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<document id="76BFEC3CCF8F52C6A1E7B7EBD24573D4" ID-CLB-Dataset="3642" ID-DOI="10.11646/zootaxa.5067.1.8" ID-GBIF-Dataset="49bf613b-5fcc-4d38-b4ae-80c9699566f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5656608" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636445647553" checkinUser="plazi" docAuthor="Moctezuma, Victor, Hernández, Benjamín, Sánchez-Huerta, José Luis, Navarrete-Heredia, José Luis &amp; Martínez-Rodríguez, Pablo Antonio" docDate="2021" docId="039958773E70CD7E0AF3F978FA31FE6F" docLanguage="en" docName="zootaxa.5067.1.8.pdf" docOrigin="Zootaxa 5067 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus acernorus Moctezuma, Hernandez, &amp; Sanchez-Huerta 2021, new species" docType="treatment" docVersion="8" lastPageNumber="128" masterDocId="FFA0200F3E70CD780A64FF87FFACFFD1" masterDocTitle="Onthophagus acernorus (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) a new dung beetle species from Jalisco, Mexico" masterLastPageNumber="128" masterPageNumber="122" pageNumber="122" updateTime="1699468065902" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D056AB4BCF136A7A4388F7BD17EF070B">
<mods:title id="97E90A9D54ABA305C4237F34EC05C6C6">Onthophagus acernorus (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) a new dung beetle species from Jalisco, Mexico</mods:title>
</mods:titleInfo>
<mods:name id="9012B7537DD083C152C7F86FF02A244B" type="personal">
<mods:role id="9A5A3E49196434D8374F9087B42F8081">
<mods:roleTerm id="D4C19B72E192B23FCF0A0B4D89FF7A4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4ACBA660AC9ECF0937A3218B95F18912">Moctezuma, Victor</mods:namePart>
<mods:affiliation id="639F06BB0D8FCFE1A7EDE9A891F59BB4">Centro Tlaxcala de Biología de la Conducta, Universidad Autónoma de Tlaxcala. Km 1.5 Carretera Tlaxcala-Puebla S / N, La Loma Xicotencatl, 90070, Tlaxcala, Tlaxcala, Mexico.</mods:affiliation>
</mods:name>
<mods:name id="838256DB79D3C173CF0BF85B57DB41B2" type="personal">
<mods:role id="1D2603C4D920DA8E7C7D2484DB1A828F">
<mods:roleTerm id="BDA04B32FD2F8D43C6167E4116D7CF92">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="73F27189DF296555DAD316D8990D6ABC">Hernández, Benjamín</mods:namePart>
<mods:affiliation id="537FEAC1A8BE75BAB422AC1FC0183563">Departamento de Ciencias Básicas, Instituto Tecnológico de Tlajomulco, Tecnológico Nacional de México. Carretera Tlajomulco-San Miguel Cuyutlán Km. 10, C. P. 45640, Tlajomulco de Zúñiga, Jalisco, Mexico. &amp; Centro de Estudios en Zoología, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara. Apdo. Postal 134, 45100 Zapopan, Jalisco, México.</mods:affiliation>
</mods:name>
<mods:name id="B6D8BAA57F59E201E1152760740D17DC" type="personal">
<mods:role id="56B6C20DBFE803DEB97424F2218C1B11">
<mods:roleTerm id="3F1607D9DA1DC82D60DB6707000EDC0B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2ABC4DD9422C222A24282B5E39CE38AF">Sánchez-Huerta, José Luis</mods:namePart>
<mods:affiliation id="D3AA6AEE0063DEC4E5FAD8E05FD0F447">Calle Enrique Cesar Velarde # 14, Col. María Esther, Xalapa, Veracruz, 91030, Mexico.</mods:affiliation>
</mods:name>
<mods:name id="5534F50C3E24B52781105F55B0A69999" type="personal">
<mods:role id="D2D2BEE6D01D2CFD41AFF0B81CFB0F8F">
<mods:roleTerm id="34D98E7025C6D5FDD97470D4A1012CCE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2AA3A82368357C07B3115397E3F15B0">Navarrete-Heredia, José Luis</mods:namePart>
<mods:affiliation id="90150F42FCED13FB1EB534AD1D745778">Centro de Estudios en Zoología, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara. Apdo. Postal 134, 45100 Zapopan, Jalisco, México.</mods:affiliation>
</mods:name>
<mods:name id="A9F1F222D122708C46A632A3E47482C9" type="personal">
<mods:role id="2C303F997B26620D04CFC86B18427EA7">
<mods:roleTerm id="91EDC5EC26A97AC4637BAB16F7C7EC89">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE31919A79C3952ECCCCB24897E8BD4B">Martínez-Rodríguez, Pablo Antonio</mods:namePart>
<mods:affiliation id="527F7954F07B2C7C83233173ADE28612">Centro de Estudios en Zoología, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara. Apdo. Postal 134, 45100 Zapopan, Jalisco, México.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C657788A472A98C3C00C2A3561253B54">text</mods:typeOfResource>
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<mods:title id="5D0DD3B1FAE268F4B5F9FA7CF8AB1EEA">Zootaxa</mods:title>
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<mods:part id="89A188F41248CB0179AE9A6255AD31AD">
<mods:date id="2E1B0C3F60A3089CCC4280A285E38D01">2021</mods:date>
<mods:detail id="1DCBAF9721F3AF5ED2CF8F8E1D850018" type="pubDate">
<mods:number id="14F4E7F39D8B6B4B03CB0B74B268D73B">2021-11-09</mods:number>
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<mods:detail id="8BA0E25E1FDA0209D98A0E3F02C4A0D5" type="volume">
<mods:number id="E007CFE698E60F0678DFD4F0F5728F8C">5067</mods:number>
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<mods:classification id="309C724FC44CABD44D53DF6B7C32BAE6">journal article</mods:classification>
<mods:identifier id="444ECA93F944C3E6F4AC8C9DBE6209C1" type="CLB-Dataset">3642</mods:identifier>
<mods:identifier id="08B2E8849866F93CE605A47D50A0666E" type="DOI">10.11646/zootaxa.5067.1.8</mods:identifier>
<mods:identifier id="AD698C9D8885BE1627D3C0A8A426503D" type="GBIF-Dataset">49bf613b-5fcc-4d38-b4ae-80c9699566f7</mods:identifier>
<mods:identifier id="4942A26D92DA8098FAD8FB58141ECFDA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039958773E70CD7E0AF3F978FA31FE6F" ID-DOI="http://doi.org/10.5281/zenodo.5693537" ID-GBIF-Taxon="190304901" ID-Zenodo-Dep="5693537" LSID="urn:lsid:plazi:treatment:039958773E70CD7E0AF3F978FA31FE6F" httpUri="http://treatment.plazi.org/id/039958773E70CD7E0AF3F978FA31FE6F" lastPageId="6" lastPageNumber="128" pageId="0" pageNumber="122">
<subSubSection id="C32ABAEA3E70CD780AF3F978FC3FF8ED" pageId="0" pageNumber="122" type="nomenclature">
<paragraph id="8B8FE9613E70CD780AF3F978FABEF8C8" blockId="0.[151,1436,1790,2027]" box="[151,1298,1790,1818]" pageId="0" pageNumber="122">
<heading id="D0C75E0D3E70CD780AF3F978FABEF8C8" bold="true" box="[151,1298,1790,1818]" fontSize="11" level="1" pageId="0" pageNumber="122" reason="1">
<emphasis id="B94435733E70CD780AF3F978FABEF8C8" bold="true" box="[151,1298,1790,1818]" pageId="0" pageNumber="122">
<taxonomicName id="4C3092E23E70CD780AF3F978FC47F8C8" ID-CoL="8SSQS" authority="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[151,1003,1790,1818]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="122" phylum="Arthropoda" rank="species" species="acernorus" status="sp. nov.">
<emphasis id="B94435733E70CD780AF3F978FE11F8CB" bold="true" box="[151,445,1791,1818]" italics="true" pageId="0" pageNumber="122">Onthophagus acernorus</emphasis>
Moctezuma, Hernández, &amp; Sánchez-Huerta
</taxonomicName>
,
<taxonomicNameLabel id="A27788083E70CD78099CF978FB24F8C8" box="[1016,1160,1791,1817]" pageId="0" pageNumber="122" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="130BF5E43E70CD780EF3F978FAA5F8C8" box="[1175,1289,1791,1817]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1943,1967]" captionStartId-1="2.[151,250,1398,1422]" captionStartId-2="3.[151,250,1438,1462]" captionStartId-3="4.[151,250,1069,1093]" captionStartId-4="4.[151,250,1999,2023]" captionTargetBox-0="[188,1398,195,1907]" captionTargetBox-1="[151,1436,181,1361]" captionTargetBox-2="[159,1422,344,1404]" captionTargetBox-3="[231,1352,182,1045]" captionTargetBox-4="[242,1345,1121,1974]" captionTargetId-0="figure-26@1.[160,1428,180,1919]" captionTargetId-1="figure-19@2.[151,1436,181,1375]" captionTargetId-2="figure-97@3.[151,1436,344,1414]" captionTargetId-3="figure-19@4.[224,1364,181,1045]" captionTargetId-4="figure-28@4.[238,1348,1117,1975]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="FIGURE 1. Onthophagus acernorus holotype male (dorsal view)." captionText-1="FIGURE 2. Onthophagus acernorus holotype male (frontal view)." captionText-2="FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite." captionText-3="FIGURE 4. Onthophagus acernorus paratype female." captionText-4="FIGURE 5. Distribution of the Onthophagus chevrolati species complex in central Mexico." figureDoi-0="http://doi.org/10.5281/zenodo.5656610" figureDoi-1="http://doi.org/10.5281/zenodo.5656612" figureDoi-2="http://doi.org/10.5281/zenodo.5656614" figureDoi-3="http://doi.org/10.5281/zenodo.5656616" figureDoi-4="http://doi.org/10.5281/zenodo.5656618" httpUri-0="https://zenodo.org/record/5656610/files/figure.png" httpUri-1="https://zenodo.org/record/5656612/files/figure.png" httpUri-2="https://zenodo.org/record/5656614/files/figure.png" httpUri-3="https://zenodo.org/record/5656616/files/figure.png" httpUri-4="https://zenodo.org/record/5656618/files/figure.png" pageId="0" pageNumber="122">Figs. 1–5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8FE9613E70CD780AF3F8A3FC3FF8ED" blockId="0.[151,1436,1790,2027]" box="[151,915,1828,1852]" pageId="0" pageNumber="122">
<uri id="FFA1E5633E70CD780AF3F8A3FC3FF8ED" box="[151,915,1828,1852]" pageId="0" pageNumber="122">
urn:lsid:zoobank.org:act:
<uuid id="FF96D3B43E70CD780BF9F8A3FC3FF8ED" box="[413,915,1828,1852]" pageId="0" pageNumber="122">3AC33DB0-093F-4B86-8CEA-8A8F0243812</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32ABAEA3E70CD780AF3F8C0FBB4F88F" box="[151,1048,1862,1887]" pageId="0" pageNumber="122" type="materials_examined">
<paragraph id="8B8FE9613E70CD780AF3F8C0FBB4F88F" blockId="0.[151,1436,1790,2027]" box="[151,1048,1862,1887]" pageId="0" pageNumber="122">
<emphasis id="B94435733E70CD780AF3F8C0FE82F88F" bold="true" box="[151,302,1862,1887]" inLineHeading="true" pageId="0" pageNumber="122" reason="1">Type locality.</emphasis>
<materialsCitation id="3B58E33C3E70CD780B50F8C1FBBFF88F" ID-GBIF-Occurrence="3400174304" box="[308,1043,1862,1886]" country="Mexico" latitude="20.214779" location="Jalisco" longLatPrecision="1" longitude="-104.75469" pageId="0" pageNumber="122" specimenCount="1" stateProvince="Jalisco">
<collectingCountry id="F327A9F13E70CD780B50F8C1FE24F88F" box="[308,392,1862,1886]" name="Mexico" pageId="0" pageNumber="122">Mexico</collectingCountry>
,
<collectingRegion id="49F427833E70CD780BF7F8C1FE72F88F" box="[403,478,1862,1886]" country="Mexico" name="Jalisco" pageId="0" pageNumber="122">Jalisco</collectingRegion>
, Talpa de Allende,
<geoCoordinate id="EE048FA63E70CD7808CEF8C1FCFDF88F" box="[682,849,1862,1886]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="0" pageNumber="122" precision="1" seconds="53.20" value="20.214779">20°12’53.20”N</geoCoordinate>
,
<geoCoordinate id="EE048FA63E70CD780938F8C1FBBFF88F" box="[860,1043,1862,1886]" degrees="104" direction="west" minutes="45" orientation="longitude" pageId="0" pageNumber="122" precision="1" seconds="16.88" value="-104.75469">104°45’16.88”W</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32ABAEA3E70CD7A0AA3F8EEFD2AF8B2" lastPageId="2" lastPageNumber="124" pageId="0" pageNumber="122" type="description">
<paragraph id="8B8FE9613E70CD7A0AA3F8EEFA31F992" blockId="0.[151,1436,1790,2027]" lastBlockId="2.[151,1437,1471,2035]" lastPageId="2" lastPageNumber="124" pageId="0" pageNumber="122">
<emphasis id="B94435733E70CD780AA3F8EEFEFEF850" bold="true" box="[199,338,1897,1921]" inLineHeading="true" pageId="0" pageNumber="122" reason="1">Description.</emphasis>
<typeStatus id="548B57C33E70CD780B33F8EEFE16F850" box="[343,442,1897,1921]" pageId="0" pageNumber="122" type="holotype">Holotype</typeStatus>
major male (
<figureCitation id="130BF5E43E70CD780821F8EEFD07F850" box="[581,683,1897,1921]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1943,1967]" captionStartId-1="2.[151,250,1398,1422]" captionStartId-2="3.[151,250,1438,1462]" captionTargetBox-0="[188,1398,195,1907]" captionTargetBox-1="[151,1436,181,1361]" captionTargetBox-2="[159,1422,344,1404]" captionTargetId-0="figure-26@1.[160,1428,180,1919]" captionTargetId-1="figure-19@2.[151,1436,181,1375]" captionTargetId-2="figure-97@3.[151,1436,344,1414]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Onthophagus acernorus holotype male (dorsal view)." captionText-1="FIGURE 2. Onthophagus acernorus holotype male (frontal view)." captionText-2="FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite." figureDoi-0="http://doi.org/10.5281/zenodo.5656610" figureDoi-1="http://doi.org/10.5281/zenodo.5656612" figureDoi-2="http://doi.org/10.5281/zenodo.5656614" httpUri-0="https://zenodo.org/record/5656610/files/figure.png" httpUri-1="https://zenodo.org/record/5656612/files/figure.png" httpUri-2="https://zenodo.org/record/5656614/files/figure.png" pageId="0" pageNumber="122">Figs. 1–3</figureCitation>
). Sericeus black dorsally, without metallic sheen.
<emphasis id="B94435733E70CD780EDDF8EEFB51F850" bold="true" box="[1209,1277,1897,1921]" pageId="0" pageNumber="122">Head.</emphasis>
Dorsal surface densely and superficially punctate, with sparse setae. Clypeus concave, transverse. Clypeogenal suture superficially indicated. Genae rounded. Frons flat. Clypeal carina obsolete. Frontal carina trapezoidal, without tubercles.
<emphasis id="B94435733E70CD780E8AF828FAC4F816" bold="true" box="[1262,1384,1967,1991]" pageId="0" pageNumber="122">Pronotum.</emphasis>
Pronotal punctures coarsely impressed; each puncture bearing a seta. Integument scabriculous; with sparse micropunctures. In dorsal view, anterior protuberance obtusely trapezoidal, medially transverse, and straight. Intermediate tubercles and apex of pronotal prominence aligned. Posterior tubercles situated on anterior half of pronotum. Anterior angles of pronotum acute, lateral angles rounded.
<emphasis id="B94435733E72CD7A0841F980FDD9F9CE" bold="true" box="[549,629,1543,1567]" pageId="2" pageNumber="124">Elytra.</emphasis>
Striae distinctly impressed; with evenly spaced, rounded, superficially impressed punctures. Interstriae surface flat, superficially punctate; each puncture bearing a seta. Integument scabriculous.
</paragraph>
<caption id="DF4FB9E93E71CD790AF3F810FC95F861" ID-DOI="http://doi.org/10.5281/zenodo.5656610" ID-Zenodo-Dep="5656610" box="[151,825,1943,1968]" httpUri="https://zenodo.org/record/5656610/files/figure.png" pageId="1" pageNumber="123" startId="1.[151,250,1943,1967]" targetBox="[188,1398,195,1907]" targetPageId="1">
<paragraph id="8B8FE9613E71CD790AF3F810FC95F861" blockId="1.[151,825,1943,1968]" box="[151,825,1943,1968]" pageId="1" pageNumber="123">
<emphasis id="B94435733E71CD790AF3F810FEBFF87E" bold="true" box="[151,275,1943,1967]" pageId="1" pageNumber="123">FIGURE 1.</emphasis>
<taxonomicName id="4C3092E23E71CD790B7DF81FFDA3F87E" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[281,527,1944,1968]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="123" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E71CD790B7DF81FFDA3F87E" box="[281,527,1944,1968]" italics="true" pageId="1" pageNumber="123">Onthophagus acernorus</emphasis>
</taxonomicName>
holotype male (dorsal view).
</paragraph>
</caption>
<caption id="DF4FB9E93E72CD7A0AF3FAF1FC93FA5E" ID-DOI="http://doi.org/10.5281/zenodo.5656612" ID-Zenodo-Dep="5656612" box="[151,831,1398,1423]" httpUri="https://zenodo.org/record/5656612/files/figure.png" pageId="2" pageNumber="124" startId="2.[151,250,1398,1422]" targetBox="[151,1436,181,1361]" targetPageId="2">
<paragraph id="8B8FE9613E72CD7A0AF3FAF1FC93FA5E" blockId="2.[151,831,1398,1423]" box="[151,831,1398,1423]" pageId="2" pageNumber="124">
<emphasis id="B94435733E72CD7A0AF3FAF1FEBFFA5F" bold="true" box="[151,275,1398,1422]" pageId="2" pageNumber="124">FIGURE 2.</emphasis>
<taxonomicName id="4C3092E23E72CD7A0B7DFAF0FDA3FA5F" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[281,527,1399,1423]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="124" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E72CD7A0B7DFAF0FDA3FA5F" box="[281,527,1399,1423]" italics="true" pageId="2" pageNumber="124">Onthophagus acernorus</emphasis>
</taxonomicName>
holotype male (frontal view).
</paragraph>
</caption>
<paragraph id="8B8FE9613E72CD7A0AF3F9C9FD24F97E" blockId="2.[151,1437,1471,2035]" pageId="2" pageNumber="124">
<emphasis id="B94435733E72CD7A0AF3F9C9FE84F9B6" bold="true" box="[151,296,1614,1639]" pageId="2" pageNumber="124">Tergite VIII.</emphasis>
Surface densely and superficially punctate, each puncture bearing a seta. Integument scabriculous.
<emphasis id="B94435733E72CD7A0F04F9C8FA30F9B6" bold="true" box="[1376,1436,1615,1639]" pageId="2" pageNumber="124">Legs.</emphasis>
Protibiae slender and quadridentate; inwardly bent on apical third. Protibial apical spur obliquely curved ventrally.
<emphasis id="B94435733E72CD7A0F39F9F5FF7DF97F" bold="true" pageId="2" pageNumber="124">Genitalia.</emphasis>
Apex of parameres obtusely trapezoidal.
</paragraph>
<paragraph id="8B8FE9613E72CD7A0AA3F93DFE67F902" blockId="2.[151,1437,1471,2035]" box="[199,459,1722,1747]" pageId="2" pageNumber="124">
<emphasis id="B94435733E72CD7A0AA3F93DFEF8F902" bold="true" box="[199,340,1722,1747]" inLineHeading="true" pageId="2" pageNumber="124" reason="1">Minor male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8FE9613E72CD7A0AA3F958FE29F8EE" blockId="2.[151,1437,1471,2035]" pageId="2" pageNumber="124">
<emphasis id="B94435733E72CD7A0AA3F958FE8DF926" bold="true" box="[199,289,1759,1783]" inLineHeading="true" pageId="2" pageNumber="124" reason="1">Female.</emphasis>
Similar to the male except for the clypeus elongate; clypeal carina distinctly developed; frontal carina rounded; pronotal protuberance distinctly reduced; posterior tubercles located in the anterior portion of pronotum; protibia widened (
<figureCitation id="130BF5E43E72CD7A0B52F8A0FED4F8EE" box="[310,376,1831,1855]" captionStart="FIGURE 4" captionStartId="4.[151,250,1069,1093]" captionTargetBox="[231,1352,182,1045]" captionTargetId="figure-19@4.[224,1364,181,1045]" captionTargetPageId="4" captionText="FIGURE 4. Onthophagus acernorus paratype female." figureDoi="http://doi.org/10.5281/zenodo.5656616" httpUri="https://zenodo.org/record/5656616/files/figure.png" pageId="2" pageNumber="124">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B8FE9613E72CD7A0AA3F8CDFD2AF8B2" blockId="2.[151,1437,1471,2035]" box="[199,646,1866,1891]" pageId="2" pageNumber="124">
<emphasis id="B94435733E72CD7A0AA3F8CDFE96F8B3" bold="true" box="[199,314,1866,1890]" inLineHeading="true" pageId="2" pageNumber="124" reason="1">Variation.</emphasis>
Mean length 9.2 (
<quantity id="4CC844843E72CD7A0B98F8CCFDD6F8B2" box="[508,634,1867,1891]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" metricValueMax="11.0" metricValueMin="6.6" pageId="2" pageNumber="124" unit="mm" value="8.8" valueMax="11.0" valueMin="6.6">6.6–11 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C32ABAEA3E72CD7B0AA3F8E8FB2BFE91" lastPageId="3" lastPageNumber="125" pageId="2" pageNumber="124" type="materials_examined">
<paragraph id="8B8FE9613E72CD7A0AA3F8E8FC30F81E" blockId="2.[151,1437,1471,2035]" pageId="2" pageNumber="124">
<emphasis id="B94435733E72CD7A0AA3F8E8FE48F857" bold="true" box="[199,484,1902,1927]" inLineHeading="true" pageId="2" pageNumber="124" reason="1">Type material examined.</emphasis>
<materialsCitation id="3B58E33C3E72CD7A0B8FF8E8FC30F81E" ID-GBIF-Occurrence="3400174301" collectingDate="2016-03-03" collectionCode="CZUG" collectorName="Arce. P. A. Martinez-Rodriguez &amp; J. L. Navarrete-Heredia Col." country="Mexico" elevation="1786" latitude="20.214779" location="Bosque Mesofilo de Montana" longLatPrecision="1" longitude="-104.75469" municipality="Bosque" pageId="2" pageNumber="124" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="548B57C33E72CD7A0B8FF8E8FDE2F856" box="[491,590,1903,1927]" pageId="2" pageNumber="124" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B94435733E72CD7A08C8F8E9FC2DF857" bold="true" box="[684,897,1902,1926]" pageId="2" pageNumber="124">
<collectingCountry id="F327A9F13E72CD7A08C8F8E9FCB3F857" box="[684,799,1902,1926]" name="Mexico" pageId="2" pageNumber="124">MEXICO</collectingCountry>
.
<collectingRegion id="49F427833E72CD7A094FF8E9FCD0F857" box="[811,892,1902,1926]" country="Mexico" name="Jalisco" pageId="2" pageNumber="124">Jalisco</collectingRegion>
:
</emphasis>
“Talpa de Allende:
<collectingMunicipality id="6BEB731B3E72CD7A0E3FF8E8FB07F857" box="[1115,1195,1903,1926]" pageId="2" pageNumber="124">Bosque</collectingMunicipality>
estatal de Arce, NTP-80-6-calamar, 30-I/
<date id="FF8ECFA13E72CD7A0B01F814FE49F87A" box="[357,485,1939,1963]" pageId="2" pageNumber="124" value="2016-03-03">
<collectingDate id="EFCA36493E72CD7A0B01F814FE49F87A" box="[357,485,1939,1963]" pageId="2" pageNumber="124" value="2016-03-03">03-III-2016</collectingDate>
</date>
,
<geoCoordinate id="EE048FA63E72CD7A0B8BF815FD39F87B" box="[495,661,1938,1962]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="124" precision="1" seconds="53.20" value="20.214779">20°12’53.20”N</geoCoordinate>
,
<geoCoordinate id="EE048FA63E72CD7A08C4F815FCF4F87B" box="[672,856,1938,1962]" degrees="104" direction="west" minutes="45" orientation="longitude" pageId="2" pageNumber="124" precision="1" seconds="16.88" value="-104.75469">104°45’16.88”W</geoCoordinate>
,
<quantity id="4CC844843E72CD7A0907F814FC1AF87B" box="[867,950,1939,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.786" pageId="2" pageNumber="124" unit="m" value="1786.0">
<elevation id="001D0E523E72CD7A0907F814FC1AF87B" box="[867,950,1939,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.786" pageId="2" pageNumber="124" unit="m" value="1786.0">1786 m</elevation>
</quantity>
,
<location id="8EEFBFBA3E72CD7A09A5F814FB51F87B" LSID="urn:lsid:plazi:treatment:039958773E70CD7E0AF3F978FA31FE6F:8EEFBFBA3E72CD7A09A5F814FB51F87B" box="[961,1277,1939,1963]" country="Mexico" latitude="20.214779" longLatPrecision="1" longitude="-104.75469" municipality="Bosque" name="Bosque Mesofilo de Montana" pageId="2" pageNumber="124" stateProvince="Jalisco">Bosque Mesófilo de Montaña</location>
con bosque de
<collectorName id="26C58CB73E72CD7A0AF3F830FE4FF81E" box="[151,483,1975,1999]" pageId="2" pageNumber="124">Arce. P.A. Martínez-Rodríguez</collectorName>
y
<collectorName id="26C58CB73E72CD7A0B9AF830FC85F81E" box="[510,809,1975,1999]" pageId="2" pageNumber="124">J.L. Navarrete-Heredia Col.</collectorName>
” (
<collectionCode id="ED2171A43E72CD7A0925F830FC3CF81E" box="[833,912,1975,1999]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="2" pageNumber="124">CZUG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8FE9613E72CD7B0AA3F85CFB2BFE91" blockId="2.[151,1437,1471,2035]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="125" pageId="2" pageNumber="124">
<typeStatus id="548B57C33E72CD7A0AA3F85CFE83F822" box="[199,303,2011,2035]" pageId="2" pageNumber="124" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D3622E83E72CD7A0B24F85CFE3BF822" box="[320,407,2011,2035]" pageId="2" pageNumber="124" type="male">3 males</specimenCount>
,
<specimenCount id="9D3622E83E72CD7A0BC0F85CFDB5F822" box="[420,537,2011,2035]" pageId="2" pageNumber="124" type="female">11 females</specimenCount>
):
<materialsCitation id="3B58E33C3E72CD7B0827F85DFECFFF61" ID-GBIF-Occurrence="3400174305" collectingDate="2016-03-03" collectingDateMax="2016-04-08" collectingDateMin="2016-03-03" collectionCode="CZUG, CMNC, VMC" collectorName="Arce. P. A. Martinez-Rodriguez &amp; J. L. Navarrete-Heredia Col." country="Mexico" county="Bosque" elevation="1750" lastPageId="3" lastPageNumber="125" latitude="20.216272" location="Bosque Mesofilo de Montana" longLatPrecision="1" longitude="-104.75589" municipality="Bosque Mesofilo de Montana" pageId="2" pageNumber="124" specimenCount="14" specimenCount-female="11" specimenCount-male="3" stateProvince="Jalisco" typeStatus="paratype">
<emphasis id="B94435733E72CD7A0827F85DFCB4F823" bold="true" box="[579,792,2010,2034]" pageId="2" pageNumber="124">
<collectingCountry id="F327A9F13E72CD7A0827F85DFD1AF823" box="[579,694,2010,2034]" name="Mexico" pageId="2" pageNumber="124">MEXICO</collectingCountry>
.
<collectingRegion id="49F427833E72CD7A08A6F85DFCBFF823" box="[706,787,2010,2034]" country="Mexico" name="Jalisco" pageId="2" pageNumber="124">Jalisco</collectingRegion>
:
</emphasis>
same data as holotype (
<specimenCount id="9D3622E83E72CD7A0E40F85CFBC2F822" box="[1060,1134,2011,2035]" pageId="2" pageNumber="124" type="male">1 male</specimenCount>
,
<specimenCount id="9D3622E83E72CD7A0E1FF85CFB4BF822" box="[1147,1255,2011,2035]" pageId="2" pageNumber="124" type="female">4 females</specimenCount>
,
<collectionCode id="ED2171A43E72CD7A0E97F85CFA93F822" box="[1267,1343,2011,2035]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="2" pageNumber="124">CZUG</collectionCode>
;
<specimenCount id="9D3622E83E72CD7A0F28F85CFA34F822" box="[1356,1432,2011,2035]" pageId="2" pageNumber="124" type="male">1 male</specimenCount>
,
<specimenCount id="9D3622E83E73CD7B0AF3FF1FFF54FF7E" box="[151,248,151,176]" pageId="3" pageNumber="125" type="female">1 female</specimenCount>
,
<collectionCode id="ED2171A43E73CD7B0B60FF1FFEFBFF61" box="[260,343,152,176]" pageId="3" pageNumber="125">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B58E33C3E73CD7B0B0FFF10FCD5FF29" ID-GBIF-Occurrence="3400174303" collectingDate="2016-03-03" collectingDateMax="2016-04-08" collectingDateMin="2016-03-03" collectionCode="CZUG, CMNC, VMC" collectorName="Arce. P. A. Martinez-Rodriguez &amp; J. L. Navarrete-Heredia Col." country="Mexico" county="Bosque" elevation="1750" latitude="20.216272" location="Bosque Mesofilo de Montana" longLatPrecision="1" longitude="-104.75589" municipality="Bosque Mesofilo de Montana" pageId="3" pageNumber="125" specimenCount="14" specimenCount-female="11" specimenCount-male="3" stateProvince="Jalisco" typeStatus="paratype">
“
<collectingRegion id="49F427833E73CD7B0B11FF10FE60FF7E" box="[373,460,151,175]" country="Mexico" name="Mexico" pageId="3" pageNumber="125">México</collectingRegion>
:
<collectingRegion id="49F427833E73CD7B0BBCFF10FD89FF7E" box="[472,549,151,175]" country="Mexico" name="Jalisco" pageId="3" pageNumber="125">Jalisco</collectingRegion>
: Talpa de Allende:
<collectingCounty id="62EE91ED3E73CD7B0892FF1FFCEAFF7E" box="[758,838,152,175]" pageId="3" pageNumber="125">Bosque</collectingCounty>
estatal de Arce, NTP-80-5-calamar, 03-III/
<date id="FF8ECFA13E73CD7B0F7EFF10FA34FF7E" box="[1306,1432,151,175]" pageId="3" pageNumber="125" value="2016-04-08">
<collectingDate id="EFCA36493E73CD7B0F7EFF10FA34FF7E" box="[1306,1432,151,175]" pageId="3" pageNumber="125" value="2016-04-08">08-IV-2016</collectingDate>
</date>
,
<geoCoordinate id="EE048FA63E73CD7B0AF3FF3CFE91FF02" box="[151,317,187,211]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="125" precision="1" seconds="58.58" value="20.216272">20°12’58.58”N</geoCoordinate>
,
<geoCoordinate id="EE048FA63E73CD7B0B2DFF3CFDACFF02" box="[329,512,187,211]" degrees="104" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="125" precision="1" seconds="21.21" value="-104.75589">104°45’21.21”W</geoCoordinate>
,
<quantity id="4CC844843E73CD7B0868FF3CFDF3FF02" box="[524,607,187,211]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="125" unit="m" value="1750.0">
<elevation id="001D0E523E73CD7B0868FF3CFDF3FF02" box="[524,607,187,211]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="125" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<collectingMunicipality id="6BEB731B3E73CD7B080FFF3BFC05FF02" box="[619,937,187,211]" pageId="3" pageNumber="125">Bosque Mesófilo de Montaña</collectingMunicipality>
con bosque de
<collectorName id="26C58CB73E73CD7B0E2BFF3BFA30FF02" box="[1103,1436,187,212]" pageId="3" pageNumber="125">Arce. P.A. Martínez-Rodríguez</collectorName>
y
<collectorName id="26C58CB73E73CD7B0ACFFF67FE79FF26" box="[171,469,223,247]" pageId="3" pageNumber="125">J.L. Navarrete-Heredia Col.</collectorName>
” (
<specimenCount id="9D3622E83E73CD7B0B89FF67FDF4FF26" box="[493,600,223,248]" pageId="3" pageNumber="125" type="female">4 females</specimenCount>
,
<collectionCode id="ED2171A43E73CD7B0806FF67FD02FF29" box="[610,686,224,248]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="3" pageNumber="125">CZUG</collectionCode>
;
<specimenCount id="9D3622E83E73CD7B08DCFF67FC8EFF26" box="[696,802,223,248]" pageId="3" pageNumber="125" type="female">2 females</specimenCount>
,
<collectionCode id="ED2171A43E73CD7B0948FF67FCC1FF29" box="[812,877,224,248]" pageId="3" pageNumber="125">VMC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B58E33C3E73CD7B091BFF58FB2BFE91" ID-GBIF-Occurrence="3400174302" collectingDate="2016-03-03" collectingDateMax="2016-04-08" collectingDateMin="2016-03-03" collectionCode="CZUG, CMNC, VMC" collectorName="Arce. P. A. Martinez-Rodriguez &amp; J. L. Navarrete-Heredia Col." country="Mexico" county="Bosque" elevation="1750" latitude="20.216272" location="Bosque Mesofilo de Montana" longLatPrecision="1" longitude="-104.75589" municipality="Bosque Mesofilo de Montana" pageId="3" pageNumber="125" specimenCount="14" specimenCount-female="11" specimenCount-male="3" stateProvince="Jalisco" typeStatus="paratype">
“
<collectingRegion id="49F427833E73CD7B09EDFF58FC4CFF26" box="[905,992,223,247]" country="Mexico" name="Mexico" pageId="3" pageNumber="125">México</collectingRegion>
:
<collectingRegion id="49F427833E73CD7B098EFF58FB9BFF26" box="[1002,1079,223,247]" country="Mexico" name="Jalisco" pageId="3" pageNumber="125">Jalisco</collectingRegion>
: Talpa de Allende: Bosque estatal de Arce, NTP-80-4-calamar, 30-I/
<date id="FF8ECFA13E73CD7B0865FE84FD2DFECA" box="[513,641,259,283]" pageId="3" pageNumber="125" value="2016-03-03">
<collectingDate id="EFCA36493E73CD7B0865FE84FD2DFECA" box="[513,641,259,283]" pageId="3" pageNumber="125" value="2016-03-03">03-III-2016</collectingDate>
</date>
,
<geoCoordinate id="EE048FA63E73CD7B08E8FE84FC9EFECA" box="[652,818,259,283]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="125" precision="1" seconds="58.58" value="20.216272">20°12’58.58”N</geoCoordinate>
,
<geoCoordinate id="EE048FA63E73CD7B0959FE84FC59FECA" box="[829,1013,259,283]" degrees="104" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="125" precision="1" seconds="21.21" value="-104.75589">104°45’21.21”W</geoCoordinate>
,
<quantity id="4CC844843E73CD7B0E64FE84FBFFFECA" box="[1024,1107,259,283]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="125" unit="m" value="1750.0">
<elevation id="001D0E523E73CD7B0E64FE84FBFFFECA" box="[1024,1107,259,283]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="125" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<location id="8EEFBFBA3E73CD7B0E3BFE83FA30FECA" LSID="urn:lsid:plazi:treatment:039958773E70CD7E0AF3F978FA31FE6F:8EEFBFBA3E73CD7B0E3BFE83FA30FECA" box="[1119,1436,259,283]" country="Mexico" county="Bosque" latitude="20.216272" longLatPrecision="1" longitude="-104.75589" municipality="Bosque Mesofilo de Montana" name="Bosque Mesofilo de Montana" pageId="3" pageNumber="125" stateProvince="Jalisco">Bosque Mesófilo de Montaña</location>
con bosque de
<collectorName id="26C58CB73E73CD7B0B51FEAFFD2DFEEE" box="[309,641,295,320]" pageId="3" pageNumber="125">Arce. P.A. Martínez-Rodríguez</collectorName>
y
<collectorName id="26C58CB73E73CD7B08F8FEAFFC6BFEEE" box="[668,967,295,319]" pageId="3" pageNumber="125">J.L. Navarrete-Heredia Col.</collectorName>
” (
<specimenCount id="9D3622E83E73CD7B0986FEAFFB86FEEE" box="[994,1066,295,320]" pageId="3" pageNumber="125" type="male">1 male</specimenCount>
,
<collectionCode id="ED2171A43E73CD7B0E52FEAFFB2CFE91" box="[1078,1152,296,320]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="3" pageNumber="125">CZUG</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF4FB9E93E73CD7B0AF3FA19FBD9FA0B" ID-DOI="http://doi.org/10.5281/zenodo.5656614" ID-Zenodo-Dep="5656614" httpUri="https://zenodo.org/record/5656614/files/figure.png" pageId="3" pageNumber="125" startId="3.[151,250,1438,1462]" targetBox="[159,1422,344,1404]" targetPageId="3">
<paragraph id="8B8FE9613E73CD7B0AF3FA19FBD9FA0B" blockId="3.[151,1437,1438,1498]" pageId="3" pageNumber="125">
<emphasis id="B94435733E73CD7B0AF3FA19FEB8FA67" bold="true" box="[151,276,1438,1462]" pageId="3" pageNumber="125">FIGURE 3.</emphasis>
<taxonomicName id="4C3092E23E73CD7B0B78FA19FDBFFA67" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[284,531,1438,1462]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E73CD7B0B78FA19FDBFFA67" box="[284,531,1438,1462]" italics="true" pageId="3" pageNumber="125">Onthophagus acernorus</emphasis>
</taxonomicName>
holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite.
<taxonomicName id="4C3092E23E73CD7B0AF3FA45FE20FA0B" authorityName="Harold" authorityYear="1869" box="[151,396,1474,1498]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E73CD7B0AF3FA45FE20FA0B" box="[151,396,1474,1498]" italics="true" pageId="3" pageNumber="125">Onthophagus chevrolati</emphasis>
</taxonomicName>
: C, aedeagus; D, medial endophallite and additional medial endophallite.
</paragraph>
</caption>
<subSubSection id="C32ABAEA3E73CD7B0AA3F98DFD65F997" pageId="3" pageNumber="125" type="distribution">
<paragraph id="8B8FE9613E73CD7B0AA3F98DFD65F997" blockId="3.[151,1437,1546,1930]" pageId="3" pageNumber="125">
<emphasis id="B94435733E73CD7B0AA3F98DFE49F9F3" bold="true" box="[199,485,1546,1570]" inLineHeading="true" pageId="3" pageNumber="125" reason="1">Distribution and ecology.</emphasis>
The new species is found in the Jaliscan district subprovince, Sierra Madre del Sur province (
<emphasis id="B94435733E73CD7B0AB7F9A8FEA2F997" box="[211,270,1583,1606]" italics="true" pageId="3" pageNumber="125">sensu</emphasis>
<bibRefCitation id="EFA194903E73CD7B0B71F9A9FE1EF997" author="Morrone, J. J." box="[277,434,1582,1606]" pageId="3" pageNumber="125" pagination="710 - 714" refId="ref2200" refString="Morrone, J. J. (2017) Biogeographic regionalization of the Sierra Madre del Sur province, Mexico. Revista Mexicana de Biodiversidad, 88, 710 - 714. https: // doi. org / 10.1016 / j. rmb. 2017.07.012" type="journal article" year="2017">Morrone 2017</bibRefCitation>
),
<collectingRegion id="49F427833E73CD7B0BA1F9A9FDBCF997" box="[453,528,1582,1606]" country="Mexico" name="Jalisco" pageId="3" pageNumber="125">Jalisco</collectingRegion>
,
<collectingCountry id="F327A9F13E73CD7B087FF9A9FDC0F997" box="[539,620,1582,1606]" name="Mexico" pageId="3" pageNumber="125">Mexico</collectingCountry>
(
<figureCitation id="130BF5E43E73CD7B081FF9A9FD17F997" box="[635,699,1582,1606]" captionStart="FIGURE 5" captionStartId="4.[151,250,1999,2023]" captionTargetBox="[242,1345,1121,1974]" captionTargetId="figure-28@4.[238,1348,1117,1975]" captionTargetPageId="4" captionText="FIGURE 5. Distribution of the Onthophagus chevrolati species complex in central Mexico." figureDoi="http://doi.org/10.5281/zenodo.5656618" httpUri="https://zenodo.org/record/5656618/files/figure.png" pageId="3" pageNumber="125">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32ABAEA3E73CD7B0AA3F9D5FC7AF95F" pageId="3" pageNumber="125" type="etymology">
<paragraph id="8B8FE9613E73CD7B0AA3F9D5FC7AF95F" blockId="3.[151,1437,1546,1930]" pageId="3" pageNumber="125">
<emphasis id="B94435733E73CD7B0AA3F9D5FEE4F9BB" bold="true" box="[199,328,1618,1642]" inLineHeading="true" pageId="3" pageNumber="125" reason="1">Etymology.</emphasis>
The specific epithet is an adjective in the nominative singular referring to the maple forest where the species occurs. The forest is dominated by
<taxonomicName id="4C3092E23E73CD7B0806F9F1FC7AF95F" authority="Vargas-Rodriguez." authorityName="Vargas-Rodriguez." box="[610,982,1654,1678]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="125" phylum="Tracheophyta" rank="species" species="binzayedii">
<emphasis id="B94435733E73CD7B0806F9F1FCA5F95F" box="[610,777,1654,1678]" italics="true" pageId="3" pageNumber="125">Acer binzayedii</emphasis>
Vargas-Rodriguez.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32ABAEA3E73CD7E0AA3F91DFA31FE6F" lastPageId="6" lastPageNumber="128" pageId="3" pageNumber="125" type="diagnosis">
<paragraph id="8B8FE9613E73CD7B0AA3F91DFA30F85B" blockId="3.[151,1437,1546,1930]" pageId="3" pageNumber="125">
<emphasis id="B94435733E73CD7B0AA3F91DFE90F963" bold="true" box="[199,316,1690,1714]" inLineHeading="true" pageId="3" pageNumber="125" reason="1">Diagnosis.</emphasis>
Major males of
<taxonomicName id="4C3092E23E73CD7B0B93F91DFD2BF963" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[503,647,1690,1714]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E73CD7B0B93F91DFD2BF963" box="[503,647,1690,1714]" italics="true" pageId="3" pageNumber="125">O. acernorus</emphasis>
</taxonomicName>
are easily distinguished from the members of the
<taxonomicName id="4C3092E23E73CD7B0ED3F91DFAEAF963" authorityName="Harold" authorityYear="1869" box="[1207,1350,1690,1714]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E73CD7B0ED3F91DFAEAF963" box="[1207,1350,1690,1714]" italics="true" pageId="3" pageNumber="125">O. chevrolati</emphasis>
</taxonomicName>
species complex by the intermediate tubercles almost aligned with the apex of the pronotal protuberance (intermediate tubercles situated posterolaterally in related species,
<figureCitation id="130BF5E43E73CD7B0802F965FD03F92B" box="[614,687,1762,1786]" captionStart="FIGURE 1" captionStartId="1.[151,250,1943,1967]" captionTargetBox="[188,1398,195,1907]" captionTargetId="figure-26@1.[160,1428,180,1919]" captionTargetPageId="1" captionText="FIGURE 1. Onthophagus acernorus holotype male (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5656610" httpUri="https://zenodo.org/record/5656610/files/figure.png" pageId="3" pageNumber="125">Figs. 1</figureCitation>
,
<figureCitation id="130BF5E43E73CD7B08D9F965FD61F92B" box="[701,717,1762,1786]" captionStart="FIGURE 6" captionStartId="5.[151,255,1943,1967]" captionTargetBox="[232,1359,200,1896]" captionTargetId="figure-27@5.[214,1373,180,1919]" captionTargetPageId="5" captionText="FIGURE 6. Onthophagus chevrolati male (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5656620" httpUri="https://zenodo.org/record/5656620/files/figure.png" pageId="3" pageNumber="125">6</figureCitation>
).
<taxonomicName id="4C3092E23E73CD7B0885F965FC48F92B" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[737,996,1762,1786]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E73CD7B0885F965FC48F92B" box="[737,996,1762,1786]" italics="true" pageId="3" pageNumber="125">Onthophagus acernorus</emphasis>
</taxonomicName>
(
<figureCitation id="130BF5E43E73CD7B0990F965FB9EF92B" box="[1012,1074,1762,1786]" captionStart="FIGURE 1" captionStartId="1.[151,250,1943,1967]" captionTargetBox="[188,1398,195,1907]" captionTargetId="figure-26@1.[160,1428,180,1919]" captionTargetPageId="1" captionText="FIGURE 1. Onthophagus acernorus holotype male (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5656610" httpUri="https://zenodo.org/record/5656610/files/figure.png" pageId="3" pageNumber="125">Fig. 1</figureCitation>
) is morphologically similar to
<taxonomicName id="4C3092E23E73CD7B0FE6F965FEAEF8CF" authorityName="Harold" authorityYear="1869" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E73CD7B0FE6F965FEAEF8CF" italics="true" pageId="3" pageNumber="125">O. chevrolati</emphasis>
</taxonomicName>
(
<figureCitation id="130BF5E43E73CD7B0B76F881FEF8F8CF" box="[274,340,1798,1822]" captionStart="FIGURE 6" captionStartId="5.[151,255,1943,1967]" captionTargetBox="[232,1359,200,1896]" captionTargetId="figure-27@5.[214,1373,180,1919]" captionTargetPageId="5" captionText="FIGURE 6. Onthophagus chevrolati male (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5656620" httpUri="https://zenodo.org/record/5656620/files/figure.png" pageId="3" pageNumber="125">Fig. 6</figureCitation>
), but it can be distinguished by the pronotum more coarsely and densely punctate and less pubescent. Moreover,
<taxonomicName id="4C3092E23E73CD7B0B6FF8ADFE3BF893" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[267,407,1834,1858]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E73CD7B0B6FF8ADFE3BF893" box="[267,407,1834,1858]" italics="true" pageId="3" pageNumber="125">O. acernorus</emphasis>
</taxonomicName>
is completely black dorsally, while specimens of
<taxonomicName id="4C3092E23E73CD7B09FBF8ADFB85F893" authorityName="Harold" authorityYear="1869" box="[927,1065,1834,1858]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E73CD7B09FBF8ADFB85F893" box="[927,1065,1834,1858]" italics="true" pageId="3" pageNumber="125">O. chevrolati</emphasis>
</taxonomicName>
are typically brown or dark brown, and with or without green metallic sheen when observed under the microscope. The apical teeth of parameres are more acute and the inferior right lobe of the medial endophallite is more strongly developed and wavy in
<taxonomicName id="4C3092E23E73CD7B0ED1F8F5FAECF85B" authorityName="Harold" authorityYear="1869" box="[1205,1344,1906,1930]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E73CD7B0ED1F8F5FAECF85B" box="[1205,1344,1906,1930]" italics="true" pageId="3" pageNumber="125">O. chevrolati</emphasis>
</taxonomicName>
(
<figureCitation id="130BF5E43E73CD7B0F2AF8F5FA22F85B" box="[1358,1422,1906,1930]" captionStart="FIGURE 3" captionStartId="3.[151,250,1438,1462]" captionTargetBox="[159,1422,344,1404]" captionTargetId="figure-97@3.[151,1436,344,1414]" captionTargetPageId="3" captionText="FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite." figureDoi="http://doi.org/10.5281/zenodo.5656614" httpUri="https://zenodo.org/record/5656614/files/figure.png" pageId="3" pageNumber="125">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF4FB9E93E74CD7C0AF3FBAAFD13FB94" ID-DOI="http://doi.org/10.5281/zenodo.5656616" ID-Zenodo-Dep="5656616" box="[151,703,1069,1093]" httpUri="https://zenodo.org/record/5656616/files/figure.png" pageId="4" pageNumber="126" startId="4.[151,250,1069,1093]" targetBox="[231,1352,182,1045]" targetPageId="4">
<paragraph id="8B8FE9613E74CD7C0AF3FBAAFD13FB94" blockId="4.[151,703,1069,1093]" box="[151,703,1069,1093]" pageId="4" pageNumber="126">
<emphasis id="B94435733E74CD7C0AF3FBAAFEBFFB94" bold="true" box="[151,275,1069,1093]" pageId="4" pageNumber="126">FIGURE 4.</emphasis>
<taxonomicName id="4C3092E23E74CD7C0B7DFBAAFDA3FB94" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[281,527,1069,1093]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E74CD7C0B7DFBAAFDA3FB94" box="[281,527,1069,1093]" italics="true" pageId="4" pageNumber="126">Onthophagus acernorus</emphasis>
</taxonomicName>
paratype female.
</paragraph>
</caption>
<caption id="DF4FB9E93E74CD7C0AF3F848FB91F836" ID-DOI="http://doi.org/10.5281/zenodo.5656618" ID-Zenodo-Dep="5656618" box="[151,1085,1999,2023]" httpUri="https://zenodo.org/record/5656618/files/figure.png" pageId="4" pageNumber="126" startId="4.[151,250,1999,2023]" targetBox="[242,1345,1121,1974]" targetPageId="4">
<paragraph id="8B8FE9613E74CD7C0AF3F848FB91F836" blockId="4.[151,1085,1999,2023]" box="[151,1085,1999,2023]" pageId="4" pageNumber="126">
<emphasis id="B94435733E74CD7C0AF3F848FEBFF836" bold="true" box="[151,275,1999,2023]" pageId="4" pageNumber="126">FIGURE 5.</emphasis>
Distribution of the
<taxonomicName id="4C3092E23E74CD7C0BBFF848FD7CF836" authorityName="Harold" authorityYear="1869" box="[475,720,1999,2023]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E74CD7C0BBFF848FD7CF836" box="[475,720,1999,2023]" italics="true" pageId="4" pageNumber="126">Onthophagus chevrolati</emphasis>
</taxonomicName>
species complex in central Mexico.
</paragraph>
</caption>
<caption id="DF4FB9E93E75CD7D0AF3F810FD54F861" ID-DOI="http://doi.org/10.5281/zenodo.5656620" ID-Zenodo-Dep="5656620" box="[151,760,1943,1968]" httpUri="https://zenodo.org/record/5656620/files/figure.png" pageId="5" pageNumber="127" startId="5.[151,255,1943,1967]" targetBox="[232,1359,200,1896]" targetPageId="5">
<paragraph id="8B8FE9613E75CD7D0AF3F810FD54F861" blockId="5.[151,760,1943,1968]" box="[151,760,1943,1968]" pageId="5" pageNumber="127">
<emphasis id="B94435733E75CD7D0AF3F810FEB5F87E" bold="true" box="[151,281,1943,1967]" pageId="5" pageNumber="127">FIGURE 6.</emphasis>
<taxonomicName id="4C3092E23E75CD7D0B44F81FFD8EF861" authorityName="Harold" authorityYear="1869" box="[288,546,1944,1968]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E75CD7D0B44F81FFD8EF861" box="[288,546,1944,1968]" italics="true" pageId="5" pageNumber="127">Onthophagus chevrolati</emphasis>
</taxonomicName>
male (dorsal view).
</paragraph>
</caption>
<paragraph id="8B8FE9613E76CD7E0AF3FF10FD58FF05" blockId="6.[151,1437,151,446]" pageId="6" pageNumber="128">
The following addendum to the
<taxonomicName id="4C3092E23E76CD7E0B8CFF10FD44FF7E" authorityName="Harold" authorityYear="1869" box="[488,744,151,175]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E76CD7E0B8CFF10FD44FF7E" box="[488,744,151,175]" italics="true" pageId="6" pageNumber="128">Onthophagus chevrolati</emphasis>
</taxonomicName>
species group identification key will accommodate
<taxonomicName id="4C3092E23E76CD7E0F69FF10FEAFFF02" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="species" species="acernorus">
<emphasis id="B94435733E76CD7E0F69FF10FEAFFF02" italics="true" pageId="6" pageNumber="128">Onthophagus acernorus</emphasis>
</taxonomicName>
(modified from
<bibRefCitation id="EFA194903E76CD7E0BD0FF3BFD45FF05" author="Moctezuma, V. &amp; Halffter, G." box="[436,745,187,212]" pageId="6" pageNumber="128" pagination="245 - 261" refId="ref2149" refString="Moctezuma, V. &amp; Halffter, G. (2020) New species and redescriptions of the Onthophagus chevrolati species complex (Coleoptera: Scarabaeoidea: Scarabaeinae). Annales Zoologici, 70, 245 - 261. https: // doi. org / 10.3161 / 00034541 ANZ 2020.70.2.005" type="journal article" year="2020">Moctezuma &amp; Halffter 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8B8FE9613E76CD7E0AF3FF5CFA30FEDC" blockId="6.[151,1437,151,446]" pageId="6" pageNumber="128">
12(11). Dorsal surface completely green, or brown with bright green sheen. Apex of parameres straight.
<collectingRegion id="49F427833E76CD7E0EE4FF5CFB6DFF20" box="[1152,1217,219,241]" country="Mexico" name="Jalisco" pageId="6" pageNumber="128">Jalisco</collectingRegion>
(environs of Quila)...............................................................
<taxonomicName id="4C3092E23E76CD7E09AAFF7FFA30FEDC" authority="Moctezuma &amp; Halffter, 2020" authorityName="Moctezuma &amp; Halffter" authorityYear="2020" box="[974,1436,248,270]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="species" species="viridichevrolati">
<emphasis id="B94435733E76CD7E09AAFF7FFBD3FEDC" box="[974,1151,248,269]" italics="true" pageId="6" pageNumber="128">O. viridichevrolati</emphasis>
<bibRefCitation id="EFA194903E76CD7E0EE2FF7FFA30FEDC" author="Moctezuma, V. &amp; Halffter, G." box="[1158,1436,248,270]" pageId="6" pageNumber="128" pagination="245 - 261" refId="ref2149" refString="Moctezuma, V. &amp; Halffter, G. (2020) New species and redescriptions of the Onthophagus chevrolati species complex (Coleoptera: Scarabaeoidea: Scarabaeinae). Annales Zoologici, 70, 245 - 261. https: // doi. org / 10.3161 / 00034541 ANZ 2020.70.2.005" type="journal article" year="2020">Moctezuma &amp; Halffter, 2020</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B8FE9613E76CD7E0AF3FE91FA30FEFB" blockId="6.[151,1437,151,446]" box="[151,1436,277,299]" pageId="6" pageNumber="128">
- Dorsal surface dark brown or black, without bright sheen. Apex of parameres trapezoidal (
<figureCitation id="130BF5E43E76CD7E0E21FE92FBD2FEFA" box="[1093,1150,277,299]" captionStart="FIGURE 3" captionStartId="3.[151,250,1438,1462]" captionTargetBox="[159,1422,344,1404]" captionTargetId="figure-97@3.[151,1436,344,1414]" captionTargetPageId="3" captionText="FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite." figureDoi="http://doi.org/10.5281/zenodo.5656614" httpUri="https://zenodo.org/record/5656614/files/figure.png" pageId="6" pageNumber="128">Fig. 3</figureCitation>
)..................... 13
</paragraph>
<paragraph id="8B8FE9613E76CD7E0AF3FEB4FA30FE52" blockId="6.[151,1437,151,446]" pageId="6" pageNumber="128">
13(12). Secondary pronotal tubercles situated posterolaterally (
<figureCitation id="130BF5E43E76CD7E088BFEB4FC86FE98" box="[751,810,307,329]" captionStart="FIGURE 6" captionStartId="5.[151,255,1943,1967]" captionTargetBox="[232,1359,200,1896]" captionTargetId="figure-27@5.[214,1373,180,1919]" captionTargetPageId="5" captionText="FIGURE 6. Onthophagus chevrolati male (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5656620" httpUri="https://zenodo.org/record/5656620/files/figure.png" pageId="6" pageNumber="128">Fig. 6</figureCitation>
). Dorsal surface dark brown, occasionally with olive green sheen. Apex of parameres distinctly trapezoidal (
<figureCitation id="130BF5E43E76CD7E081BFED7FD61FEB7" box="[639,717,336,358]" captionStart="FIGURE 3" captionStartId="3.[151,250,1438,1462]" captionTargetBox="[159,1422,344,1404]" captionTargetId="figure-97@3.[151,1436,344,1414]" captionTargetPageId="3" captionText="FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite." figureDoi="http://doi.org/10.5281/zenodo.5656614" httpUri="https://zenodo.org/record/5656614/files/figure.png" pageId="6" pageNumber="128">Fig. 3C</figureCitation>
).
<collectingRegion id="49F427833E76CD7E0884FED7FDE2FE52" isEnumeration="true" pageId="6" pageNumber="128">Guanajuato, Hidalgo, Mexico City, Mexico State, Michoacán, Morelos, Puebla, Querétaro, Tlaxcala, Veracruz</collectingRegion>
(
<figureCitation id="130BF5E43E76CD7E083FFEEAFD38FE52" box="[603,660,365,387]" captionStart="FIGURE 5" captionStartId="4.[151,250,1999,2023]" captionTargetBox="[242,1345,1121,1974]" captionTargetId="figure-28@4.[238,1348,1117,1975]" captionTargetPageId="4" captionText="FIGURE 5. Distribution of the Onthophagus chevrolati species complex in central Mexico." figureDoi="http://doi.org/10.5281/zenodo.5656618" httpUri="https://zenodo.org/record/5656618/files/figure.png" pageId="6" pageNumber="128">Fig. 5</figureCitation>
).........................................
<taxonomicName id="4C3092E23E76CD7E0EFEFEE9FA30FE52" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[1178,1436,365,387]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B94435733E76CD7E0EFEFEE9FABBFE53" box="[1178,1303,365,387]" italics="true" pageId="6" pageNumber="128">O. chevrolati</emphasis>
Harold, 1869
</taxonomicName>
</paragraph>
<paragraph id="8B8FE9613E76CD7E0AF3FE0CFA31FE6F" blockId="6.[151,1437,151,446]" pageId="6" pageNumber="128">
– - metallic sheen. Apex of parameres obtusely trapezoidal (
<figureCitation id="130BF5E43E76CD7E0966FE0CFCFCFE70" box="[770,848,395,417]" captionStart="FIGURE 3" captionStartId="3.[151,250,1438,1462]" captionTargetBox="[159,1422,344,1404]" captionTargetId="figure-97@3.[151,1436,344,1414]" captionTargetPageId="3" captionText="FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite." figureDoi="http://doi.org/10.5281/zenodo.5656614" httpUri="https://zenodo.org/record/5656614/files/figure.png" pageId="6" pageNumber="128">Fig. 3A</figureCitation>
).
<collectingRegion id="49F427833E76CD7E0904FE0CFC0DFE70" box="[864,929,395,417]" country="Mexico" name="Jalisco" pageId="6" pageNumber="128">Jalisco</collectingRegion>
(environs of Talpa;
<figureCitation id="130BF5E43E76CD7E0E07FE0CFB37FE71" box="[1123,1179,394,417]" captionStart="FIGURE 5" captionStartId="4.[151,250,1999,2023]" captionTargetBox="[242,1345,1121,1974]" captionTargetId="figure-28@4.[238,1348,1117,1975]" captionTargetPageId="4" captionText="FIGURE 5. Distribution of the Onthophagus chevrolati species complex in central Mexico." figureDoi="http://doi.org/10.5281/zenodo.5656618" httpUri="https://zenodo.org/record/5656618/files/figure.png" pageId="6" pageNumber="128">Fig. 5</figureCitation>
)...............................................................
<taxonomicName id="4C3092E23E76CD7E0896FE2FFAB1FE6F" authority="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityName="Moctezuma, Hernandez, &amp; Sanchez-Huerta" authorityYear="2021" box="[754,1309,424,446]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="species" species="acernorus" status="sp. nov.">
<emphasis id="B94435733E76CD7E0896FE2FFCDDFE6F" box="[754,881,424,446]" italics="true" pageId="6" pageNumber="128">O. acernorus</emphasis>
Moctezuma, Hernández, &amp; Sánchez-Huerta
</taxonomicName>
,
<emphasis id="B94435733E76CD7E0F42FE2FFA31FE6F" bold="true" box="[1318,1437,424,446]" pageId="6" pageNumber="128">
<taxonomicNameLabel id="A27788083E76CD7E0F42FE2FFA31FE6F" box="[1318,1437,424,446]" pageId="6" pageNumber="128" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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