treatments-xml/data/03/A4/6A/03A46A7DFFED0C7AFC658B3CFCC1F998.xml

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<mods:title id="7F2C7EEB08FB1A8C85E766A660D5A1FE">Review of the subgenus Attumbrinus (Coleoptera: Leiodidae: Cholevinae: Philomessor), with description of Philomessor lackneri sp. nov. from Morocco</mods:title>
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<taxonomicName id="4C0DA0E8FFED0C71FC658B3CFD15FCFC" ID-CoL="4G9BD" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[356,753,826,850]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="88" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
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(
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) 
<emphasis id="B9790779FFED0C71FF968B3CFD15FCFC" bold="true" box="[663,753,826,850]" italics="true" pageId="3" pageNumber="88">lackneri</emphasis>
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<subSubSection id="C31788E0FFED0C71FC538B5FFC84FCC2" box="[338,864,857,876]" pageId="3" pageNumber="88" type="description">
<paragraph id="8BB2DB6BFFED0C71FC538B5FFC84FCC2" blockId="3.[338,864,857,876]" box="[338,864,857,876]" pageId="3" pageNumber="88">
(
<figureCitation id="1336C7EEFFED0C71FC598B5FFE5EFCC2" box="[344,442,857,876]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="3" pageNumber="88">Figs 1, 2, 6</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FCC28B5FFDFEFCC2" box="[451,538,857,876]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="3" pageNumber="88">14–17, 20</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FF228B5FFDBEFCC2" box="[547,602,857,876]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="3" pageNumber="88">26, 27</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FF628B5FFD7EFCC2" box="[611,666,857,876]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="3" pageNumber="88">33, 34</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FFA28B5FFD3EFCC2" box="[675,730,857,876]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="3" pageNumber="88">41–45</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FFE28B5FFD1EFCC2" box="[739,762,857,876]" captionStart="Figs 49–54. 49, 51, 53" captionStartId="14.[151,188,1161,1180]" captionTargetBox="[151,1126,219,1141]" captionTargetId="figure-10@14.[151,1126,219,1141]" captionTargetPageId="14" captionText="Figs 49–54. 49, 51, 53 – metatibia in lateral view, males; 50 – antenna, male; 52 – antenna, female; 54 – pronotum, male.49 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden,Morocco); 50–52 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 53–54 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372956" httpUri="https://zenodo.org/record/5372956/files/figure.png" pageId="3" pageNumber="88">49</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FE028B5FFCDEFCC2" box="[771,826,857,876]" captionStart="Figs 55–58. 55" captionStartId="16.[151,188,904,923]" captionTargetBox="[151,1126,219,879]" captionTargetId="figure-10@16.[151,1126,219,880]" captionTargetPageId="16" captionText="Figs 55–58. 55 – habitat of Philomessor (Attumbrinus) lackneri sp. nov. and P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco, 22.v.2009); 56 – habitat of P. (A.) bolivari (Mischlifen, Morocco, 27.v.2011); 57 – nest of Messor abdelazizi Santschi, 1921, under a stone on a pasture (Oukaïmeden, Morocco, 20.v.2011); 58 – male of P. (A.) bolivari in the nest of Messor abdelazizi (Oukaïmeden, Morocco, 20.v.2011). Photographs by P. Koniar and M. Švarc." figureDoi="http://doi.org/10.5281/zenodo.5372958" httpUri="https://zenodo.org/record/5372958/files/figure.png" pageId="3" pageNumber="88">55, 57</figureCitation>
, 
<figureCitation id="1336C7EEFFED0C71FE428B5FFCBDFCC2" box="[835,857,857,876]" captionStart="Fig" captionStartId="16.[151,182,1564,1583]" captionTargetBox="[152,1122,1132,1537]" captionTargetId="figure-95@16.[151,1127,1132,1540]" captionTargetPageId="16" captionText="Fig. 59. Distribution of Philomessor (Attumbrinus) Jeannel, 1936 in the northwest of Africa." figureDoi="http://doi.org/10.5281/zenodo.5372960" httpUri="https://zenodo.org/record/5372960/files/figure.png" pageId="3" pageNumber="88">59</figureCitation>
)
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locality.
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<collectingCountry id="F31A9BFBFFED0C71FDEE8B85FEDAFC38" box="[239,318,899,918]" name="Morocco" pageId="3" pageNumber="88">Morocco</collectingCountry>
, High Atlas Mts., Oukaïmeden, 
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a.s.l., 
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, 
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.
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<emphasis id="B9790779FFED0C71FD708B98FEECFC1F" bold="true" box="[113,264,926,945]" pageId="3" pageNumber="88">Type material.</emphasis>
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: J (
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), ‘ 
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/ OUKAÏMEDEN ENV. (
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N 
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</emphasis>
′ 
<emphasis id="B9790779FFED0C71FDDA8BBEFEA7FC65" box="[219,323,952,971]" italics="true" pageId="3" pageNumber="88">W07º49,264</emphasis>
′ / 
<collectorName id="26F8BEBDFFED0C71FC5D8BBEFE2EFC65" box="[348,458,952,971]" pageId="3" pageNumber="88">Peter Koniar</collectorName>
&amp; 
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lgt. [white label, printed]
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’. 
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: 
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[MD] (
<collectionCode id="ED1C43AEFFED0C71F9018BBEFBD3FC65" box="[1024,1079,952,971]" pageId="3" pageNumber="88">JVAC</collectionCode>
), ‘ 
<collectingCountry id="F31A9BFBFFED0C71FD758BD5FF00FC48" box="[116,228,979,998]" name="Morocco" pageId="3" pageNumber="88">MOROCCO</collectingCountry>
<date id="FFB3FDABFFED0C71FDE98BD5FEDAFC48" box="[232,318,979,998]" pageId="3" pageNumber="88" value="2011-05-20">
<collectingDate id="EFF70443FFED0C71FDE98BD5FEDAFC48" box="[232,318,979,998]" pageId="3" pageNumber="88" value="2011-05-20">20.V.2011</collectingDate>
</date>
/ OUKAÏMEDEN ENV.(
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N 
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′ 
<emphasis id="B9790779FFED0C71FFD58BD5FCD8FC48" box="[724,828,979,998]" italics="true" pageId="3" pageNumber="88">W07º49,055</emphasis>
′ / 
<collectorName id="26F8BEBDFFED0C71FE4E8BD5FC5DFC48" box="[847,953,979,998]" pageId="3" pageNumber="88">Peter Koniar</collectorName>
&amp; 
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lgt. [white label, printed]
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’. 
<materialsCitation id="3B65D136FFED0C71FC598BE8FE6BFBB5" collectingDate="1968-03-19" collectionCode="MNHN" collectorName="Ito &amp; Azrou" country="Morocco" location="MAROC" pageId="3" pageNumber="88" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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: 
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(
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), ‘ 
<collectingCountry id="F31A9BFBFFED0C71FF3D8BE8FD6AFBAF" box="[572,654,1006,1025]" name="Morocco" pageId="3" pageNumber="88">MAROC</collectingCountry>
19 3. 6 8 [
<date id="FFB3FDABFFED0C71FFE88BE8FCADFBAF" box="[745,841,1006,1025]" pageId="3" pageNumber="88" value="1968-03-19">
<collectingDate id="EFF70443FFED0C71FFE88BE8FCADFBAF" box="[745,841,1006,1025]" pageId="3" pageNumber="88" value="1968-03-19">19.iii.1968</collectingDate>
</date>
] / I T O [
<collectorName id="26F8BEBDFFED0C71FE968BE8FC55FBAF" box="[919,945,1006,1025]" pageId="3" pageNumber="88">Ito</collectorName>
near 
<collectorName id="26F8BEBDFFED0C71FEDE8BE8FBF3FBAF" box="[991,1047,1006,1025]" pageId="3" pageNumber="88">Azrou</collectorName>
] / H. COIFFAIT [white label, printed]
</materialsCitation>
’; 
<materialsCitation id="3B65D136FFED0C71FC9A8C0EFC58FB98" collectingDate="1998-06-21" collectionCode="TLAC, JVAC" collectorName="T. Lackner" country="Morocco" elevation="1900" latitude="33.45" location="Moyen Atlas" longLatPrecision="1205" longitude="-5.0333333" municipality="Ifrane" pageId="3" pageNumber="88" specimenCount="1" typeStatus="paratype">
2 JJ (
<collectionCode id="ED1C43AEFFED0C71FCD78C0EFDF5FBB5" box="[470,529,1032,1051]" pageId="3" pageNumber="88">TLAC</collectionCode>
, 
<collectionCode id="ED1C43AEFFED0C71FF1B8C0EFDB4FBB5" box="[538,592,1032,1051]" pageId="3" pageNumber="88">JVAC</collectionCode>
), ‘ 
<collectingCountry id="F31A9BFBFFED0C71FF638C0EFD7AFBB5" box="[610,670,1032,1051]" name="Morocco" pageId="3" pageNumber="88">Maroc</collectingCountry>
centr. / 
<location id="8ED28DB0FFED0C71FFE28C0EFCB4FBB5" LSID="urn:lsid:plazi:treatment:03A46A7DFFED0C7AFC658B3CFCC1F998:8ED28DB0FFED0C71FFE28C0EFCB4FBB5" box="[739,848,1032,1051]" country="Morocco" latitude="33.45" longLatPrecision="1205" longitude="-5.0333333" municipality="Ifrane" name="Moyen Atlas" pageId="3" pageNumber="88">Moyen Atlas</location>
/ 
<collectingMunicipality id="6BD64111FFED0C71FE608C0EFC70FBB5" box="[865,916,1032,1051]" pageId="3" pageNumber="88">Ifrane</collectingMunicipality>
env. [Tizi-n-Tretten near Ifrane, 
<geoCoordinate id="EE39BDACFFED0C71FDDA8C25FEC0FB98" box="[219,292,1059,1078]" degrees="33" direction="north" minutes="27" orientation="latitude" pageId="3" pageNumber="88" precision="925" value="33.45">33°27′N</geoCoordinate>
, 
<geoCoordinate id="EE39BDACFFED0C71FC2F8C25FE8BFB98" box="[302,367,1059,1078]" degrees="5" direction="west" minutes="02" orientation="longitude" pageId="3" pageNumber="88" precision="925" value="-5.0333333">5°02′W</geoCoordinate>
, 
<elevation id="00203C58FFED0C71FC788C25FE0EFB98" box="[377,490,1059,1078]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="88" unit="m" value="1900.0">
<quantity id="4CF5768EFFED0C71FC788C25FE5FFB98" box="[377,443,1059,1078]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="88" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
] 17- 
<date id="FFB3FDABFFED0C71FF0E8C25FD8CFB98" box="[527,616,1059,1078]" pageId="3" pageNumber="88" value="1998-06-21">
<collectingDate id="EFF70443FFED0C71FF0E8C25FD8CFB98" box="[527,616,1059,1078]" pageId="3" pageNumber="88" value="1998-06-21">21.6.1998</collectingDate>
</date>
/ 
<collectorName id="26F8BEBDFFED0C71FF7A8C25FD3CFB98" box="[635,728,1059,1078]" pageId="3" pageNumber="88">T. Lackner</collectorName>
lgt. [white label, printed]
</materialsCitation>
’; 
<materialsCitation id="3B65D136FFED0C71FEC88C25FEAEFBC5" collectingDate="2009-05-21" collectionCode="NMPC" collectorName="Peter Koniar &amp; Martin Svarc" country="Morocco" elevation="2245" location="MOROCCO" longitude="-31.237617" pageId="3" pageNumber="88" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D0B10E2FFED0C71FEC88C25FC08FB98" box="[969,1004,1053,1078]" pageId="3" pageNumber="88" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1C43AEFFED0C71FEF78C25FBD3FB98" box="[1014,1079,1059,1078]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="88">NMPC</collectionCode>
), ‘ 
<collectingCountry id="F31A9BFBFFED0C71FD758C38FF00FBFF" box="[116,228,1086,1105]" name="Morocco" pageId="3" pageNumber="88">MOROCCO</collectingCountry>
<date id="FFB3FDABFFED0C71FDE98C38FEDBFBFF" box="[232,319,1086,1105]" pageId="3" pageNumber="88" value="2009-05-21">
<collectingDate id="EFF70443FFED0C71FDE98C38FEDBFBFF" box="[232,319,1086,1105]" pageId="3" pageNumber="88" value="2009-05-21">21.V.2009</collectingDate>
</date>
/ OUKAÏMEDEN ENV. (
<quantity id="4CF5768EFFED0C71FF1B8C38FDBCFBFF" box="[538,600,1086,1105]" metricMagnitude="3" metricUnit="m" metricValue="2.245" pageId="3" pageNumber="88" unit="m" value="2245.0">
<elevation id="00203C58FFED0C71FF1B8C38FDBCFBFF" box="[538,600,1086,1105]" metricMagnitude="3" metricUnit="m" metricValue="2.245" pageId="3" pageNumber="88" unit="m" value="2245.0">2245m</elevation>
</quantity>
) / 
<emphasis id="B9790779FFED0C71FF6A8C38FD34FBFF" box="[619,720,1086,1105]" italics="true" pageId="3" pageNumber="88">
N 
<geoCoordinate id="EE39BDACFFED0C71FF7A8C38FD34FBFF" box="[635,720,1086,1105]" degrees="31" direction="west" minutes="14,257" orientation="longitude" pageId="3" pageNumber="88" precision="1" value="-31.237617">31º14,257</geoCoordinate>
</emphasis>
′ 
<emphasis id="B9790779FFED0C71FFD58C38FCD8FBFF" box="[724,828,1086,1105]" italics="true" pageId="3" pageNumber="88">W07º49,066</emphasis>
′ / 
<collectorName id="26F8BEBDFFED0C71FE4E8C38FC5EFBFF" box="[847,954,1086,1105]" pageId="3" pageNumber="88">Peter Koniar</collectorName>
&amp; 
<collectorName id="26F8BEBDFFED0C71FED38C38FBA4FBFF" box="[978,1088,1086,1105]" pageId="3" pageNumber="88">Martin Švarc</collectorName>
lgt. [white label, printed]
</materialsCitation>
’; 
<materialsCitation id="3B65D136FFED0C71FC578C5EFC43FB28" collectingDate="2010-05-14" collectionCode="JRUC" collectorName="Di Giulio &amp; Maurizi &amp; Bravi" country="Morocco" county="Oukaimeden" elevation="870" location="Toubkal National" municipality="Park" pageId="3" pageNumber="88" specimenCount="1" typeStatus="paratype">
1 J (
<collectionCode id="ED1C43AEFFED0C71FC7E8C5EFE53FBC5" box="[383,439,1112,1131]" pageId="3" pageNumber="88">JRUC</collectionCode>
), ‘ 
<collectingCountry id="F31A9BFBFFED0C71FCC88C5EFDDFFBC5" box="[457,571,1112,1131]" name="Morocco" pageId="3" pageNumber="88">MOROCCO</collectingCountry>
, 
<location id="8ED28DB0FFED0C71FF438C5EFD32FBC5" LSID="urn:lsid:plazi:treatment:03A46A7DFFED0C7AFC658B3CFCC1F998:8ED28DB0FFED0C71FF438C5EFD32FBC5" box="[578,726,1112,1131]" country="Morocco" county="Oukaimeden" municipality="Park" name="Toubkal National" pageId="3" pageNumber="88">Toubkal National</location>
/ 
<collectingMunicipality id="6BD64111FFED0C71FFE48C5EFCEAFBC5" box="[741,782,1112,1131]" pageId="3" pageNumber="88">Park</collectingMunicipality>
, 
<collectingCounty id="62D3A3E7FFED0C71FE178C5EFC61FBC5" box="[790,901,1112,1131]" pageId="3" pageNumber="88">Oukaimeden</collectingCounty>
, on road / 2028, 1 
<elevation id="00203C58FFED0C71F9208C5EFF54FB28" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="88" unit="m" value="870.0">
<quantity id="4CF5768EFFED0C71F9208C5EFF66FB28" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="88" unit="m" value="870.0">870 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFB3FDABFFED0C71FDB88C75FEF4FB28" box="[185,272,1139,1158]" pageId="3" pageNumber="88" value="2010-05-14">
<collectingDate id="EFF70443FFED0C71FDB88C75FEF4FB28" box="[185,272,1139,1158]" pageId="3" pageNumber="88" value="2010-05-14">14.V.2010</collectingDate>
</date>
/ 0613390E- 3457303N / 
<collectorName id="26F8BEBDFFED0C71FCEC8C75FDA4FB28" box="[493,576,1139,1158]" pageId="3" pageNumber="88">Di Giulio</collectorName>
, 
<collectorName id="26F8BEBDFFED0C71FF498C75FD6AFB28" box="[584,654,1139,1158]" pageId="3" pageNumber="88">Maurizi</collectorName>
, 
<collectorName id="26F8BEBDFFED0C71FF978C75FD21FB28" box="[662,709,1139,1158]" pageId="3" pageNumber="88">Bravi</collectorName>
leg. [white label, printed]
</materialsCitation>
’; 
<materialsCitation id="3B65D136FFED0C71FEB28C75FCFEFB0F" collectingDate="2011-05-20" collectionCode="PKOC, MSVC, JVAC" collectorName="Peter Koniar &amp; Martin Svarc" country="Morocco" elevation="2237" location="MOROCCO" longitude="-31.237267" pageId="3" pageNumber="88" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
4 JJ 
<specimenCount id="9D0B10E2FFED0C71FEE68C75FBF1FB28" box="[999,1045,1133,1158]" pageId="3" pageNumber="88" type="female">2 ♀♀</specimenCount>
(1 J 
<specimenCount id="9D0B10E2FFED0C71FD708C88FF75FB0E" box="[113,145,1159,1185]" pageId="3" pageNumber="88" type="female">1 ♀</specimenCount>
in 
<collectionCode id="ED1C43AEFFED0C71FDAD8C88FF0CFB0F" box="[172,232,1166,1185]" pageId="3" pageNumber="88">PKOC</collectionCode>
; 2 JJ, 
<specimenCount id="9D0B10E2FFED0C71FC2B8C88FEAEFB0E" box="[298,330,1159,1185]" pageId="3" pageNumber="88" type="female">1 ♀</specimenCount>
[MD] in 
<collectionCode id="ED1C43AEFFED0C71FC9A8C88FE3EFB0F" box="[411,474,1166,1185]" pageId="3" pageNumber="88">MSVC</collectionCode>
; 1 J in 
<collectionCode id="ED1C43AEFFED0C71FF1F8C88FDB0FB0F" box="[542,596,1166,1185]" pageId="3" pageNumber="88">JVAC</collectionCode>
), same data as allotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFED0C74FD708CBFFC3AFB3A" lastPageId="6" lastPageNumber="91" pageId="3" pageNumber="88" type="description">
<paragraph id="8BB2DB6BFFED0C71FD708CBFFE04FA3E" blockId="3.[113,1090,1208,1584]" pageId="3" pageNumber="88">
<emphasis id="B9790779FFED0C71FD708CBFFEA0FB7E" bold="true" box="[113,324,1208,1232]" pageId="3" pageNumber="88">
Description. 
<emphasis id="B9790779FFED0C71FC048CBEFEA0FB7E" bold="true" box="[261,324,1208,1232]" italics="true" pageId="3" pageNumber="88">Male.</emphasis>
</emphasis>
Body length 3.0–3.4 mm (
<typeStatus id="54B665C9FFED0C71FF6C8CBFFD28FB7E" box="[621,716,1209,1232]" pageId="3" pageNumber="88" type="holotype">holotype</typeStatus>
3.0 mm). Body slender (
<figureCitation id="1336C7EEFFED0C71FEE38CBFFBD0FB7E" box="[994,1076,1209,1232]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="3" pageNumber="88">Fig. 26</figureCitation>
). Winged. Head dark brown to black brown, pronotum dark brown. Elytra, legs and maxillary palpi light brown. Antennae brown, two basal antennomeres and the tip of the terminal one lighter. Pubescence light and of two kinds: entire body covered with short, dense and recumbent setae, somewhat longer on head and pronotum than on elytra; on elytra the short setae intermixed with long erect setae, which are, especially on disc, more than three times longer than the recumbent ones (
<figureCitation id="1336C7EEFFED0C71FC858D7FFE30FA3E" box="[388,468,1401,1424]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="3" pageNumber="88">Fig. 34</figureCitation>
).
</paragraph>
<paragraph id="8BB2DB6BFFED0C71FD918D9FFED4FA5E" blockId="3.[113,1090,1208,1584]" pageId="3" pageNumber="88">Head transverse, 1.2 times wider than long. Surface shiny, finely punctate, punctures separated by more than their diameter. Interspaces with very fine microsculpture. Eyes developed, feebly prominent.</paragraph>
<paragraph id="8BB2DB6BFFED0C71FD918DFFFD51F99E" blockId="3.[113,1090,1208,1584]" pageId="3" pageNumber="88">
Antennae (
<figureCitation id="1336C7EEFFED0C71FC058DFFFEB5F9BE" box="[260,337,1529,1552]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="3" pageNumber="88">Fig. 14</figureCitation>
) long and slender, 2.4–2.5 (
<typeStatus id="54B665C9FFED0C71FF7A8DFFFD3EF9BE" box="[635,730,1529,1552]" pageId="3" pageNumber="88" type="holotype">holotype</typeStatus>
2.5) times longer than pronotum. Three terminal antennomeres dorsoventrally flattened.
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB488DAFBF0FEDD" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
Pronotum transverse, 2.4–2.5 (
<typeStatus id="54B665C9FFEA0C76FCFA88DAFDBEFF5D" box="[507,602,220,243]" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
2.5) times wider than long and 1.6–1.7 (
<typeStatus id="54B665C9FFEA0C76F90688DAFB82FF5D" box="[1031,1126,220,243]" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
1.6) times wider than head. Disc with pair of shallow lateral impressions at its midlength, furter paired impressions sometimes situated closer to median axis and base (also in 
<typeStatus id="54B665C9FFEA0C76FEFA891AFBB8FE9D" box="[1019,1116,284,307]" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
). Punctation of pronotum granular, very fine and dense, punctures separated by less than one diameter, at places contiguous. Microsculpture of interspaces very fine, but distinct.
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB4897AFC7CFE3D" blockId="4.[151,1127,220,1587]" box="[181,920,380,403]" pageId="4" pageNumber="89">Scutellum distinct, triangular, in middle finely and densely granulate.</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB4899AFE98FD3D" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
Elytra oblong, feebly convex, 1.6–1.7 (
<typeStatus id="54B665C9FFEA0C76FF53899AFD55FE1D" box="[594,689,412,435]" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
1.6) times longer than wide, 2.8–2.9 (
<typeStatus id="54B665C9FFEA0C76F943899AFF3EFE7D" pageId="4" pageNumber="89" type="holotype">holotype</typeStatus>
2.9) times longer than pronotum, their maximum combined width equal to or slightly wider than the width of pronotum. 
<collectingRegion id="49C91589FFEA0C76FF1C89DAFDAFFE5D" box="[541,587,476,499]" country="Algeria" name="Tipaza" pageId="4" pageNumber="89">Tips</collectingRegion>
of elytra separately rounded, almost rectangular. Punctation of elytra (
<figureCitation id="1336C7EEFFEA0C76FC7B89FAFE2DFDBD" box="[378,457,508,531]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="4" pageNumber="89">Fig. 33</figureCitation>
) granular and of two kinds. Fine granular punctures occurring all over surface of elytra, somewhat coarser and sparser than on pronotum and separated by more than one diameter, are intermixed with distinctly raised and larger granules bearing long erect setae. Microsculpture between punctures absent and elytra therefore appear more shiny than pronotum.
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB48A9AFE9BFCDD" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
Three basal protarsomeres (
<figureCitation id="1336C7EEFFEA0C76FCDF8A9AFDC8FD1D" box="[478,556,668,691]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="4" pageNumber="89">Fig. 41</figureCitation>
) dilated, becoming gradually narrower distad. Apical margins of tarsomeres simply emarginate. First tarsomere wider than apical margin of protibia and almost twice as wide as the second. Protibia (
<figureCitation id="1336C7EEFFEA0C76FFD18ADAFCFAFD5D" box="[720,798,732,755]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="4" pageNumber="89">Fig. 42</figureCitation>
) feebly outcurved. Mesotarsus long and narrow, as long as mesotibia. Mesotibia (
<figureCitation id="1336C7EEFFEA0C76FFBA8AFAFCEEFCBD" box="[699,778,764,787]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="4" pageNumber="89">Fig. 44</figureCitation>
) feebly curved, its inner margin except for basal portion with fine, dense and erect setae. Metatarsus long and narrow, as long as metatibia. Metatibia (
<figureCitation id="1336C7EEFFEA0C76FC9F8B3AFE09FCFD" box="[414,493,828,851]" captionStart="Figs 49–54. 49, 51, 53" captionStartId="14.[151,188,1161,1180]" captionTargetBox="[151,1126,219,1141]" captionTargetId="figure-10@14.[151,1126,219,1141]" captionTargetPageId="14" captionText="Figs 49–54. 49, 51, 53 – metatibia in lateral view, males; 50 – antenna, male; 52 – antenna, female; 54 – pronotum, male.49 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden,Morocco); 50–52 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 53–54 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372956" httpUri="https://zenodo.org/record/5372956/files/figure.png" pageId="4" pageNumber="89">Fig. 49</figureCitation>
) straight, its inner margin only in apical portion with fine, dense and erect setae.
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB48B7AFE89FBFD" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
Abdominal terga without impressions. Outer margin of sternum VIII broadly emarginate in middle (
<figureCitation id="1336C7EEFFEA0C76FC128B9AFEBDFC1D" box="[275,345,924,947]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="4" pageNumber="89">Fig. 6</figureCitation>
). Genital segment oval, anterior part wide, extremely elongated, trifid anterolaterally and strongly sclerotized (
<figureCitation id="1336C7EEFFEA0C76FF4A8BBAFD59FC7D" box="[587,701,956,979]" captionStart="Figs 9–12" captionStartId="6.[151,188,842,861]" captionTargetBox="[171,1113,231,805]" captionTargetId="figure-10@6.[151,1126,216,819]" captionTargetPageId="6" captionText="Figs 9–12. Genital segments in ventral view, males. 9, 10 – Philomessor (Attumbrinus) lackneri sp. nov. (9 – PT, Tizi-n-Tretten near Ifrane, Morocco; 10 – HT); 11 – P. (A.) bolivari Jeannel, 1936 (HT); 12 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372942" httpUri="https://zenodo.org/record/5372942/files/figure.png" pageId="4" pageNumber="89">Figs 9, 10</figureCitation>
). Aedeagus in dorsal view narrowed in basal portion, widest in middle and in apical portion tapering into long acute point (
<figureCitation id="1336C7EEFFEA0C76FEFE8BDAFBD8FC5D" box="[1023,1084,988,1011]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="4" pageNumber="89">Fig. 1</figureCitation>
), in lateral view regularly arcuate with feebly S-shaped apical portion (
<figureCitation id="1336C7EEFFEA0C76FE508BFAFC74FBBD" box="[849,912,1020,1043]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="4" pageNumber="89">Fig. 2</figureCitation>
), parameres narrow, equally broad all along their length with two long setae at apex, distinctly not reaching tip of aedeagus (
<figureCitation id="1336C7EEFFEA0C76FC248C3AFE84FBFD" box="[293,352,1084,1107]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="4" pageNumber="89">Fig.1</figureCitation>
).
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB48C5DFB83FABD" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
<emphasis id="B9790779FFEA0C76FDB48C5DFEEBFBDD" bold="true" box="[181,271,1115,1139]" pageId="4" pageNumber="89">
<emphasis id="B9790779FFEA0C76FDB48C5DFEECFBDD" bold="true" box="[181,264,1115,1139]" italics="true" pageId="4" pageNumber="89">Female</emphasis>
.
</emphasis>
Body length 2.9–3.3 mm (
<typeStatus id="54B665C9FFEA0C76FF288C5AFD65FBDD" box="[553,641,1116,1139]" pageId="4" pageNumber="89" type="allotype">allotype</typeStatus>
3.0 mm). Body slender (
<figureCitation id="1336C7EEFFEA0C76FE878C5AFC31FBDD" box="[902,981,1116,1139]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="4" pageNumber="89">Fig. 27</figureCitation>
). Colouration as in males (
<typeStatus id="54B665C9FFEA0C76FC1A8C7AFE97FB3D" box="[283,371,1148,1171]" pageId="4" pageNumber="89" type="allotype">allotype</typeStatus>
and 
<specimenCount id="9D0B10E2FFEA0C76FCA78C7AFDC0FB3D" box="[422,548,1148,1171]" pageId="4" pageNumber="89" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
), or head and pronotum black brown and elytra brown (
<specimenCount id="9D0B10E2FFEA0C76FDA38C9AFEF7FB1D" box="[162,275,1180,1203]" pageId="4" pageNumber="89" type="generic" typeStatus="paratype">1 paratype</specimenCount>
).Antennae (
<figureCitation id="1336C7EEFFEA0C76FC958C9AFE04FB1D" box="[404,480,1180,1203]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="4" pageNumber="89">Fig. 15</figureCitation>
) shorter than in male, 2.4–2.5 (
<typeStatus id="54B665C9FFEA0C76FE248C9AFC99FB1D" box="[805,893,1180,1203]" pageId="4" pageNumber="89" type="allotype">allotype</typeStatus>
2.5) times longer than pronotum. Pronotum transverse, 1.8–1.9 (
<typeStatus id="54B665C9FFEA0C76FF518CBAFD4CFB7D" box="[592,680,1212,1235]" pageId="4" pageNumber="89" type="allotype">allotype</typeStatus>
1.9) times wider than long. Elytra 1.6–1.7 (
<typeStatus id="54B665C9FFEA0C76FD9F8CDAFF12FB5D" box="[158,246,1244,1267]" pageId="4" pageNumber="89" type="allotype">allotype</typeStatus>
1.6) times longer than their combined width, 3.0–3.1 (
<typeStatus id="54B665C9FFEA0C76FE4C8CDAFC41FB5D" box="[845,933,1244,1267]" pageId="4" pageNumber="89" type="allotype">allotype</typeStatus>
3.1) times longer than pronotum and their maximum combined width equal to or slightly wider than pronotum.
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB48D1AFECFFA3D" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
Three basal protarsomeres (
<figureCitation id="1336C7EEFFEA0C76FCD58D1AFDFBFA9D" box="[468,543,1308,1331]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="4" pageNumber="89">Fig. 43</figureCitation>
) dilated, but narrower than in males, gradually narrowed distad. First tarsomere slightly narrower than apical margin of protibia and only a little wider than the second. Apical margins of the first and second protarsomere (
<figureCitation id="1336C7EEFFEA0C76FE848D5AFC37FADD" box="[901,979,1372,1395]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="4" pageNumber="89">Fig. 45</figureCitation>
) with distinct median tooth.
</paragraph>
<paragraph id="8BB2DB6BFFEA0C76FDB48D9AFE73F99D" blockId="4.[151,1127,220,1587]" pageId="4" pageNumber="89">
Sternum VIII (
<figureCitation id="1336C7EEFFEA0C76FC538D9AFE44FA1D" box="[338,416,1436,1459]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="4" pageNumber="89">Fig. 20</figureCitation>
) as wide as long, on inner margin with long and tapering spiculum ventrale, outer margin rounded with long setae. Terga IX and X (
<figureCitation id="1336C7EEFFEA0C76FE498DBAFC70FA7D" box="[840,916,1468,1491]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="4" pageNumber="89">Fig. 17</figureCitation>
) feebly sclerotized, only apex of tergum X and lateral margins of tergum IX more sclerotized. Tergum X long and narrow, dilated in apical portion, apex rounded with long setae. Stylus (
<figureCitation id="1336C7EEFFEA0C76FEB08DFAFC19F9BD" box="[945,1021,1532,1555]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="4" pageNumber="89">Fig. 16</figureCitation>
) long and narrow, with long setae.
</paragraph>
<caption id="DF728BE3FFEB0C77FD708DE8FBA6F999" ID-DOI="http://doi.org/10.5281/zenodo.5372940" ID-Zenodo-Dep="5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="5" pageNumber="90" startId="5.[113,150,1518,1537]" targetBox="[133,1074,219,1457]" targetPageId="5">
<paragraph id="8BB2DB6BFFEB0C77FD708DE8FBA6F999" blockId="5.[113,1090,1518,1591]" pageId="5" pageNumber="90">
Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – 
<taxonomicName id="4C0DA0E8FFEB0C77FDFD8E0FFDD5F9B2" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[252,561,1545,1564]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
<emphasis id="B9790779FFEB0C77FDFD8E0FFE83F9B2" box="[252,359,1545,1564]" italics="true" pageId="5" pageNumber="90">Philomessor</emphasis>
(
<emphasis id="B9790779FFEB0C77FC758E0FFE39F9B2" box="[372,477,1545,1564]" italics="true" pageId="5" pageNumber="90">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFEB0C77FCEA8E0FFDD5F9B2" box="[491,561,1545,1564]" italics="true" pageId="5" pageNumber="90">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFEB0C77FF368E0FFD9EF9B2" box="[567,634,1545,1564]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
(HT); 3, 4, 8 – 
<emphasis id="B9790779FFEB0C77FE028E0FFCF4F9B2" box="[771,784,1545,1564]" italics="true" pageId="5" pageNumber="90">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFEB0C77FE1D8E0FFBE6F9B2" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[796,1026,1545,1564]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="bolivari" subGenus="Attumbrinus">
(
<emphasis id="B9790779FFEB0C77FE228E0FFCD4F9B2" box="[803,816,1545,1564]" italics="true" pageId="5" pageNumber="90">A</emphasis>
.) 
<emphasis id="B9790779FFEB0C77FE438E0FFC62F9B2" box="[834,902,1545,1564]" italics="true" pageId="5" pageNumber="90">bolivari</emphasis>
<bibRefCitation id="EF9CA69AFFEB0C77FE8D8E0FFBE6F9B2" author="JEANNEL R." box="[908,1026,1545,1564]" pageId="5" pageNumber="90" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
(3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – 
<taxonomicName id="4C0DA0E8FFEB0C77FF2D8E22FCD0F999" authority="(Portevin, 1907)" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" box="[556,820,1572,1591]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
<emphasis id="B9790779FFEB0C77FF2D8E22FDDDF999" box="[556,569,1572,1591]" italics="true" pageId="5" pageNumber="90">P</emphasis>
. (
<emphasis id="B9790779FFEB0C77FF4A8E22FDBCF999" box="[587,600,1572,1591]" italics="true" pageId="5" pageNumber="90">A</emphasis>
.) 
<emphasis id="B9790779FFEB0C77FF688E22FD47F999" box="[617,675,1572,1591]" italics="true" pageId="5" pageNumber="90">cloueti</emphasis>
(
<bibRefCitation id="EF9CA69AFFEB0C77FFAF8E22FCCBF999" author="PORTEVIN G." box="[686,815,1572,1591]" pageId="5" pageNumber="90" pagination="67 - 82" refId="ref8747" refString="PORTEVIN G. 1907: Clavicornes nouveaux du groupe des necrophages. Annales de la Societe Entomologique de France 76: 67 - 82." type="journal article" year="1907">Portevin, 1907</bibRefCitation>
)
</taxonomicName>
(5 – HT; 7 – Kairous, Algeria).
</paragraph>
</caption>
<caption id="DF728BE3FFE80C74FD968B4CFE42FC3C" ID-DOI="http://doi.org/10.5281/zenodo.5372942" ID-Zenodo-Dep="5372942" httpUri="https://zenodo.org/record/5372942/files/figure.png" pageId="6" pageNumber="91" startId="6.[151,188,842,861]" targetBox="[171,1113,231,805]" targetPageId="6">
<paragraph id="8BB2DB6BFFE80C74FD968B4CFE42FC3C" blockId="6.[151,1127,842,914]" pageId="6" pageNumber="91">
Figs 9–12. Genital segments in ventral view, males. 9, 10 – 
<taxonomicName id="4C0DA0E8FFE80C74FF9F8B4CFC37FCF3" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[670,979,842,861]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FF9F8B4CFCEDFCF3" box="[670,777,842,861]" italics="true" pageId="6" pageNumber="91">Philomessor</emphasis>
(
<emphasis id="B9790779FFE80C74FE178B4CFC9BFCF3" box="[790,895,842,861]" italics="true" pageId="6" pageNumber="91">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFE80C74FE8C8B4CFC37FCF3" box="[909,979,842,861]" italics="true" pageId="6" pageNumber="91">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE80C74FED88B4CFBF8FCF3" box="[985,1052,842,861]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
(9 – PT, Tizi-n-Tretten near Ifrane, Morocco; 10 – HT); 11 – 
<emphasis id="B9790779FFE80C74FF588B62FD82FCD9" box="[601,614,868,887]" italics="true" pageId="6" pageNumber="91">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE80C74FF708B62FCB3FCD9" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[625,855,868,887]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="bolivari" subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE80C74FF788B62FD62FCD9" box="[633,646,868,887]" italics="true" pageId="6" pageNumber="91">A</emphasis>
.) 
<emphasis id="B9790779FFE80C74FF998B62FD38FCD9" box="[664,732,868,887]" italics="true" pageId="6" pageNumber="91">bolivari</emphasis>
<bibRefCitation id="EF9CA69AFFE80C74FFE08B62FCB3FCD9" author="JEANNEL R." box="[737,855,868,887]" pageId="6" pageNumber="91" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
(HT); 12 – 
<taxonomicName id="4C0DA0E8FFE80C74FEBC8B62FF1AFC3C" authority="(Portevin, 1907)" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FEBC8B62FC2EFCD9" box="[957,970,868,887]" italics="true" pageId="6" pageNumber="91">P</emphasis>
. (
<emphasis id="B9790779FFE80C74FEDD8B62FC0DFCD9" box="[988,1001,868,887]" italics="true" pageId="6" pageNumber="91">A</emphasis>
.) 
<emphasis id="B9790779FFE80C74FEFA8B62FBD1FCD9" box="[1019,1077,868,887]" italics="true" pageId="6" pageNumber="91">cloueti</emphasis>
(
<bibRefCitation id="EF9CA69AFFE80C74F9408B62FF1DFC3C" author="PORTEVIN G." pageId="6" pageNumber="91" pagination="67 - 82" refId="ref8747" refString="PORTEVIN G. 1907: Clavicornes nouveaux du groupe des necrophages. Annales de la Societe Entomologique de France 76: 67 - 82." type="journal article" year="1907">Portevin, 1907</bibRefCitation>
)
</taxonomicName>
(Kairous, Algeria).
</paragraph>
</caption>
<paragraph id="8BB2DB6BFFE80C74FD968BDBFC3AFB3A" blockId="6.[151,1127,989,1588]" pageId="6" pageNumber="91">
<emphasis id="B9790779FFE80C74FD968BDBFE7AFC5A" bold="true" box="[151,414,989,1012]" pageId="6" pageNumber="91">Geographic variability.</emphasis>
Populations from Tizi-n-Tretten near Ifrane and Oukaïmeden somewhat differ in the size and the shape of the pronotum. Specimens from Tizi-n-Tretten are in average larger, their pronota are more convex, less narrowed anteriorly and posterior margin more emarginate at posterior angles than in the specimens from Oukaïmeden. With respect to coincident habitual proportions and further morphological characters given above I consider both populations conspecific and the mentioned differences to be local variation.
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFE80C7AFD968C9BFC86FBD8" lastPageId="8" lastPageNumber="93" pageId="6" pageNumber="91" type="diagnosis">
<paragraph id="8BB2DB6BFFE80C74FD968C9BFC96F99A" blockId="6.[151,1127,989,1588]" pageId="6" pageNumber="91">
<emphasis id="B9790779FFE80C74FD968C9BFE75FB1A" bold="true" box="[151,401,1181,1204]" pageId="6" pageNumber="91">Differential diagnosis.</emphasis>
<taxonomicName id="4C0DA0E8FFE80C74FC978C9BFD9CFB1A" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[406,632,1181,1204]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov.">
<emphasis id="B9790779FFE80C74FC978C9BFD9CFB1A" box="[406,632,1181,1204]" italics="true" pageId="6" pageNumber="91">Philomessor lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE80C74FF7F8C9BFD34FB1A" box="[638,720,1181,1204]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
is placed in the subgenus 
<taxonomicName id="4C0DA0E8FFE80C74FEE28C9BFECCFB7A" authority="Jeannel, 1936" authorityName="Bolivari Jeannel" authorityYear="1936" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="subGenus" subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FEE28C9BFB83FB1A" box="[995,1127,1181,1204]" italics="true" pageId="6" pageNumber="91">Attumbrinus</emphasis>
<bibRefCitation id="EF9CA69AFFE80C74FD968CBBFECCFB7A" author="JEANNEL R." box="[151,296,1213,1236]" pageId="6" pageNumber="91" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
because of the long and erect pubescence of elytra (
<figureCitation id="1336C7EEFFE80C74FE478CBBFC76FB7A" box="[838,914,1213,1236]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="6" pageNumber="91">Fig. 34</figureCitation>
) and fine, dense and long pubescence of mesotibiae (
<figureCitation id="1336C7EEFFE80C74FCEE8CDBFDA5FB5A" box="[495,577,1245,1268]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="6" pageNumber="91">Fig. 44</figureCitation>
). It differs from 
<taxonomicName id="4C0DA0E8FFE80C74FFF08CDBFECBFABA" authority="Ruzicka, 1996" authorityName="Ruzicka" authorityYear="1996" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="henroti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FFF08CDBFC92FB5A" box="[753,886,1245,1268]" italics="true" pageId="6" pageNumber="91">Philomessor</emphasis>
(
<emphasis id="B9790779FFE80C74FE868CDBFBEFFB5A" box="[903,1035,1245,1268]" italics="true" pageId="6" pageNumber="91">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFE80C74F91B8CDBFB82FB5A" box="[1050,1126,1245,1268]" italics="true" pageId="6" pageNumber="91">henroti</emphasis>
<bibRefCitation id="EF9CA69AFFE80C74FD968CFBFECBFABA" author="RUZICKA J." box="[151,303,1277,1300]" pageId="6" pageNumber="91" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">Růžička, 1996</bibRefCitation>
</taxonomicName>
first of all in the shape of pronotum, which is not deeply emarginate at the base besides posterior angles (
<figureCitation id="1336C7EEFFE80C74FCA08D1BFDFDFA9A" box="[417,537,1309,1332]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="6" pageNumber="91">Figs 26, 27</figureCitation>
vs. 32, 40), different pubescence and double punctation of elytra (
<figureCitation id="1336C7EEFFE80C74FDFF8D3BFE92FAFA" box="[254,374,1341,1364]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="6" pageNumber="91">Figs 33, 34</figureCitation>
vs. 39) and also in narrow body and different shape of aedeagus. From 
<taxonomicName id="4C0DA0E8FFE80C74FD968D5BFE01FADA" authority="(Portevin, 1907)" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" box="[151,485,1373,1396]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FD968D5BFF48FADA" box="[151,172,1373,1396]" italics="true" pageId="6" pageNumber="91">P.</emphasis>
(
<emphasis id="B9790779FFE80C74FDBD8D5BFF37FADA" box="[188,211,1373,1396]" italics="true" pageId="6" pageNumber="91">A.</emphasis>
) 
<emphasis id="B9790779FFE80C74FDE58D5BFEC9FADA" box="[228,301,1373,1396]" italics="true" pageId="6" pageNumber="91">cloueti</emphasis>
(
<bibRefCitation id="EF9CA69AFFE80C74FC3D8D5BFE3AFADA" author="PORTEVIN G." box="[316,478,1373,1396]" pageId="6" pageNumber="91" pagination="67 - 82" refId="ref8747" refString="PORTEVIN G. 1907: Clavicornes nouveaux du groupe des necrophages. Annales de la Societe Entomologique de France 76: 67 - 82." type="journal article" year="1907">Portevin, 1907</bibRefCitation>
)
</taxonomicName>
it differs especially in the shape of the posterior margin of pronotum, which is straight in 
<taxonomicName id="4C0DA0E8FFE80C74FCDB8D7BFD8EFA3A" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" box="[474,618,1405,1428]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FCDB8D7BFE0AFA3A" box="[474,494,1405,1428]" italics="true" pageId="6" pageNumber="91">P.</emphasis>
(
<emphasis id="B9790779FFE80C74FCFD8D7BFDF7FA3A" box="[508,531,1405,1428]" italics="true" pageId="6" pageNumber="91">A.</emphasis>
) 
<emphasis id="B9790779FFE80C74FF208D7BFD8EFA3A" box="[545,618,1405,1428]" italics="true" pageId="6" pageNumber="91">cloueti</emphasis>
</taxonomicName>
(
<figureCitation id="1336C7EEFFE80C74FF798D7BFD22FA3A" box="[632,710,1405,1428]" captionStart="Figs 49–54. 49, 51, 53" captionStartId="14.[151,188,1161,1180]" captionTargetBox="[151,1126,219,1141]" captionTargetId="figure-10@14.[151,1126,219,1141]" captionTargetPageId="14" captionText="Figs 49–54. 49, 51, 53 – metatibia in lateral view, males; 50 – antenna, male; 52 – antenna, female; 54 – pronotum, male.49 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden,Morocco); 50–52 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 53–54 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372956" httpUri="https://zenodo.org/record/5372956/files/figure.png" pageId="6" pageNumber="91">Fig. 54</figureCitation>
), but convex in 
<emphasis id="B9790779FFE80C74FE6C8D7BFC99FA3A" box="[877,893,1405,1428]" italics="true" pageId="6" pageNumber="91">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE80C74FE8B8D7BFBEBFA3A" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[906,1039,1405,1428]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE80C74FE928D7BFC47FA3A" box="[915,931,1405,1428]" italics="true" pageId="6" pageNumber="91">A</emphasis>
.) 
<emphasis id="B9790779FFE80C74FEB98D7BFBEBFA3A" box="[952,1039,1405,1428]" italics="true" pageId="6" pageNumber="91">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE80C74F9148D7BFB83FA3A" box="[1045,1127,1405,1428]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1336C7EEFFE80C74FD9E8D9BFEF3FA1A" box="[159,279,1437,1460]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="6" pageNumber="91">Figs 26, 27</figureCitation>
); pronotum is as wide as elytra at humeri in 
<taxonomicName id="4C0DA0E8FFE80C74FFDE8D9BFC94FA1A" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" box="[735,880,1437,1460]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE80C74FFDE8D9BFD0BFA1A" box="[735,751,1437,1460]" italics="true" pageId="6" pageNumber="91">P</emphasis>
. (
<emphasis id="B9790779FFE80C74FE028D9BFCF7FA1A" box="[771,787,1437,1460]" italics="true" pageId="6" pageNumber="91">A</emphasis>
.) 
<emphasis id="B9790779FFE80C74FE298D9BFC94FA1A" box="[808,880,1437,1460]" italics="true" pageId="6" pageNumber="91">cloueti</emphasis>
</taxonomicName>
(
<figureCitation id="1336C7EEFFE80C74FE7C8D9BFC2FFA1A" box="[893,971,1437,1460]" captionStart="Figs 30–32" captionStartId="11.[113,150,1515,1534]" captionTargetBox="[113,1088,222,1430]" captionTargetId="figure-11@11.[113,1088,222,1452]" captionTargetPageId="11" captionText="Figs 30–32. General habitus of types with original labels, males. 30 – Philomessor (Attumbrinus) cloueti (Portevin, 1907), holotype; 31 – P. (A.) bolivari Jeannel, 1936, lectotype; 32 – P. (A.) henroti Růžička, 1996, holotype. An asterisk ‘*’ denotes upper and under side of the same label." figureDoi="http://doi.org/10.5281/zenodo.5372950" httpUri="https://zenodo.org/record/5372950/files/figure.png" pageId="6" pageNumber="91">Fig. 30</figureCitation>
), but distinctly wider in 
<emphasis id="B9790779FFE80C74FDF78DBBFEE2FA7A" box="[246,262,1469,1492]" italics="true" pageId="6" pageNumber="91">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE80C74FC158DBBFE7FFA7A" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[276,411,1469,1492]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE80C74FC1C8DBBFEC9FA7A" box="[285,301,1469,1492]" italics="true" pageId="6" pageNumber="91">A</emphasis>
.) 
<emphasis id="B9790779FFE80C74FC458DBBFE7FFA7A" box="[324,411,1469,1492]" italics="true" pageId="6" pageNumber="91">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE80C74FCA38DBBFE11FA7A" box="[418,501,1469,1492]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1336C7EEFFE80C74FF028DBBFD9BFA7A" box="[515,639,1469,1492]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="6" pageNumber="91">Figs 26, 27</figureCitation>
); both species further differ also in the length of terminal antennomeres (
<figureCitation id="1336C7EEFFE80C74FCB58DDBFDC8FA5A" box="[436,556,1501,1524]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="6" pageNumber="91">Figs 14, 15</figureCitation>
vs. 13), more outstanding difference between the two kinds of punctation and longer setae on elytra in 
<emphasis id="B9790779FFE80C74FF9A8DFBFD4FF9BA" box="[667,683,1533,1556]" italics="true" pageId="6" pageNumber="91">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE80C74FFB98DFBFCDAF9BA" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[696,830,1533,1556]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE80C74FFC08DFBFD35F9BA" box="[705,721,1533,1556]" italics="true" pageId="6" pageNumber="91">A</emphasis>
.) 
<emphasis id="B9790779FFE80C74FFE68DFBFCDAF9BA" box="[743,830,1533,1556]" italics="true" pageId="6" pageNumber="91">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE80C74FE458DFBFC72F9BA" box="[836,918,1533,1556]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1336C7EEFFE80C74FEA28DFBFBFFF9BA" box="[931,1051,1533,1556]" captionStart="Figs 33–40. 33, 35, 37, 39" captionStartId="12.[151,188,1484,1503]" captionTargetBox="[151,1126,221,1455]" captionTargetId="figure-10@12.[151,1126,221,1456]" captionTargetPageId="12" captionText="Figs 33–40. 33, 35, 37, 39 – punctation of elytron in dorsal view, males; 34, 36, 38 – pubescence of elytron in lateral view, males; 40 – posterior margin of pronotum in posterolateral view, male. 33, 34 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden, Morocco); 35, 36 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 37, 38 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria); 39, 40 – P. (A.) henroti Růžička, 1996 (HT)." figureDoi="http://doi.org/10.5281/zenodo.5372952" httpUri="https://zenodo.org/record/5372952/files/figure.png" pageId="6" pageNumber="91">Figs 33, 34</figureCitation>
vs. 37, 38), and in the shape of the male genital segment (
<figureCitation id="1336C7EEFFE80C74FFB18E1BFCF8F99A" box="[688,796,1565,1588]" captionStart="Figs 9–12" captionStartId="6.[151,188,842,861]" captionTargetBox="[171,1113,231,805]" captionTargetId="figure-10@6.[151,1126,216,819]" captionTargetPageId="6" captionText="Figs 9–12. Genital segments in ventral view, males. 9, 10 – Philomessor (Attumbrinus) lackneri sp. nov. (9 – PT, Tizi-n-Tretten near Ifrane, Morocco; 10 – HT); 11 – P. (A.) bolivari Jeannel, 1936 (HT); 12 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372942" httpUri="https://zenodo.org/record/5372942/files/figure.png" pageId="6" pageNumber="91">Figs 9, 10</figureCitation>
vs. 12).
</paragraph>
<caption id="DF728BE3FFE90C75FD708DB3FEF4F99D" ID-DOI="http://doi.org/10.5281/zenodo.5372944" ID-Zenodo-Dep="5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="7" pageNumber="92" startId="7.[113,150,1461,1480]" targetBox="[133,1072,219,1402]" targetPageId="7">
<paragraph id="8BB2DB6BFFE90C75FD708DB3FEF4F99D" blockId="7.[113,1089,1461,1587]" pageId="7" pageNumber="92">
Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – 
<taxonomicName id="4C0DA0E8FFE90C75FED78DD6FE16FA50" authority="(Portevin, 1907) (HT)" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE90C75FED78DD6FBA5FA4D" box="[982,1089,1488,1507]" italics="true" pageId="7" pageNumber="92">Philomessor</emphasis>
(
<emphasis id="B9790779FFE90C75FD798DEDFF05FA50" box="[120,225,1515,1534]" italics="true" pageId="7" pageNumber="92">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFE90C75FDF18DEDFECEFA50" box="[240,298,1515,1534]" italics="true" pageId="7" pageNumber="92">cloueti</emphasis>
(
<bibRefCitation id="EF9CA69AFFE90C75FC368DEDFE5DFA50" author="PORTEVIN G." box="[311,441,1515,1534]" pageId="7" pageNumber="92" pagination="67 - 82" refId="ref8747" refString="PORTEVIN G. 1907: Clavicornes nouveaux du groupe des necrophages. Annales de la Societe Entomologique de France 76: 67 - 82." type="journal article" year="1907">Portevin, 1907</bibRefCitation>
) (HT)
</taxonomicName>
; 14–17, 20 – 
<emphasis id="B9790779FFE90C75FF6C8DEDFD9EFA50" box="[621,634,1515,1534]" italics="true" pageId="7" pageNumber="92">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE90C75FF878DEDFD10FA50" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[646,756,1515,1534]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE90C75FF8C8DEDFD7EFA50" box="[653,666,1515,1534]" italics="true" pageId="7" pageNumber="92">A</emphasis>
.) 
<emphasis id="B9790779FFE90C75FFAF8DEDFD10FA50" box="[686,756,1515,1534]" italics="true" pageId="7" pageNumber="92">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE90C75FFFA8DEDFCDBFA50" box="[763,831,1515,1534]" pageId="7" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
(14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – 
<emphasis id="B9790779FFE90C75FFFE8E03FCE8F9B6" box="[767,780,1541,1560]" italics="true" pageId="7" pageNumber="92">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE90C75FE178E03FC1EF9B6" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[790,1018,1541,1560]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="bolivari" subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE90C75FE1C8E03FCCEF9B6" box="[797,810,1541,1560]" italics="true" pageId="7" pageNumber="92">A</emphasis>
.) 
<emphasis id="B9790779FFE90C75FE3D8E03FC64F9B6" box="[828,896,1541,1560]" italics="true" pageId="7" pageNumber="92">bolivari</emphasis>
<bibRefCitation id="EF9CA69AFFE90C75FE858E03FC1EF9B6" author="JEANNEL R." box="[900,1018,1541,1560]" pageId="7" pageNumber="92" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
(Oukaïmeden, Morocco).
</paragraph>
</caption>
<caption id="DF728BE3FFE60C7AFD968AE2FE69FCE9" ID-DOI="http://doi.org/10.5281/zenodo.5372946" ID-Zenodo-Dep="5372946" httpUri="https://zenodo.org/record/5372946/files/figure.png" pageId="8" pageNumber="93" startId="8.[151,188,740,759]" targetBox="[162,1118,224,725]" targetPageId="8">
<paragraph id="8BB2DB6BFFE60C7AFD968AE2FE69FCE9" blockId="8.[151,1127,740,839]" pageId="8" pageNumber="93">
Figs 21–25. 21 – antenna, male; 22 – aedeagus in dorsal view; 23 – aedeagus in lateral view; 24 – genital segment in ventral view, male; 25 – aedeagus in dorsolateral view. 21–24 – 
<taxonomicName id="4C0DA0E8FFE60C7AFFBD8AF8FB83FCBF" authority="Ruzicka, 1996" authorityName="Ruzicka" authorityYear="1996" box="[700,1127,766,785]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="henroti" subGenus="Attumbrinus">
<emphasis id="B9790779FFE60C7AFFBD8AF8FCC3FCBF" box="[700,807,766,785]" italics="true" pageId="8" pageNumber="93">Philomessor</emphasis>
(
<emphasis id="B9790779FFE60C7AFE328AF8FC78FCBF" box="[819,924,766,785]" italics="true" pageId="8" pageNumber="93">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFE60C7AFEA88AF8FC02FCBF" box="[937,998,766,785]" italics="true" pageId="8" pageNumber="93">henroti</emphasis>
<bibRefCitation id="EF9CA69AFFE60C7AFEEA8AF8FB83FCBF" author="RUZICKA J." box="[1003,1127,766,785]" pageId="8" pageNumber="93" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">Růžička, 1996</bibRefCitation>
</taxonomicName>
(HT); 25 – 
<emphasis id="B9790779FFE60C7AFDF68B1FFEE0FC82" box="[247,260,793,812]" italics="true" pageId="8" pageNumber="93">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE60C7AFC0F8B1FFDD4FC82" authority="(Portevin, 1907) (HT)" authorityName="HT" baseAuthorityName="Portevin" baseAuthorityYear="1907" box="[270,560,793,812]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="cloueti" subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE60C7AFC148B1FFEC6FC82" box="[277,290,793,812]" italics="true" pageId="8" pageNumber="93">A</emphasis>
.) 
<emphasis id="B9790779FFE60C7AFC358B1FFE8AFC82" box="[308,366,793,812]" italics="true" pageId="8" pageNumber="93">cloueti</emphasis>
(
<bibRefCitation id="EF9CA69AFFE60C7AFC788B1FFE1DFC82" author="PORTEVIN G." box="[377,505,793,812]" pageId="8" pageNumber="93" pagination="67 - 82" refId="ref8747" refString="PORTEVIN G. 1907: Clavicornes nouveaux du groupe des necrophages. Annales de la Societe Entomologique de France 76: 67 - 82." type="journal article" year="1907">Portevin, 1907</bibRefCitation>
) (HT)
</taxonomicName>
. Figs 21–24 modified from 
<bibRefCitation id="EF9CA69AFFE60C7AFE1D8B1FFC46FC82" author="RUZICKA J." box="[796,930,793,812]" pageId="8" pageNumber="93" pagination="623 - 628" refId="ref8772" refString="RUZICKA J. 1996: Description of Philomessor (Attumbrinus) henroti sp. n. from Morocco and notes on P. (A.) bolivari (Coleoptera: Leiodidae: Cholevinae). European Journal of Entomology 93: 623 - 628." type="journal article" year="1996">RŮŽIČKA (1996)</bibRefCitation>
, Fig. 25 modified from 
<bibRefCitation id="EF9CA69AFFE60C7AFD968B32FEFFFCE9" author="JEANNEL R." box="[151,283,820,839]" pageId="8" pageNumber="93" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">JEANNEL (1936)</bibRefCitation>
; not to scale.
</paragraph>
</caption>
<paragraph id="8BB2DB6BFFE60C7AFDB48B79FC86FBD8" blockId="8.[151,1127,895,1590]" pageId="8" pageNumber="93">
<taxonomicName id="4C0DA0E8FFE60C7AFDB48B79FDDCFC38" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[181,568,895,918]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
<emphasis id="B9790779FFE60C7AFDB48B79FEDEFC38" box="[181,314,895,918]" italics="true" pageId="8" pageNumber="93">Philomessor</emphasis>
(
<emphasis id="B9790779FFE60C7AFC4D8B79FE34FC38" box="[332,464,895,918]" italics="true" pageId="8" pageNumber="93">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFE60C7AFCE08B79FDDCFC38" box="[481,568,895,918]" italics="true" pageId="8" pageNumber="93">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE60C7AFF408B79FD71FC38" box="[577,661,895,918]" pageId="8" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to 
<emphasis id="B9790779FFE60C7AFE628B79FC97FC38" box="[867,883,895,918]" italics="true" pageId="8" pageNumber="93">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE60C7AFE838B79FF2BFC18" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="bolivari" subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE60C7AFE8A8B79FC7FFC38" box="[907,923,895,918]" italics="true" pageId="8" pageNumber="93">A</emphasis>
.) 
<emphasis id="B9790779FFE60C7AFEB28B79FBE3FC38" box="[947,1031,895,918]" italics="true" pageId="8" pageNumber="93">bolivari</emphasis>
<bibRefCitation id="EF9CA69AFFE60C7AF9118B79FF2BFC18" author="JEANNEL R." pageId="8" pageNumber="93" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
, from which it differs in the narrower shape of body (
<figureCitation id="1336C7EEFFE60C7AFE188B99FC71FC18" box="[793,917,927,950]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="8" pageNumber="93">Figs 26, 27</figureCitation>
vs. 28, 29) and the pubescence of metatibiae (
<figureCitation id="1336C7EEFFE60C7AFCB18BB9FDE4FC78" box="[432,512,959,982]" captionStart="Figs 49–54. 49, 51, 53" captionStartId="14.[151,188,1161,1180]" captionTargetBox="[151,1126,219,1141]" captionTargetId="figure-10@14.[151,1126,219,1141]" captionTargetPageId="14" captionText="Figs 49–54. 49, 51, 53 – metatibia in lateral view, males; 50 – antenna, male; 52 – antenna, female; 54 – pronotum, male.49 – Philomessor (Attumbrinus) lackneri sp. nov. (PT, Oukaïmeden,Morocco); 50–52 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 53–54 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372956" httpUri="https://zenodo.org/record/5372956/files/figure.png" pageId="8" pageNumber="93">Figs 49</figureCitation>
vs. 51), in males in its straight metatibiae (
<figureCitation id="1336C7EEFFE60C7AFEC28BB9FBF7FC78" box="[963,1043,959,982]" captionStart="Figs 26–29. 26, 28" captionStartId="10.[151,188,1535,1554]" captionTargetBox="[228,1045,236,1484]" captionTargetId="figure-10@10.[202,1075,208,1506]" captionTargetPageId="10" captionText="Figs 26–29. 26, 28 – general habitus, males; 27, 29 – general habitus, females. 26, 27 – Philomessor (Attumbrinus) lackneri sp. nov. (27 – HT, 29 – AT); 28, 29 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372948" httpUri="https://zenodo.org/record/5372948/files/figure.png" pageId="8" pageNumber="93">Figs 26</figureCitation>
vs. 28), the shape of aedeagus and the length of parameres (
<figureCitation id="1336C7EEFFE60C7AFFB18BD9FD16FC58" box="[688,754,991,1014]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="8" pageNumber="93">Figs 1</figureCitation>
vs. 3), the shape of genital segment (
<figureCitation id="1336C7EEFFE60C7AFD9E8BF9FEEFFBB8" box="[159,267,1023,1046]" captionStart="Figs 9–12" captionStartId="6.[151,188,842,861]" captionTargetBox="[171,1113,231,805]" captionTargetId="figure-10@6.[151,1126,216,819]" captionTargetPageId="6" captionText="Figs 9–12. Genital segments in ventral view, males. 9, 10 – Philomessor (Attumbrinus) lackneri sp. nov. (9 – PT, Tizi-n-Tretten near Ifrane, Morocco; 10 – HT); 11 – P. (A.) bolivari Jeannel, 1936 (HT); 12 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372942" httpUri="https://zenodo.org/record/5372942/files/figure.png" pageId="8" pageNumber="93">Figs 9, 10</figureCitation>
vs. 11) and the emargination of the outer margin of the sternum VIII (
<figureCitation id="1336C7EEFFE60C7AFEFC8BF9FBA5FBB8" box="[1021,1089,1023,1046]" captionStart="Figs 1–8. 1, 3" captionStartId="5.[113,150,1518,1537]" captionTargetBox="[133,1074,219,1457]" captionTargetId="figure-11@5.[117,1084,200,1476]" captionTargetPageId="5" captionText="Figs 1–8. 1, 3 – aedeagus in dorsal view; 2, 4, 5 – aedeagus in lateral view; 6–8 – sternum VIII in ventral view, males. 1, 2, 6 – Philomessor (Attumbrinus) lackneri sp. nov. (HT); 3, 4, 8 – P. (A.) bolivari Jeannel, 1936 (3, 8 – LT; 4 – Tizi-n-Tretten near Ifrane, Morocco); 5, 7 – P. (A.) cloueti (Portevin, 1907) (5 – HT; 7 – Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372940" httpUri="https://zenodo.org/record/5372940/files/figure.png" pageId="8" pageNumber="93">Figs 6</figureCitation>
vs. 8); in females in dilated protarsi (
<figureCitation id="1336C7EEFFE60C7AFCF88C19FDADFB98" box="[505,585,1055,1078]" captionStart="Figs 41–48. 41" captionStartId="13.[113,150,1486,1505]" captionTargetBox="[113,1088,221,1455]" captionTargetId="figure-11@13.[113,1088,221,1456]" captionTargetPageId="13" captionText="Figs 41–48. 41 – protarsus, male; 42 – protibia in dorsolateral view, male; 43, 47 – protarsus, females; 44, 46, 48 – mesotibia in lateral view, males; 45 – detail of basal protarsomeres, female. 41–45 – Philomessor (Attumbrinus) lackneri sp. nov. (41, 42, 44 – PT, Oukaïmeden, Morocco; 43, 45 – AT); 46, 47 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco); 48 – P. (A.) cloueti (Portevin, 1907) (Kairous, Algeria)." figureDoi="http://doi.org/10.5281/zenodo.5372954" httpUri="https://zenodo.org/record/5372954/files/figure.png" pageId="8" pageNumber="93">Figs 43</figureCitation>
vs. 47), in the shape of terga IX and X (
<figureCitation id="1336C7EEFFE60C7AFEF08C19FBA6FB98" box="[1009,1090,1055,1078]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="8" pageNumber="93">Figs 17</figureCitation>
vs. 19), and in the shape of spiculum ventrale on the sternum VIII (
<figureCitation id="1336C7EEFFE60C7AFE458C39FC71FBF8" box="[836,917,1087,1110]" captionStart="Figs 13–20. 13, 14" captionStartId="7.[113,150,1461,1480]" captionTargetBox="[133,1072,219,1402]" captionTargetId="figure-11@7.[113,1088,216,1412]" captionTargetPageId="7" captionText="Figs 13–20. 13, 14 – antenna, males; 15 – antenna, female; 16 – stylus in dorsal view, female; 17, 19 – terga IX and X in dorsal view, females; 18, 20 – sternum VIII in ventral view, females, sv – spiculum ventrale. 13 – Philomessor (Attumbrinus) cloueti (Portevin, 1907) (HT); 14–17, 20 – P. (A.) lackneri sp. nov. (14 – PT, Tizi-n-Tretten near Ifrane, Morocco; 15, 16, 20 – AT; 17 – PT, Oukaïmeden, Morocco); 18, 19 – P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco)." figureDoi="http://doi.org/10.5281/zenodo.5372944" httpUri="https://zenodo.org/record/5372944/files/figure.png" pageId="8" pageNumber="93">Figs 20</figureCitation>
vs. 18). Diagnostic key to all four species of the subgenus 
<taxonomicName id="4C0DA0E8FFE60C7AFF378C59FD5EFBD8" authorityName="Bolivari Jeannel" authorityYear="1936" box="[566,698,1119,1142]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="subGenus" subGenus="Attumbrinus">
<emphasis id="B9790779FFE60C7AFF378C59FD5EFBD8" box="[566,698,1119,1142]" italics="true" pageId="8" pageNumber="93">Attumbrinus</emphasis>
</taxonomicName>
is given above.
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFE60C7AFD968C79FC8BFA58" pageId="8" pageNumber="93" type="etymology">
<paragraph id="8BB2DB6BFFE60C7AFD968C79FCB4FB18" blockId="8.[151,1127,895,1590]" pageId="8" pageNumber="93">
<emphasis id="B9790779FFE60C7AFD968C79FEFCFB38" bold="true" box="[151,280,1151,1174]" pageId="8" pageNumber="93">Etymology.</emphasis>
Dedicated to the renowned specialist on the family 
<taxonomicName id="4C0DA0E8FFE60C7AFE528C79FC25FB38" authorityName="Gyllenhal" authorityYear="1808" box="[851,961,1151,1174]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
and one of the collectors of the new species, Tomáš Lackner (Košice, 
<collectingCountry id="F31A9BFBFFE60C7AFFE38C99FCA0FB18" box="[738,836,1183,1206]" name="Slovakia" pageId="8" pageNumber="93">Slovakia</collectingCountry>
).
</paragraph>
<paragraph id="8BB2DB6BFFE60C7AFD968CB9FE2FFA38" blockId="8.[151,1127,895,1590]" pageId="8" pageNumber="93">
<emphasis id="B9790779FFE60C7AFD968CB9FE5CFB78" bold="true" box="[151,440,1215,1238]" pageId="8" pageNumber="93">Collecting circumstances.</emphasis>
All specimens of the new species were collected in the ant nest of the genus 
<taxonomicName id="4C0DA0E8FFE60C7AFDFE8CD9FE2EFB58" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" box="[255,458,1247,1270]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B9790779FFE60C7AFDFE8CD9FEA9FB58" box="[255,333,1247,1270]" italics="true" pageId="8" pageNumber="93">Messor</emphasis>
Forel, 1890
</taxonomicName>
(
<taxonomicName id="4C0DA0E8FFE60C7AFCD78CD9FD88FB58" box="[470,620,1247,1270]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
: 
<taxonomicName id="4C0DA0E8FFE60C7AFF728CD9FD15FB58" box="[627,753,1247,1270]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
). The specimens from Oukaïmeden, collected by Peter Koniar and Martin Švarc were found in the nests of 
<taxonomicName id="4C0DA0E8FFE60C7AFEA28CF9FED0FA98" authority="Santschi, 1921" authorityName="Santschi" authorityYear="1921" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="abdelazizi">
<emphasis id="B9790779FFE60C7AFEA28CF9FB82FAB8" box="[931,1126,1279,1302]" italics="true" pageId="8" pageNumber="93">Messor abdelazizi</emphasis>
Santschi, 1921
</taxonomicName>
on pastures (
<figureCitation id="1336C7EEFFE60C7AFCC28D19FDDAFA98" box="[451,574,1311,1334]" captionStart="Figs 55–58. 55" captionStartId="16.[151,188,904,923]" captionTargetBox="[151,1126,219,879]" captionTargetId="figure-10@16.[151,1126,219,880]" captionTargetPageId="16" captionText="Figs 55–58. 55 – habitat of Philomessor (Attumbrinus) lackneri sp. nov. and P. (A.) bolivari Jeannel, 1936 (Oukaïmeden, Morocco, 22.v.2009); 56 – habitat of P. (A.) bolivari (Mischlifen, Morocco, 27.v.2011); 57 – nest of Messor abdelazizi Santschi, 1921, under a stone on a pasture (Oukaïmeden, Morocco, 20.v.2011); 58 – male of P. (A.) bolivari in the nest of Messor abdelazizi (Oukaïmeden, Morocco, 20.v.2011). Photographs by P. Koniar and M. Švarc." figureDoi="http://doi.org/10.5281/zenodo.5372958" httpUri="https://zenodo.org/record/5372958/files/figure.png" pageId="8" pageNumber="93">Figs 55, 57</figureCitation>
) at altitude 
<quantity id="4CF5768EFFE60C7AFFBF8D19FCB6FA98" box="[702,850,1311,1334]" metricMagnitude="3" metricUnit="m" metricValue="2.1134999999999997" metricValueMax="2.357" metricValueMin="1.8699999999999999" pageId="8" pageNumber="93" unit="m" value="2113.5" valueMax="2357.0" valueMin="1870.0">1870–2357 m</quantity>
a.s.l. Specimens from Tizi-n-Tretten near Ifrane were found in the nest of 
<taxonomicName id="4C0DA0E8FFE60C7AFFAA8D39FCFBFAF8" box="[683,799,1343,1366]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9790779FFE60C7AFFAA8D39FD1EFAF8" box="[683,762,1343,1366]" italics="true" pageId="8" pageNumber="93">Messor</emphasis>
sp.
</taxonomicName>
under a big stone on a pasture at the forest border at 
<quantity id="4CF5768EFFE60C7AFC818D59FE35FAD8" box="[384,465,1375,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="93" unit="m" value="1900.0">1900 m</quantity>
a.s.l. Detailed data on the collection of the single female from the locality Ito are unknown.
</paragraph>
<paragraph id="8BB2DB6BFFE60C7AFDB48D99FC8BFA58" blockId="8.[151,1127,895,1590]" pageId="8" pageNumber="93">
<taxonomicName id="4C0DA0E8FFE60C7AFDB48D99FDD1FA18" authority="Vávra, 2015" authorityName="Vávra" authorityYear="2015" box="[181,565,1439,1462]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="lackneri" status="sp. nov." subGenus="Attumbrinus">
<emphasis id="B9790779FFE60C7AFDB48D99FEDEFA18" box="[181,314,1439,1462]" italics="true" pageId="8" pageNumber="93">Philomessor</emphasis>
(
<emphasis id="B9790779FFE60C7AFC4A8D99FE2BFA18" box="[331,463,1439,1462]" italics="true" pageId="8" pageNumber="93">Attumbrinus</emphasis>
) 
<emphasis id="B9790779FFE60C7AFCDF8D99FDD1FA18" box="[478,565,1439,1462]" italics="true" pageId="8" pageNumber="93">lackneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24ABA02FFE60C7AFF3C8D99FD74FA18" box="[573,656,1439,1462]" pageId="8" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
seems to be sympatric and syntopic with 
<emphasis id="B9790779FFE60C7AF94E8D99FBBBFA18" box="[1103,1119,1439,1462]" italics="true" pageId="8" pageNumber="93">P</emphasis>
. 
<taxonomicName id="4C0DA0E8FFE60C7AFD968DB9FE5FFA78" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[151,443,1471,1494]" class="Insecta" family="Leiodidae" genus="Philomessor" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="bolivari" subGenus="Attumbrinus">
(
<emphasis id="B9790779FFE60C7AFDA18DB9FF54FA78" box="[160,176,1471,1494]" italics="true" pageId="8" pageNumber="93">A</emphasis>
.) 
<emphasis id="B9790779FFE60C7AFDC98DB9FEF8FA78" box="[200,284,1471,1494]" italics="true" pageId="8" pageNumber="93">bolivari</emphasis>
<bibRefCitation id="EF9CA69AFFE60C7AFC258DB9FE5FFA78" author="JEANNEL R." box="[292,443,1471,1494]" pageId="8" pageNumber="93" pagination="1 - 433" refId="ref8596" refString="JEANNEL R. 1936: Monographie des Catopidae. Memoires du Museum National d Histoire Naturelle (N. S.) 1: 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
; both species were found together in the same ant nests on two localities (Oukaïmeden and Tizi-n-Tretten near Ifrane) (see 
<figureCitation id="1336C7EEFFE60C7AFE158DD9FC87FA58" box="[788,867,1503,1526]" captionStart="Fig" captionStartId="16.[151,182,1564,1583]" captionTargetBox="[152,1122,1132,1537]" captionTargetId="figure-95@16.[151,1127,1132,1540]" captionTargetPageId="16" captionText="Fig. 59. Distribution of Philomessor (Attumbrinus) Jeannel, 1936 in the northwest of Africa." figureDoi="http://doi.org/10.5281/zenodo.5372960" httpUri="https://zenodo.org/record/5372960/files/figure.png" pageId="8" pageNumber="93">Fig. 59</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31788E0FFE60C7AFD968DF9FCC1F998" pageId="8" pageNumber="93" type="distribution">
<paragraph id="8BB2DB6BFFE60C7AFD968DF9FCC1F998" blockId="8.[151,1127,895,1590]" pageId="8" pageNumber="93">
<emphasis id="B9790779FFE60C7AFD968DF9FECEF9B8" bold="true" box="[151,298,1535,1558]" pageId="8" pageNumber="93">Distribution.</emphasis>
<collectingCountry id="F31A9BFBFFE60C7AFC318DF9FE71F9B8" box="[304,405,1535,1558]" name="Morocco" pageId="8" pageNumber="93">Morocco</collectingCountry>
: Oukaïmeden in the High Atlas Mts.; Ito near Azrou, Tizi-n-Tretten near Ifrane, both localities in the Middle Atlas Mts. (
<figureCitation id="1336C7EEFFE60C7AFFC98E19FCFCF998" box="[712,792,1567,1590]" captionStart="Fig" captionStartId="16.[151,182,1564,1583]" captionTargetBox="[152,1122,1132,1537]" captionTargetId="figure-95@16.[151,1127,1132,1540]" captionTargetPageId="16" captionText="Fig. 59. Distribution of Philomessor (Attumbrinus) Jeannel, 1936 in the northwest of Africa." figureDoi="http://doi.org/10.5281/zenodo.5372960" httpUri="https://zenodo.org/record/5372960/files/figure.png" pageId="8" pageNumber="93">Fig. 59</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>