="93" latitude="7.3783" location="Suriname" longLatPrecision="7" longitude="-54.3583" pageId="61" pageNumber="62" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[151,385,1661,1686]" pageId="61" pageNumber="62">Material examined.</emphasis>
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<typeStatus box="[393,497,1661,1686]" pageId="61" pageNumber="62">Holotype</typeStatus>
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<collectionCode box="[505,589,1662,1685]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="61" pageNumber="62">RMNH</collectionCode>
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Por. 6301,
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<collectingCountry box="[720,830,1661,1686]" name="Suriname" pageId="61" pageNumber="62">Suriname</collectingCountry>
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, ‘
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<collectorName box="[846,1049,1662,1686]" pageId="61" pageNumber="62">Luymes O.C.P.S.</collectorName>
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’
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<collectingCountry box="[1062,1149,1662,1686]" name="Guyana" pageId="61" pageNumber="62">Guyana</collectingCountry>
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Shelf Expedition, station K101B,
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<geoCoordinate box="[246,354,1698,1722]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="5" value="7.3783">7.3783°N</geoCoordinate>
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<geoCoordinate box="[361,490,1697,1722]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="5" value="-54.3583">54.3583°W</geoCoordinate>
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, depth
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<quantity box="[570,630,1698,1722]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="61" pageNumber="62" unit="m" value="93.0">
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<elevation box="[570,630,1698,1722]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="61" pageNumber="62" unit="m" value="93.0">93 m</elevation>
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</quantity>
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, rectangular dredge,
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<date box="[864,1024,1697,1722]" pageId="61" pageNumber="62" value="1969-04-17">
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<collectingDate box="[864,1024,1697,1722]" pageId="61" pageNumber="62" value="1969-04-17">17 April 1969</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="61.[151,1437,1661,2010]" box="[199,784,1733,1758]" pageId="61" pageNumber="62">
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<materialsCitation ID-GBIF-Occurrence="1424726434" box="[199,784,1733,1758]" collectingDate="1969-04-17" collectionCode="RMNH" collectorName="Luymes O. C. P. S." country="Suriname" elevation="93" latitude="7.3783" location="Suriname" longLatPrecision="7" longitude="-54.3583" pageId="61" pageNumber="62" specimenCount="1" typeStatus="paratype">
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<typeStatus box="[199,297,1734,1758]" pageId="61" pageNumber="62">Paratype</typeStatus>
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<collectionCode box="[304,388,1734,1757]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="61" pageNumber="62">RMNH</collectionCode>
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Por. 10550, same data as holotype.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="description">
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<paragraph blockId="61.[151,1437,1661,2010]" pageId="61" pageNumber="62">
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<emphasis bold="true" box="[199,345,1769,1794]" pageId="61" pageNumber="62">Description.</emphasis>
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(
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<figureCitation box="[361,444,1769,1794]" captionStart="FIGURE 40" captionStartId="62.[151,250,1115,1137]" captionTargetBox="[151,1436,193,1093]" captionTargetId="figure@62.[151,1436,193,1093]" captionTargetPageId="62" captionText="FIGURE 40. Raspailia (Parasyringella) thamnopilosa sp. nov., a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b – d, SEM images of spicules, b, style, b 1, details of b, c, oxea, c 1, details of c, d, rhabdostyle, d 1, details of d." httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="61" pageNumber="62">Fig. 40</figureCitation>
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a) Small arborescent hairy sponges (two almost identical specimens), with short stubby branches partially anastomosed. Size about
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<quantity box="[659,718,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="62" unit="cm" value="2.0">2 cm</quantity>
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high,
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<quantity box="[796,876,1805,1830]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="61" pageNumber="62" unit="cm" value="1.5">1.5 cm</quantity>
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wide, individual branches
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<quantity box="[1193,1290,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="61" pageNumber="62" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
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in diameter. Color in alcohol light beige. Consistency firm, slightly compressible.
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</paragraph>
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<paragraph blockId="61.[151,1437,1661,2010]" pageId="61" pageNumber="62">
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<emphasis bold="true" box="[199,309,1877,1902]" pageId="61" pageNumber="62">Skeleton.</emphasis>
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Densely spiculous, with a central axis of styles, curved oxeas and rhabdostyles in a vague reticulation, without visible spongin. From the axis long styles verge toward the surface and protrude far beyond it causing hispidity and a hairy aspect.
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</paragraph>
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<paragraph blockId="61.[151,1437,1661,2010]" box="[199,771,1985,2010]" pageId="61" pageNumber="62">
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<emphasis bold="true" box="[199,304,1985,2010]" pageId="61" pageNumber="62">Spicules.</emphasis>
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(
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<figureCitation box="[319,404,1985,2010]" captionStart="FIGURE 40" captionStartId="62.[151,250,1115,1137]" captionTargetBox="[151,1436,193,1093]" captionTargetId="figure@62.[151,1436,193,1093]" captionTargetPageId="62" captionText="FIGURE 40. Raspailia (Parasyringella) thamnopilosa sp. nov., a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b – d, SEM images of spicules, b, style, b 1, details of b, c, oxea, c 1, details of c, d, rhabdostyle, d 1, details of d." httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="61" pageNumber="62">Figs 40</figureCitation>
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b–d) Styles, oxeas, rhabdostyles.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="62" pageNumber="63" targetBox="[151,1436,193,1093]" targetPageId="62">
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<paragraph blockId="62.[151,1436,1115,1168]" pageId="62" pageNumber="63">
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<emphasis bold="true" box="[151,289,1115,1137]" pageId="62" pageNumber="63">FIGURE 40.</emphasis>
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<taxonomicName box="[296,716,1115,1137]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="62" pageNumber="63" phylum="Porifera" rank="species" species="thamnopilosa" status="sp. nov." subGenus="Parasyringella">
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<emphasis box="[296,716,1115,1137]" italics="true" pageId="62" pageNumber="63">Raspailia (Parasyringella) thamnopilosa</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[723,805,1115,1136]" pageId="62" pageNumber="63">
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<taxonomicNameLabel box="[723,805,1115,1136]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b–d, SEM images of spicules, b, style, b1, details of b, c, oxea, c1, details of c, d, rhabdostyle, d1, details of d.
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</paragraph>
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</caption>
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<paragraph blockId="62.[151,1436,1207,1988]" pageId="62" pageNumber="63">
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Styles (
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<figureCitation box="[285,378,1207,1232]" captionStart="FIGURE 40" captionStartId="62.[151,250,1115,1137]" captionTargetBox="[151,1436,193,1093]" captionTargetId="figure@62.[151,1436,193,1093]" captionTargetPageId="62" captionText="FIGURE 40. Raspailia (Parasyringella) thamnopilosa sp. nov., a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b – d, SEM images of spicules, b, style, b 1, details of b, c, oxea, c 1, details of c, d, rhabdostyle, d 1, details of d." httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="62" pageNumber="63">Figs 40</figureCitation>
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b,b1) of the central column and surface, usually curved, occasionally more or less straight, slightly fusiform, with the rounded head marginally thinner than the middle part of the shaft, 612–
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<emphasis box="[1300,1342,1243,1267]" italics="true" pageId="62" pageNumber="63">923</emphasis>
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–
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<date pageId="62" pageNumber="63" value="1092-10-18">1092 x 18</date>
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–
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<emphasis box="[193,242,1279,1303]" italics="true" pageId="62" pageNumber="63">23.2</emphasis>
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–28 µm.
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</paragraph>
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<paragraph blockId="62.[151,1436,1207,1988]" pageId="62" pageNumber="63">
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Oxeas (
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<figureCitation box="[285,378,1315,1340]" captionStart="FIGURE 40" captionStartId="62.[151,250,1115,1137]" captionTargetBox="[151,1436,193,1093]" captionTargetId="figure@62.[151,1436,193,1093]" captionTargetPageId="62" captionText="FIGURE 40. Raspailia (Parasyringella) thamnopilosa sp. nov., a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b – d, SEM images of spicules, b, style, b 1, details of b, c, oxea, c 1, details of c, d, rhabdostyle, d 1, details of d." httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="62" pageNumber="63">Figs 40</figureCitation>
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c,c1), strongly curved, often a bit crooked with abrupt bends, but never sinuous, occasionally one of the apices is bluntly rounded, but the majority are sharply pointed, in a large size range, 217–
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<emphasis box="[1315,1357,1351,1376]" italics="true" pageId="62" pageNumber="63">336</emphasis>
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–464
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<date pageId="62" pageNumber="63">x 8</date>
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–
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<emphasis box="[179,224,1387,1411]" italics="true" pageId="62" pageNumber="63">11.3</emphasis>
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–14 µm.
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</paragraph>
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<paragraph blockId="62.[151,1436,1207,1988]" pageId="62" pageNumber="63">
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Rhabdostyles (
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<figureCitation box="[367,457,1423,1448]" captionStart="FIGURE 40" captionStartId="62.[151,250,1115,1137]" captionTargetBox="[151,1436,193,1093]" captionTargetId="figure@62.[151,1436,193,1093]" captionTargetPageId="62" captionText="FIGURE 40. Raspailia (Parasyringella) thamnopilosa sp. nov., a, habitus of holotype RMNH Por. 6301 (scale bar = 1 cm), b – d, SEM images of spicules, b, style, b 1, details of b, c, oxea, c 1, details of c, d, rhabdostyle, d 1, details of d." httpUri="https://zenodo.org/record/272991/files/figure.png" pageId="62" pageNumber="63">Figs 40</figureCitation>
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d,d1) smooth, short and fat, with abrupt curve just below the rounded head, in a large size range, 177–
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<emphasis box="[333,375,1459,1484]" italics="true" pageId="62" pageNumber="63">326</emphasis>
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–509
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<date box="[438,485,1459,1483]" pageId="62" pageNumber="63" value="2011-10">x 11</date>
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–
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<emphasis box="[500,549,1459,1483]" italics="true" pageId="62" pageNumber="63">18.2</emphasis>
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–26 µm.
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,1027,1495,1520]" pageId="62" pageNumber="63" type="distribution">
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<paragraph blockId="62.[151,1436,1207,1988]" box="[199,1027,1495,1520]" pageId="62" pageNumber="63">
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<emphasis bold="true" box="[199,498,1495,1520]" pageId="62" pageNumber="63">Distribution and ecology.</emphasis>
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<collectingCountry box="[505,592,1495,1519]" name="Guyana" pageId="62" pageNumber="63">Guyana</collectingCountry>
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Shelf, on sandy bottom at
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<quantity box="[893,950,1495,1519]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="62" pageNumber="63" unit="m" value="93.0">93 m</quantity>
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depth.
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,1234,1531,1556]" pageId="62" pageNumber="63" type="etymology">
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<paragraph blockId="62.[151,1436,1207,1988]" box="[199,1234,1531,1556]" pageId="62" pageNumber="63">
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<emphasis bold="true" box="[199,334,1531,1556]" pageId="62" pageNumber="63">Etymology.</emphasis>
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Thamnos (Gr.) = bush, pilosus (L.) = hairy, referring to the shape of the sponge.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="62" pageNumber="63" type="discussion">
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<paragraph blockId="62.[151,1436,1207,1988]" pageId="62" pageNumber="63">
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<emphasis bold="true" box="[199,315,1567,1592]" pageId="62" pageNumber="63">Remarks.</emphasis>
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None of the described
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<collectingRegion box="[586,669,1567,1592]" country="Sri Lanka" name="Central" pageId="62" pageNumber="63">Central</collectingRegion>
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<collectingRegion box="[677,732,1568,1591]" country="Tanzania" name="Mjini Magharibi" pageId="62" pageNumber="63">West</collectingRegion>
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Atlantic sponges appear to be similar to the above-described specimen. It is difficult to assign to any genus with certainty. On overall spiculation it could conform to
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<taxonomicName box="[1316,1409,1603,1628]" class="Demospongiae" family="Axinellidae" genus="Axinella" kingdom="Animalia" order="Halichondrida" pageId="62" pageNumber="63" phylum="Porifera" rank="genus">
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<emphasis box="[1316,1409,1603,1628]" italics="true" pageId="62" pageNumber="63">Axinella</emphasis>
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</taxonomicName>
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, a genus, which has been demonstrated on molecular grounds to be polyphyletic. However, the spined surface and the axial skeleton of strongly curved oxeas and rhabdostyles of the present specimen do not resemble that of any of the described
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||
<collectingRegion box="[265,356,1712,1735]" country="Sri Lanka" name="Western" pageId="62" pageNumber="63">Western</collectingRegion>
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Atlantic
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<taxonomicName box="[459,552,1711,1736]" class="Demospongiae" family="Axinellidae" genus="Axinella" kingdom="Animalia" order="Halichondrida" pageId="62" pageNumber="63" phylum="Porifera" rank="genus">
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<emphasis box="[459,552,1711,1736]" italics="true" pageId="62" pageNumber="63">Axinella</emphasis>
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</taxonomicName>
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species (cf.
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<bibRefCitation author="Alvarez" box="[691,908,1711,1736]" pageId="62" pageNumber="63" refString="Alvarez, B., van Soest, R. W. M. & Rutzler, K. (1998) A revision of the Axinellidae (Porifera: Demospongiae) of the Central- West Atlantic region. Smithsonian Contributions to Zoology, 598, 1 - 47. http: // dx. doi. org / 10.5479 / si. 00810282.598" type="journal article" year="1998">
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Alvarez
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<emphasis box="[786,842,1711,1736]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
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1998
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</bibRefCitation>
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). The spined surface caused by protruding long styles reminds of
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<taxonomicName box="[352,460,1747,1772]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="62" pageNumber="63" phylum="Porifera" rank="genus">
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<emphasis box="[352,460,1747,1772]" italics="true" pageId="62" pageNumber="63">Raspailia</emphasis>
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</taxonomicName>
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, but its lack of acanthostyles and lack of typical raspailiid bouquets of thinner shorter spicules (either styles or oxeas) makes membership of that genus less likely. The subgenus
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<emphasis italics="true" pageId="62" pageNumber="63">
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<taxonomicName box="[1329,1436,1783,1808]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="62" pageNumber="63" phylum="Porifera" rank="genus">Raspailia</taxonomicName>
|
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(
|
||
<taxonomicName box="[159,329,1819,1844]" class="Demospongiae" family="Raspailiidae" genus="Parasyringella" higherTaxonomySource="CoL" kingdom="Animalia" order="Axinellida" pageId="62" pageNumber="63" phylum="Porifera" rank="genus">Parasyringella</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
unites
|
||
<taxonomicName box="[428,535,1819,1844]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="62" pageNumber="63" phylum="Porifera" rank="genus">
|
||
<emphasis box="[428,535,1819,1844]" italics="true" pageId="62" pageNumber="63">Raspailia</emphasis>
|
||
</taxonomicName>
|
||
species lacking acanthostyles, and with some hesitation I assign the present specimens to that group.
|
||
</paragraph>
|
||
<paragraph blockId="62.[151,1436,1207,1988]" pageId="62" pageNumber="63">
|
||
<taxonomicName authority="Lerner et al., 2006" authorityName="Lerner et al." authorityYear="2006" box="[199,846,1891,1916]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="62" pageNumber="63" phylum="Porifera" rank="species" species="bouryesnaultae" subGenus="Raspaxilla">
|
||
<emphasis box="[199,632,1891,1916]" italics="true" pageId="62" pageNumber="63">Raspailia (Raspaxilla) bouryesnaultae</emphasis>
|
||
Lerner
|
||
<emphasis box="[720,776,1891,1916]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
|
||
, 2006
|
||
</taxonomicName>
|
||
(replacement name for
|
||
<taxonomicName box="[1114,1281,1891,1916]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" kingdom="Animalia" order="Poecilosclerida" pageId="62" pageNumber="63" phylum="Porifera" rank="species" species="elegans" subGenus="Raspaxilla">
|
||
<emphasis box="[1114,1281,1891,1916]" italics="true" pageId="62" pageNumber="63">R. (R.) elegans</emphasis>
|
||
</taxonomicName>
|
||
sensu Boury-
|
||
<bibRefCitation author="Esnault" box="[151,306,1927,1952]" pageId="62" pageNumber="63" refString="Boury-Esnault, N. (1973) Resultats scientifiques des campagnes de la " Calypso "." type="book" year="1973">Esnault, 1973</bibRefCitation>
|
||
) from NE
|
||
<collectingCountry box="[428,496,1927,1952]" name="Brazil" pageId="62" pageNumber="63">Brazil</collectingCountry>
|
||
also has rhabdostyles, but these are spined, unlike the entirely smooth rhabdostyles of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |