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<document id="E25B2818A4704FA6DC5B3D4B15FF0B0A" ID-DOI="10.5281/zenodo.7666039" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666039" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677089034480" checkinUser="felipe" docAuthor="Sinclair, Bradley J." docDate="2002" docId="03A39140FFF2FFC0C6A89B5FFDA0FA5C" docLanguage="en" docName="AfrInvertebr.43.1.109122.pdf" docOrigin="African Invertebrates 43" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Edenophorus irwini Sinclair 2002, sp. n." docType="treatment" docVersion="3" lastPageNumber="116" masterDocId="FF9AE938FFF4FFC7C77F9D5BFFBEFFE3" masterDocTitle="Revision of the South African endemic genus Edenophorus Smith (Diptera: Empididae)" masterLastPageNumber="122" masterPageNumber="109" pageNumber="115" updateTime="1699455179651" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8E4FBB0948CD4F52B7C0EC4A3FE89B8D">Revision of the South African endemic genus Edenophorus Smith (Diptera: Empididae)</mods:title>
</mods:titleInfo>
<mods:name id="D74610A534CBE83F248C4ED2FE8732BC" type="personal">
<mods:role id="270BA4599326AFCE78DA69406D8D9102">
<mods:roleTerm id="92A676356A993DB3F428D7166CDC45F1">Author</mods:roleTerm>
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<mods:namePart id="C31EE5D30F2191E2DD96EF7499280227">Sinclair, Bradley J.</mods:namePart>
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<mods:typeOfResource id="797B7F0430B940A285494E6A0EBB3B92">text</mods:typeOfResource>
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<mods:titleInfo id="5CE3834924F67A4170219E19A27A204A">
<mods:title id="AF545E522DEEF68C54161014E5374782">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="60EAEC01918A87655F954F2E97C71530">
<mods:date id="E82DE567F75DC7F3E92C495C2B505169">2002</mods:date>
<mods:detail id="2A1561177F5B9F661360805CFB5D2526" type="pubDate">
<mods:number id="3D1ED857CA6BCF8B9CFC00A71C5A4A65">2002-12-31</mods:number>
</mods:detail>
<mods:detail id="C18E13621981CCEA5F0C60F80443776D" type="volume">
<mods:number id="6E6FB30F03972F6E4E274C5B00E918BF">43</mods:number>
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<mods:start id="F2793C7E9B67705A477A03700DBF2136">109</mods:start>
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<mods:classification id="8CCCA109B67AAD43E7885DF51C369677">journal article</mods:classification>
<mods:identifier id="74F9F8A8732A36B535F72ADFE40B3A8C" type="DOI">10.5281/zenodo.7666039</mods:identifier>
<mods:identifier id="3F78464796570A5FA0A510348BADD746" type="ISSN">2305-2562</mods:identifier>
<mods:identifier id="B88AF288AF857D2833C09FB40EB10848" type="Zenodo-Dep">7666039</mods:identifier>
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<treatment id="03A39140FFF2FFC0C6A89B5FFDA0FA5C" ID-DOI="http://doi.org/10.5281/zenodo.7666062" ID-Zenodo-Dep="7666062" LSID="urn:lsid:plazi:treatment:03A39140FFF2FFC0C6A89B5FFDA0FA5C" httpUri="http://treatment.plazi.org/id/03A39140FFF2FFC0C6A89B5FFDA0FA5C" lastPageId="7" lastPageNumber="116" pageId="6" pageNumber="115">
<subSubSection id="C31073DDFFF2FFC1C6A89B5FFCB6F9FD" box="[471,776,1540,1566]" pageId="6" pageNumber="115" type="nomenclature">
<paragraph id="8BB52056FFF2FFC1C6A89B5FFCB6F9FD" blockId="6.[144,1104,1540,1678]" box="[471,776,1540,1566]" pageId="6" pageNumber="115">
<heading id="D0FD973AFFF2FFC1C6A89B5FFCB6F9FD" box="[471,776,1540,1566]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="115" reason="2">
<taxonomicName id="4C0A5BD5FFF2FFC1C6A89B5FFD78F9FE" ID-CoL="38SRV" authority="Sinclair, 2002" authorityName="Sinclair" authorityYear="2002" box="[471,710,1540,1565]" class="Insecta" family="Empididae" genus="Edenophorus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov.">
<emphasis id="B97EFC44FFF2FFC1C6A89B5FFD78F9FE" bold="true" box="[471,710,1540,1565]" pageId="6" pageNumber="115">Edenophorus irwini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D413FFFF2FFC1C5B39B5EFCB6F9FD" box="[716,776,1541,1566]" pageId="6" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31073DDFFF2FFC0C5499B76FDA0FA5C" lastPageId="7" lastPageNumber="116" pageId="6" pageNumber="115" type="description">
<paragraph id="8BB52056FFF2FFC1C5499B76FD14F9A5" blockId="6.[144,1104,1540,1678]" box="[566,682,1581,1606]" pageId="6" pageNumber="115">
(
<figureCitation id="13313CD3FFF2FFC1C5419B76FD1FF9A5" box="[574,673,1581,1606]" captionStart="Figs 27" captionStartId="3.[182,221,1551,1571]" captionTargetBox="[244,1095,340,1534]" captionTargetId="figure-9@3.[238,1095,312,1605]" captionTargetPageId="3" captionText="Figs 27. Edenophorus characters. 2. E. spinosus sp. n., head. 3. E. hiemalis sp. n., antenna. 4. E. knysna Smith, wing. 5. E. hiemalis sp. n., wing. 67. E. irwini sp. n., male terminalia. 6. Lateral view. 7. Posterior view. Scale bar = 0.1 mm, except Figs 2, 4, 5, where scale bar = 0.5 mm. [Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; epand = epandrium; epand lb = epandrial lobe; goncx apod = gonocoxal apodeme; pgt = postgonite; ph = phallus; sp ch = sperm chamber.]" figureDoi="http://doi.org/10.5281/zenodo.7666043" httpUri="https://zenodo.org/record/7666043/files/figure.png" pageId="6" pageNumber="115">Figs 6, 7</figureCitation>
)
</paragraph>
<paragraph id="8BB52056FFF2FFC1C7EF9B0EFD74F96D" blockId="6.[144,1104,1540,1678]" pageId="6" pageNumber="115">
Etymology: Named in honour of the collector, Mike E. Irwin, who has contributed greatly to the knowledge of South African
<taxonomicName id="4C0A5BD5FFF2FFC1C5119B2EFD78F96D" box="[622,710,1653,1678]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="6" pageNumber="115" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
.
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9D95FE22FEC4" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
<materialsCitation id="3B622A0BFFF3FFC0C7CA9D95FE22FEC4" collectingDate="1974-06-17" collectionCode="NMSA" collectorName="ME Irwin" country="South Africa" county="South Africa" location="Karkloof Range" municipality="Natal Prov" pageId="7" pageNumber="116" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<typeStatus id="54B19EF4FFF3FFC0C7CA9D95FEA3FF04" box="[181,285,206,231]" pageId="7" pageNumber="116" type="holotype">Holotype</typeStatus>
male labelled:
<collectingCountry id="F31D60C6FFF3FFC0C6BB9D95FD2EFF04" box="[452,656,206,231]" name="South Africa" pageId="7" pageNumber="116">SOUTH AFRICA</collectingCountry>
:
<emphasis id="B97EFC44FFF3FFC0C5E69D95FCF9FF05" box="[665,839,206,230]" italics="true" pageId="7" pageNumber="116">
<collectingRegion id="49CEEEB4FFF3FFC0C5E69D95FCF9FF05" box="[665,839,206,230]" country="South Africa" name="KwaZulu-Natal" pageId="7" pageNumber="116">KwaZulu-Natal</collectingRegion>
</emphasis>
:
<collectingCounty id="62D458DAFFF3FFC0C4269D95FC53FF04" box="[857,1005,206,231]" pageId="7" pageNumber="116">South Africa</collectingCounty>
,
<collectingMunicipality id="6BD1BA2CFFF3FFC0C4889D95FBD3FF04" box="[1015,1133,206,231]" pageId="7" pageNumber="116">Natal Prov</collectingMunicipality>
/
<location id="8ED5768DFFF3FFC0C7CA9DB5FED7FEE4" LSID="urn:lsid:plazi:treatment:03A39140FFF2FFC0C6A89B5FFDA0FA5C:8ED5768DFFF3FFC0C7CA9DB5FED7FEE4" box="[181,361,238,263]" country="South Africa" county="South Africa" municipality="Natal Prov" name="Karkloof Range" pageId="7" pageNumber="116" stateProvince="KwaZulu-Natal">Karkloof Range</location>
,near
<location id="8ED5768DFFF3FFC0C6DA9DB5FE7BFEE4" LSID="urn:lsid:plazi:treatment:03A39140FFF2FFC0C6A89B5FFDA0FA5C:8ED5768DFFF3FFC0C6DA9DB5FE7BFEE4" box="[421,453,238,263]" country="South Africa" county="South Africa" municipality="Natal Prov" name="Mt" pageId="7" pageNumber="116" stateProvince="KwaZulu-Natal">Mt</location>
/
<location id="8ED5768DFFF3FFC0C6AD9DB5FDAAFEE4" LSID="urn:lsid:plazi:treatment:03A39140FFF2FFC0C6A89B5FFDA0FA5C:8ED5768DFFF3FFC0C6AD9DB5FDAAFEE4" box="[466,532,238,263]" country="South Africa" county="South Africa" municipality="Natal Prov" name="Alida" pageId="7" pageNumber="116" stateProvince="KwaZulu-Natal">Alida</location>
; temp.forest assoc/
<date id="FFB40696FFF3FFC0C5929DB5FC3DFEE4" box="[749,899,238,263]" pageId="7" pageNumber="116" value="1974-06-17">
<collectingDate id="EFF0FF7EFFF3FFC0C5929DB5FC3DFEE4" box="[749,899,238,263]" pageId="7" pageNumber="116" value="1974-06-17">17 June 1974</collectingDate>
</date>
, near pond/
<collectorName id="26FF4580FFF3FFC0C3769DB5FBC8FEE4" box="[1033,1142,238,263]" pageId="7" pageNumber="116">ME Irwin</collectorName>
(2930Ab) (
<collectionCode id="ED1BB893FFF3FFC0C6469C55FE2EFEC4" box="[313,400,270,295]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="116" type="Museum">NMSA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9C6EFB9AFE8D" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">Recognition: This species is distinguished by its short proboscis, presence of cell dm, and the enormous, erect male terminalia, with their cerci positioned anteriorly.</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9C27FEFFFE76" blockId="7.[181,1142,206,1471]" box="[181,321,380,405]" pageId="7" pageNumber="116">Description:</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9CF9FF49FE58" blockId="7.[181,1142,206,1471]" box="[181,247,418,443]" pageId="7" pageNumber="116">Male:</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9C92FED4FD41" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
<emphasis id="B97EFC44FFF3FFC0C7CA9C92FF4CFE02" box="[181,242,457,481]" italics="true" pageId="7" pageNumber="116">Head</emphasis>
: Eyes nearly touching above antennae; facets above antennae enlarged; setulae on face; 4 pairs of setulae on frons, upper-most longest. Antenna with basal segments slightly paler than the postpedicel; scape very short, pedicel globular with fringe of setulae; postpedicel conical, clothed in fine pubescence; twice length of pedicel, gradually tapered to style. Stout occipital setae in several rows. Proboscis light brown, slightly longer than height of head; palpus one-quarter length of proboscis, pointed bearing long, pale setae.
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9FEAFC51FC69" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
<emphasis id="B97EFC44FFF3FFC0C7CA9FEAFEB8FD2A" box="[181,262,689,713]" italics="true" pageId="7" pageNumber="116">Thorax</emphasis>
: Dusted entirely with brownish grey pruinescence. Thoracic setae generally short, reduced to numerous rows: Acr biserial; dc uniserial, with 4 pairs of long bristles at prescutellar depression; inter-alar setulae scattered, extending to wing base; 2 pprn (lower directed forward, upper directed posteriorly) and numerous short setulae; 1 ph; 3 npl, posterior bristle short; 1 pal; 4 sctl, inner pair stronger, nearly twice length of outer. Antepronotum with 1 pair of short, stout setae. Upper half of anepisternum with greater than 10 setulae; precoxal bridge with 1 stout seta above fore coxa.
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7AF9ECAFE30FC09" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">Legs: Yellowish, darker on apical tarsal segments; clothed in fine setulae, lacking modified setae; fore coxa with 12 pairs of long pale lateral setae on lower half, longer than width of coxa.</paragraph>
<paragraph id="8BB52056FFF3FFC0C7AF9EAAFE7BFBC9" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
Wings (length
<quantity id="4CF28DB3FFF3FFC0C60C9EAAFE73FBE9" box="[371,461,1009,1034]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="116" unit="mm" value="2.4">2.4 mm</quantity>
): Stigma faint. Venation similar to
<taxonomicName id="4C0A5BD5FFF3FFC0C4339EAAFC06FBEA" box="[844,952,1009,1033]" class="Insecta" family="Empididae" genus="Edenophorus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="species" species="knysna">
<emphasis id="B97EFC44FFF3FFC0C4339EAAFC06FBEA" box="[844,952,1009,1033]" italics="true" pageId="7" pageNumber="116">E. knysna</emphasis>
</taxonomicName>
; cell dm present. Halter yellowish brown.
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9962FE13FB91" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
<emphasis id="B97EFC44FFF3FFC0C7CA9962FEA1FBB2" box="[181,287,1081,1105]" italics="true" pageId="7" pageNumber="116">Abdomen</emphasis>
: Pale brown, concolorous with thorax. T8 membranous, S8 divided into 2 small square sclerites.
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7AF9922FDDCFAB1" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
Terminalia (
<figureCitation id="13313CD3FFF3FFC0C6279922FE03FB71" box="[344,445,1145,1170]" captionStart="Figs 27" captionStartId="3.[182,221,1551,1571]" captionTargetBox="[244,1095,340,1534]" captionTargetId="figure-9@3.[238,1095,312,1605]" captionTargetPageId="3" captionText="Figs 27. Edenophorus characters. 2. E. spinosus sp. n., head. 3. E. hiemalis sp. n., antenna. 4. E. knysna Smith, wing. 5. E. hiemalis sp. n., wing. 67. E. irwini sp. n., male terminalia. 6. Lateral view. 7. Posterior view. Scale bar = 0.1 mm, except Figs 2, 4, 5, where scale bar = 0.5 mm. [Abbreviations: cerc = cercus; ej apod = ejaculatory apodeme; epand = epandrium; epand lb = epandrial lobe; goncx apod = gonocoxal apodeme; pgt = postgonite; ph = phallus; sp ch = sperm chamber.]" figureDoi="http://doi.org/10.5281/zenodo.7666043" httpUri="https://zenodo.org/record/7666043/files/figure.png" pageId="7" pageNumber="116">Figs 6, 7</figureCitation>
): Held upright, greatly enlarged compared to abdomen, cercus anterior; cercus and epandrium yellowish. Cercus thinly sclerotised, setose. Epandrium with slightly twisted finger-like lobe on inner margin. Hypandrium reduced. Phallus with large basal, inflated sperm chamber; apex encircled by subapical toothed crown; ejaculatory apodeme broad, stout, articulated at base of chamber, one-third length of terminalia. Postgonite triangular in posterior view, divergently encircling tip of phallus; fused posteriorly with base of phallus.
</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA9804FE33FA9B" blockId="7.[181,1142,206,1471]" box="[181,397,1375,1400]" pageId="7" pageNumber="116">Female: Unknown.</paragraph>
<paragraph id="8BB52056FFF3FFC0C7CA98DDFDA0FA5C" blockId="7.[181,1142,206,1471]" pageId="7" pageNumber="116">
Additional material examined:
<materialsCitation id="3B622A0BFFF3FFC0C57198DDFDA4FA5C" collectingDate="1974-06-17" collectionCode="NMSA" collectorName="ME Irwin" country="South Africa" county="South Africa" location="Karkloof Range" municipality="Natal Prov" pageId="7" pageNumber="116" specimenCount="4" specimenCount-male="4" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<typeStatus id="54B19EF4FFF3FFC0C57198DDFDC1FA7C" box="[526,639,1414,1439]" pageId="7" pageNumber="116" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F31D60C6FFF3FFC0C5F698DDFCEBFA7C" box="[649,853,1414,1439]" name="South Africa" pageId="7" pageNumber="116">SOUTH AFRICA</collectingCountry>
:
<emphasis id="B97EFC44FFF3FFC0C42198DDFBB2FA7D" box="[862,1036,1414,1438]" italics="true" pageId="7" pageNumber="116">
<collectingRegion id="49CEEEB4FFF3FFC0C42198DDFBB2FA7D" box="[862,1036,1414,1438]" country="South Africa" name="KwaZulu-Natal" pageId="7" pageNumber="116">KwaZulu-Natal</collectingRegion>
</emphasis>
:
<specimenCount id="9D0CEBDFFFF3FFC0C36598DDFBCFFA7C" box="[1050,1137,1414,1439]" pageId="7" pageNumber="116" type="male">4 males</specimenCount>
, same data as holotype (
<collectionCode id="ED1BB893FFF3FFC0C6C498FDFDACFA5C" box="[443,530,1446,1471]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="7" pageNumber="116" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>