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<document id="2DB62D340F3CF8439DF36B7EC8691334" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13842FF9F2B39FEE5FB03FCBD" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium itoi Sawada 2008" docType="treatment" docVersion="7" lastPageNumber="82" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="81" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D19819634E72F77E81CB1FFC2292DFFC">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<mods:namePart id="79AA6C9CFC300715C643802F23DD8F7F">Darby, Michael</mods:namePart>
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<mods:title id="34E335C489F1A4A09F740BF5D431A865">European Journal of Taxonomy</mods:title>
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<mods:date id="576892F2BBBDE645B6B73E69509FF698">2020</mods:date>
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<mods:number id="EBD2D4D86CECC37B22945DD6EBA04857">2020-04-02</mods:number>
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<treatment id="039987A13842FF9F2B39FEE5FB03FCBD" ID-DOI="http://doi.org/10.5281/zenodo.3795582" ID-GBIF-Taxon="163877862" ID-Zenodo-Dep="3795582" LSID="urn:lsid:plazi:treatment:039987A13842FF9F2B39FEE5FB03FCBD" httpUri="http://treatment.plazi.org/id/039987A13842FF9F2B39FEE5FB03FCBD" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
<subSubSection id="C32A653C3842FF9E2B39FEE5FC58FEBD" box="[624,964,267,294]" pageId="80" pageNumber="81" type="nomenclature">
<paragraph id="8B8F36B73842FF9E2B39FEE5FC58FEBD" blockId="80.[624,964,267,327]" box="[624,964,267,294]" pageId="80" pageNumber="81">
<heading id="D0C781DB3842FF9E2B39FEE5FC58FEBD" box="[624,964,267,294]" centered="true" fontSize="11" level="2" pageId="80" pageNumber="81" reason="2">
<taxonomicName id="4C304D343842FF9E2B39FEE5FC58FEBD" ID-CoL="VK8D" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[624,964,267,294]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="B944EAA53842FF9E2B39FEE5FC8BFEBD" bold="true" box="[624,791,267,294]" italics="true" pageId="80" pageNumber="81">Cissidium itoi</emphasis>
<bibRefCitation id="EFA14B463842FF9E2A56FEE5FC58FEBD" author="Sawada Y." box="[799,964,267,293]" pageId="80" pageNumber="81" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3842FF9E2BB9FEC3FCDFFEDF" box="[752,835,300,327]" pageId="80" pageNumber="81" type="description">
<paragraph id="8B8F36B73842FF9E2BB9FEC3FCDFFEDF" blockId="80.[624,964,267,327]" box="[752,835,300,327]" pageId="80" pageNumber="81">
<figureCitation id="130B2A323842FF9E2BB9FEC3FCDFFEDF" box="[752,835,300,327]" captionStart="Fig" captionStartId="80.[189,232,1964,1990]" captionTargetBox="[191,1395,1046,1920]" captionTargetId="figure@80.[186,1395,1045,1923]" captionTargetPageId="80" captionText="Fig. 51. Cissidium itoi Sawada, 2008. A. Habitus. B. Pronotum, × 375. C. Mesoventrite showing median process of collar, mid keel, keel and lateral margin, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777386" httpUri="https://zenodo.org/record/3777386/files/figure.png" pageId="80" pageNumber="81">Fig. 51</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3842FF9F29F4FE82FAEAFD12" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="8B8F36B73842FF9E29F4FE82FE36FE1E" blockId="80.[189,426,364,390]" box="[189,426,364,390]" pageId="80" pageNumber="81">
<heading id="D0C781DB3842FF9E29F4FE82FE36FE1E" bold="true" box="[189,426,364,390]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B944EAA53842FF9E29F4FE82FE36FE1E" bold="true" box="[189,426,364,390]" pageId="80" pageNumber="81">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73842FF9E29A3FE77FEFBFE2B" blockId="80.[189,1311,409,469]" box="[234,359,409,435]" pageId="80" pageNumber="81">
<materialsCitation id="3B583CEA3842FF9E29A3FE77FEFBFE2B" ID-GBIF-Occurrence="2609493686" box="[234,359,409,435]" collectingDate="1997-05-01" collectingDateMax="1997-05-03" collectingDateMin="1997-05-01" collectionCode="YS, BMNH" collectorName="T. Ito" country="Japan" location="JAPAN" pageId="80" pageNumber="81" specimenCount="17" typeStatus="paratype">
<emphasis id="B944EAA53842FF9E29A3FE77FEFBFE2B" bold="true" box="[234,359,409,435]" pageId="80" pageNumber="81">
<typeStatus id="548B88153842FF9E29A3FE77FEFBFE2B" box="[234,359,409,435]" pageId="80" pageNumber="81" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FE54FA83FE4D" blockId="80.[189,1311,409,469]" box="[189,1311,442,469]" pageId="80" pageNumber="81">
<collectingCountry id="F32776273842FF9E29F4FE54FE8AFE4C" box="[189,278,442,468]" name="Japan" pageId="80" pageNumber="81">JAPAN</collectingCountry>
•
<specimenCount id="9D36FD3E3842FF9E2867FE54FEE5FE4D" box="[302,377,442,469]" pageId="80" pageNumber="81" type="generic">17 ex.</specimenCount>
; Shikoku, Ashizuri, Cape Kohchi Pref.;
<date id="FF8E10773842FF9E2A18FE54FB9CFE4C" box="[849,1024,442,469]" pageId="80" pageNumber="81" value="1997-05-01" valueMax="1997-05-03" valueMin="1997-05-01">
<collectingDate id="EFCAE99F3842FF9E2A18FE54FB9CFE4C" box="[849,1024,442,469]" pageId="80" pageNumber="81" value="1997-05-01" valueMax="1997-05-03" valueMin="1997-05-01">1–3 May 1997</collectingDate>
</date>
;
<collectorName id="26C553613842FF9E2D42FE55FBD7FE4D" box="[1035,1099,443,469]" pageId="80" pageNumber="81">T. Ito</collectorName>
leg.;
<collectionCode id="ED21AE723842FF9E2DC2FE54FB2FFE4C" box="[1163,1203,442,468]" pageId="80" pageNumber="81">YS</collectionCode>
,
<collectionCode id="ED21AE723842FF9E2DF6FE55FA87FE4D" box="[1215,1307,443,469]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="80" pageNumber="81">BMNH</collectionCode>
.
</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FE14FD8DFD8C" blockId="80.[189,1398,506,610]" box="[189,529,506,532]" pageId="80" pageNumber="81">
<heading id="D0C781DB3842FF9E29F4FE14FD8DFD8C" bold="true" box="[189,529,506,532]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B944EAA53842FF9E29F4FE14FD8DFD8C" bold="true" box="[189,529,506,532]" pageId="80" pageNumber="81">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FDC8FE36FDFA" blockId="80.[189,1398,506,610]" pageId="80" pageNumber="81">
The following description amplifies the
<typeStatus id="548B88153842FF9E2BDEFDC9FD56FDD9" box="[663,714,551,577]" pageId="80" pageNumber="81">type</typeStatus>
description of
<bibRefCitation id="EFA14B463842FF9E2A35FDC8FBB1FDD9" author="Sawada Y." box="[892,1069,550,577]" pageId="80" pageNumber="81" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada (2008)</bibRefCitation>
to conform with the entries in the present paper.
</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FD69FD36FD3A" blockId="80.[189,682,647,674]" box="[189,682,647,674]" pageId="80" pageNumber="81">
SIZE. Habitus (
<figureCitation id="130B2A323842FF9E282EFD69FE48FD3A" box="[359,468,647,674]" captionStart="Fig" captionStartId="80.[189,232,1964,1990]" captionTargetBox="[191,1395,1046,1920]" captionTargetId="figure@80.[186,1395,1045,1923]" captionTargetPageId="80" captionText="Fig. 51. Cissidium itoi Sawada, 2008. A. Habitus. B. Pronotum, × 375. C. Mesoventrite showing median process of collar, mid keel, keel and lateral margin, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777386" httpUri="https://zenodo.org/record/3777386/files/figure.png" pageId="80" pageNumber="81">Fig. 51A</figureCitation>
), length
<quantity id="4CC89B523842FF9E2B73FD69FD39FD3A" box="[570,677,647,674]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="80" pageNumber="81" unit="mm" value="0.66">0.66 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FD29FCECFD79" blockId="80.[189,880,711,737]" box="[189,880,711,737]" pageId="80" pageNumber="81">COLOUR. Dark brown, antennae, legs and pubescence paler.</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FCE9FB98FCDA" blockId="80.[189,1398,774,834]" pageId="80" pageNumber="81">
HEAD. With a row of indistinctly formed foveolae between the eyes; width across eyes
<quantity id="4CC89B523842FF9E2C48FCE9FAEEFCB9" box="[1281,1394,775,801]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="80" pageNumber="81" unit="mm" value="0.2">0.20 mm</quantity>
; antennomeres III–XI, length
<quantity id="4CC89B523842FF9E2B58FCC6FDE1FCDA" box="[529,637,808,834]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="80" pageNumber="81" unit="mm" value="0.27">0.27 mm</quantity>
, terminal antennomeres globular.
</paragraph>
<paragraph id="8B8F36B73842FF9E29F4FC86FA85FC5F" blockId="80.[189,1398,871,967]" pageId="80" pageNumber="81">
PRONOTUM. Length
<quantity id="4CC89B523842FF9E28D4FC86FD94FC1A" box="[413,520,872,898]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="80" pageNumber="81" unit="mm" value="0.21">0.21 mm</quantity>
, width
<quantity id="4CC89B523842FF9E2B15FC86FD5BFC1A" box="[604,711,872,898]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="80" pageNumber="81" unit="mm" value="0.29">0.29 mm</quantity>
, densely foveolate and pubescent throughout; side margins rounded to slightly concave before acute hind angles, border shallowly crenulate, not continued along the hind margin which has a shallow sinuous emargination in front of the scutellum (
<figureCitation id="130B2A323842FF9E2DE9FC42FA97FC5F" box="[1184,1291,940,967]" captionStart="Fig" captionStartId="80.[189,232,1964,1990]" captionTargetBox="[191,1395,1046,1920]" captionTargetId="figure@80.[186,1395,1045,1923]" captionTargetPageId="80" captionText="Fig. 51. Cissidium itoi Sawada, 2008. A. Habitus. B. Pronotum, × 375. C. Mesoventrite showing median process of collar, mid keel, keel and lateral margin, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777386" httpUri="https://zenodo.org/record/3777386/files/figure.png" pageId="80" pageNumber="81">Fig. 51B</figureCitation>
).
</paragraph>
<caption id="DF4F663F3842FF9E29F4F842FCF4F872" ID-DOI="http://doi.org/10.5281/zenodo.3777386" ID-Zenodo-Dep="3777386" httpUri="https://zenodo.org/record/3777386/files/figure.png" pageId="80" pageNumber="81" startId="80.[189,232,1964,1990]" targetBox="[191,1395,1046,1920]" targetPageId="80">
<paragraph id="8B8F36B73842FF9E29F4F842FCF4F872" blockId="80.[189,1398,1964,2026]" pageId="80" pageNumber="81">
<emphasis id="B944EAA53842FF9E29F4F842FE84F85E" bold="true" box="[189,280,1964,1990]" pageId="80" pageNumber="81">Fig. 51.</emphasis>
<taxonomicName id="4C304D343842FF9E2857F842FDF1F85E" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[286,621,1964,1990]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="B944EAA53842FF9E2857F842FE5CF85E" box="[286,448,1964,1990]" italics="true" pageId="80" pageNumber="81">Cissidium itoi</emphasis>
<bibRefCitation id="EFA14B463842FF9E288FF842FDF1F85E" author="Sawada Y." box="[454,621,1964,1990]" pageId="80" pageNumber="81" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
.
<emphasis id="B944EAA53842FF9E2B3EF842FD10F85E" bold="true" box="[631,652,1964,1990]" pageId="80" pageNumber="81">A</emphasis>
. Habitus.
<emphasis id="B944EAA53842FF9E2A4BF842FC8AF85E" bold="true" box="[770,790,1964,1990]" pageId="80" pageNumber="81">B</emphasis>
. Pronotum, × 375.
<emphasis id="B944EAA53842FF9E2ABAF842FB94F85E" bold="true" box="[1011,1032,1964,1990]" pageId="80" pageNumber="81">C</emphasis>
. Mesoventrite showing median process of collar, mid keel, keel and lateral margin, × 590.
</paragraph>
</caption>
<paragraph id="8B8F36B73843FF9F29F4FEE2FBB6FEBD" blockId="81.[189,1066,267,294]" box="[189,1066,267,294]" pageId="81" pageNumber="82">
ELYTRA. Length
<quantity id="4CC89B523843FF9F2833FEE5FE7AFEBE" box="[378,486,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="81" pageNumber="82" unit="mm" value="0.41">0.41 mm</quantity>
, width
<quantity id="4CC89B523843FF9F2B72FEE5FD3BFEBE" box="[571,679,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="81" pageNumber="82" unit="mm" value="0.32">0.32 mm</quantity>
, pubescent but without foveolae.
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FEBEFEA3FE44" blockId="81.[189,1399,335,476]" pageId="81" pageNumber="82">
MESOVENTRITE. Median extension of collar short; mid-keel and keel united without extensions to the anterior mesocoxal borders, strongly setose and sculpted, terminating in a rounded point between the mesocoxae; humeri shallowly toothed; mesoventral lateral borders smoothly rounded without serrations (
<figureCitation id="130B2A323843FF9F298FFE2FFEAEFE44" box="[198,306,449,476]" captionStart="Fig" captionStartId="80.[189,232,1964,1990]" captionTargetBox="[191,1395,1046,1920]" captionTargetId="figure@80.[186,1395,1045,1923]" captionTargetPageId="80" captionText="Fig. 51. Cissidium itoi Sawada, 2008. A. Habitus. B. Pronotum, × 375. C. Mesoventrite showing median process of collar, mid keel, keel and lateral margin, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777386" httpUri="https://zenodo.org/record/3777386/files/figure.png" pageId="81" pageNumber="82">Fig. 51C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FDE8FCC6FDDE" blockId="81.[189,1398,518,582]" pageId="81" pageNumber="82">
METAVENTRITE. Sparsely pubescent, length
<quantity id="4CC89B523843FF9F2B81FDE8FCA5FDB8" box="[712,825,518,544]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="81" pageNumber="82" unit="mm" value="0.11">0.11 mm</quantity>
, width across spines
<quantity id="4CC89B523843FF9F2D00FDE8FB27FDB8" box="[1097,1211,518,544]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="81" pageNumber="82" unit="mm" value="0.1">0.10 mm</quantity>
, disc foveolate medially; mesocoxal posterior borders without serrations
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FD9EFAEAFD12" blockId="81.[189,1398,624,650]" box="[189,1398,624,650]" pageId="81" pageNumber="82">
GENITALIA. Male aedeagus pointed, not beak-shaped in profile; female spermatheca elongate as
<figureCitation id="130B2A323843FF9F2C5FFD9EFAEFFD12" box="[1302,1395,624,650]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="81" pageNumber="82">Fig. 6D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3843FF9F29F4FD5AFB03FCBD" pageId="81" pageNumber="82" type="discussion">
<paragraph id="8B8F36B73843FF9F29F4FD5AFEB3FD56" blockId="81.[189,303,692,718]" box="[189,303,692,718]" pageId="81" pageNumber="82">
<heading id="D0C781DB3843FF9F29F4FD5AFEB3FD56" bold="true" box="[189,303,692,718]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="B944EAA53843FF9F29F4FD5AFEB3FD56" bold="true" box="[189,303,692,718]" pageId="81" pageNumber="82">Remarks</emphasis>
</heading>
</paragraph>
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This is one of three species in this group from
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and differs from
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<emphasis id="B944EAA53843FF9F2AB3FD08FBFDFD67" box="[1018,1121,741,768]" italics="true" pageId="81" pageNumber="82">C. latum</emphasis>
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Sawada,
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in being narrower and from
<taxonomicName id="4C304D343843FF9F28E9FCE2FD4EFCBE" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[416,722,779,806]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B944EAA53843FF9F28E9FCE2FDB9FCBD" box="[416,549,780,806]" italics="true" pageId="81" pageNumber="82">C. nomurai</emphasis>
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Sawada,
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in the form of the female spermatheca.
</paragraph>
</subSubSection>
</treatment>
</document>
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