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<emphasis id="B953EADEFFCAE42AFE96F8C5FD9AF89F" bold="true" box="[302,539,1810,1837]" italics="true" pageId="44" pageNumber="537">Nucellopsis parisiensis</emphasis>
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<taxonomicNameLabel id="A26057A5FFCAE42AFD9AF8C4FDDCF89F" box="[546,605,1810,1836]" pageId="44" pageNumber="537" rank="species">n. sp.</taxonomicNameLabel>
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(
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A-F; 28F, G)
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<taxonomicName id="4C274D4FFFCAE42AFF3CF86FFEE7F862" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[132,358,1977,2001]" class="Gastropoda" family="Muricidae" genus="Muricopsis" kingdom="Animalia" order="Neogastropoda" pageId="44" pageNumber="537" phylum="Mollusca" rank="species" species="depauperatus">
<emphasis id="B953EADEFFCAE42AFF3CF86FFEE7F862" box="[132,358,1977,2001]" italics="true" pageId="44" pageNumber="537">Muricopsis depauperatus</emphasis>
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<bibRefCitation id="EFB64B3DFFCAE42AFEC2F86FFD00F862" author="COSSMANN M. &amp; PISSARRO G." box="[378,641,1977,2001]" pageId="44" pageNumber="537" pagination="26 - 45" refId="ref43883" refString="COSSMANN M. &amp; PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene du Bassin de Paris. Tome 2. Hermann, Paris: pl. 26 - 45. https: // www. biodiversitylibrary. org / page / 36319938" type="book chapter" year="1911">Cossmann &amp; Pissarro 1911</bibRefCitation>
: fig. 169bis-2 [
<emphasis id="B953EADEFFCAE42AFF34F802FF31F858" box="[140,176,2004,2027]" italics="true" pageId="44" pageNumber="537">non</emphasis>
<bibRefCitation id="EFB64B3DFFCAE42AFF0EF805FED5F858" author="DESHAYES G. - P." box="[182,340,2003,2027]" pageId="44" pageNumber="537" pagination="101 - 107" refId="ref44001" refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]." type="journal article" year="1865">Deshayes, 1865</bibRefCitation>
].
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sp.
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non
<taxonomicName id="4C274D4FFFCAE42AFC53F8C7FBDEF89A" ID-CoL="8YMTF" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[1003,1119,1809,1833]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="44" pageNumber="537" phylum="Mollusca" rank="species" species="depauperata">depauperata</taxonomicName>
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Merle
<emphasis id="B953EADEFFCAE42AFB78F8C4FB55F89A" box="[1216,1236,1810,1833]" italics="true" pageId="44" pageNumber="537">in</emphasis>
<bibRefCitation id="EFB64B3DFFCAE42AFB63F8C4FC15F8F7" author="LE RENARD J. &amp; PACAUD J. - M." pageId="44" pageNumber="537" pagination="65 - 132" refId="ref45932" refString="LE RENARD J. &amp; PACAUD J. - M. 1995. - Revision des Mollusques paleogenes du Bassin de Paris. 2: Liste des references primaires des especes. Cossmanniana 3 (3): 65 - 132." type="journal article" year="1995">Le Renard &amp; Pacaud 1995: 114</bibRefCitation>
, réf.: GA 169bis-2.
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<taxonomicName id="4C274D4FFFCAE42AFC95F881FB83F8DC" ID-CoL="86ZBQ" baseAuthorityName="Strauzs" baseAuthorityYear="1966" box="[813,1026,1879,1903]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="44" pageNumber="537" phylum="Mollusca" rank="species" species="dudariensis">
<emphasis id="B953EADEFFCAE42AFC95F881FB83F8DC" box="[813,1026,1879,1903]" italics="true" pageId="44" pageNumber="537">Nucellopsis dudariensis</emphasis>
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<bibRefCitation id="EFB64B3DFFCAE42AFBA4F881FB19F8DC" author="MERLE D." box="[1052,1176,1879,1903]" pageId="44" pageNumber="537" pagination="177 - 189" refId="ref46903" refString="MERLE D. 2005 c. - Nucellopsis (Gastropoda, Muricidae), a new genus from the paralic domain of the European Palaeogene. Comptes Rendus Palevol 4 (1 - 2): 177 - 189. https: // doi. org / 10.1016 / j. crpv. 2004.11.005" type="journal article" year="2005">Merle 2005c</bibRefCitation>
[
<emphasis id="B953EADEFFCAE42AFB1FF88EFB4AF8DC" box="[1191,1227,1880,1903]" italics="true" pageId="44" pageNumber="537">non</emphasis>
<bibRefCitation id="EFB64B3DFFCAE42AFB69F88EFAD8F8DC" author="STRAUSZ L." box="[1233,1369,1879,1904]" pageId="44" pageNumber="537" pagination="1 - 200" refId="ref48690" refString="STRAUSZ L. 1966. - Dudari Eocen Csigak. Geologica Hungarica (series Palaeontologica) 33: 1 - 200." type="journal article" year="1966">Strausz, 1966</bibRefCitation>
].
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. —
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.
<collectingCountry id="F330765CFFCAE42AFBF3F855FB0AF828" box="[1099,1163,1923,1947]" name="France" pageId="44" pageNumber="537">France</collectingCountry>
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• Paris Basin, Val dOise, Saint-Witz (Le Guépelle); Sable du Guépelle (biozone NP16); middle Eocene (early Bartonian, Auversian); MNHN.F.R64955 (
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coll.), figured specimen in
<bibRefCitation id="EFB64B3DFFCAE42AFC19F805FBA0F858" author="MERLE D." box="[929,1057,2003,2027]" pageId="44" pageNumber="537" pagination="177 - 189" refId="ref46903" refString="MERLE D. 2005 c. - Nucellopsis (Gastropoda, Muricidae), a new genus from the paralic domain of the European Palaeogene. Comptes Rendus Palevol 4 (1 - 2): 177 - 189. https: // doi. org / 10.1016 / j. crpv. 2004.11.005" type="journal article" year="2005">
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(2005c
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: fig. 2H), H:
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(
<figureCitation id="131C2A49FFCAE42AFABEF805FAF2F858" box="[1286,1395,2003,2027]" captionStart="FIG" captionStartId="42.[132,143,1281,1298]" captionTargetBox="[132,1455,215,1223]" captionTargetId="figure-489@42.[143,795,242,771]" captionTargetPageId="42" captionText="FIG. 23. — Spiral cords and internal denticles of the outer lip in Nucellopsis parisiensis n. sp. (A-F) and N. dudariensis (Strauzs, 1966) (G, H): A, B, paratype, MNHN.F.A90964 (Faullummel coll.), Saint-Witz (Le Guépelle), Val dOise, Paris Basin, France, middle Eocene (Bartonian); C, D, paratype, MNHN.F.A90963 (Faullummel coll.),Saint-Witz (Le Guépelle);E, F, holotype,MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle);G, H, holotype of Cantharus (Pollia) dudariensis, MGSH E.467, Dudar, Hungary, middle Eocene (upper Lutetian). Scale bars: 5 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E, F, D. Serrette (MNHN); G, H, courtesy of Z. Kovács." figureDoi="http://doi.org/10.5281/zenodo.13985819" httpUri="https://zenodo.org/record/13985819/files/figure.png" pageId="44" pageNumber="537">Figs 23E, F</figureCitation>
;
<figureCitation id="131C2A49FFCAE42AFAC1F805FA25F85F" box="[1401,1444,2003,2028]" captionStart="FIG" captionStartId="48.[132,143,1301,1318]" captionTargetBox="[145,1443,235,1238]" captionTargetId="figure-535@48.[700,1258,235,783]" captionTargetPageId="48" captionText="FIG. 28. — New species of Flexopteron Shuto, 1969, Crassimurex (Pliocrassimurex) n. subgen., Nucellopsis Merle, 2005 and Pronucellopsis Merle, n. gen.: A, B, Flexopteron liancurtense n. sp.: A, holotype, MNHN.F.J02578 (Cossmann coll.), Liancourt-Saint-Pierre, France, Ypresian; B, paratype: MNHN.F.A77758 (Merle coll.), Liancourt-Saint-Pierre; C-E, Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp.: C, paratype (juvenile), MNHN.F.A77753, Molins del Rei, Spain, Zanclean; D, paratype, MNHN.F.A77752, Molins del Rei; E, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei; F, G, Nucellopsis parisiensis n. sp.: F, holotype, MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle), France, Bartonian; G, paratype, MNHN.F.A90963 (Faullummel coll.), Saint-Witz (Le Guépelle); H, I, Pronucellopsis pacaudi Merle, Ledon &amp; Goret, n. gen., n. sp.: H, holotype, MNHN.F.J02587 (Cossmann coll.), Châlons-sur-Vesle, France, Thanetian; I, paratype, MNHN.F.A90829 (Pacaud coll.), Jonchery-sur-Vesle, France, Thanetian. Scale bars: 5 mm. Scale bar: A, B, 3 mm; C, D, F-I, 5 mm; E, 10 mm. Credits: P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985833" httpUri="https://zenodo.org/record/13985833/files/figure.png" pageId="44" pageNumber="537">28F</figureCitation>
).
</materialsCitation>
<materialsCitation id="3B4F3C91FFCBE42BFF3CF852FD2AF804" ID-GBIF-Occurrence="4958671366" pageId="45" pageNumber="538">
<emphasis id="B953EADEFFCBE42BFF3CF852FEA7F82F" bold="true" box="[132,294,1924,1948]" pageId="45" pageNumber="538">
<typeStatus id="549C886EFFCBE42BFF3CF852FF61F82F" box="[132,224,1924,1948]" pageId="45" pageNumber="538" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F330765CFFCBE42BFF50F852FEA7F82F" box="[232,294,1924,1948]" name="France" pageId="45" pageNumber="538">France</collectingCountry>
</emphasis>
• 1 spm; same as for the holotype; MNHN.F.A71797 (Pons coll.), figured specimen in
<bibRefCitation id="EFB64B3DFFCBE42BFE73F848FDCEF805" author="MERLE D." box="[459,591,1950,1974]" pageId="45" pageNumber="538" pagination="177 - 189" refId="ref46903" refString="MERLE D. 2005 c. - Nucellopsis (Gastropoda, Muricidae), a new genus from the paralic domain of the European Palaeogene. Comptes Rendus Palevol 4 (1 - 2): 177 - 189. https: // doi. org / 10.1016 / j. crpv. 2004.11.005" type="journal article" year="2005">
<taxonomicName id="4C274D4FFFCBE42BFE73F848FD85F805" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[459,516,1950,1974]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(2005c
</bibRefCitation>
: fig. 1G)
</materialsCitation>
<materialsCitation id="3B4F3C91FFCBE42BFD7CF849FEBCF85F" ID-GBIF-Occurrence="4958671309" pageId="45" pageNumber="538">
1 spm; idem; MNHN.F.A71798 (Pons coll.), figured specimen in
<bibRefCitation id="EFB64B3DFFCBE42BFD77F86FFF48F858" author="MERLE D." pageId="45" pageNumber="538" pagination="177 - 189" refId="ref46903" refString="MERLE D. 2005 c. - Nucellopsis (Gastropoda, Muricidae), a new genus from the paralic domain of the European Palaeogene. Comptes Rendus Palevol 4 (1 - 2): 177 - 189. https: // doi. org / 10.1016 / j. crpv. 2004.11.005" type="journal article" year="2005">
<taxonomicName id="4C274D4FFFCBE42BFD77F86FFC89F862" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[719,776,1977,2001]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(2005c
</bibRefCitation>
: fig. 1H, I)
</materialsCitation>
<materialsCitation id="3B4F3C91FFCBE42BFEEDF805FA2EF82F" ID-GBIF-Occurrence="4958671412" box="[341,1455,1924,2027]" pageId="45" pageNumber="538">
1 spm; idem; MNHN.F.J02585 (Cossmann coll.); figured in
<bibRefCitation id="EFB64B3DFFCBE42BFC76F852FB60F82F" author="COSSMANN M. &amp; PISSARRO G." box="[974,1249,1924,1948]" pageId="45" pageNumber="538" pagination="26 - 45" refId="ref43883" refString="COSSMANN M. &amp; PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene du Bassin de Paris. Tome 2. Hermann, Paris: pl. 26 - 45. https: // www. biodiversitylibrary. org / page / 36319938" type="book chapter" year="1911">Cossmann &amp; Pissarro (1911</bibRefCitation>
: pl. 36 fig. 169bis-2)
</materialsCitation>
<materialsCitation id="3B4F3C91FFCBE42BFC86F849FC04F862" ID-GBIF-Occurrence="4958671360" pageId="45" pageNumber="538">
1 spm; idem; MNHN.F.A90963 (Faullummel coll.) (
<figureCitation id="131C2A49FFCBE42BFAF7F848FCC2F863" captionStart="FIG" captionStartId="42.[132,143,1281,1298]" captionTargetBox="[132,1455,215,1223]" captionTargetId="figure-489@42.[143,795,242,771]" captionTargetPageId="42" captionText="FIG. 23. — Spiral cords and internal denticles of the outer lip in Nucellopsis parisiensis n. sp. (A-F) and N. dudariensis (Strauzs, 1966) (G, H): A, B, paratype, MNHN.F.A90964 (Faullummel coll.), Saint-Witz (Le Guépelle), Val dOise, Paris Basin, France, middle Eocene (Bartonian); C, D, paratype, MNHN.F.A90963 (Faullummel coll.),Saint-Witz (Le Guépelle);E, F, holotype,MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle);G, H, holotype of Cantharus (Pollia) dudariensis, MGSH E.467, Dudar, Hungary, middle Eocene (upper Lutetian). Scale bars: 5 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E, F, D. Serrette (MNHN); G, H, courtesy of Z. Kovács." figureDoi="http://doi.org/10.5281/zenodo.13985819" httpUri="https://zenodo.org/record/13985819/files/figure.png" pageId="45" pageNumber="538">Figs 23C, D</figureCitation>
;
<figureCitation id="131C2A49FFCBE42BFCF7F86FFCFCF862" box="[847,893,1977,2001]" captionStart="FIG" captionStartId="48.[132,143,1301,1318]" captionTargetBox="[145,1443,235,1238]" captionTargetId="figure-535@48.[700,1258,235,783]" captionTargetPageId="48" captionText="FIG. 28. — New species of Flexopteron Shuto, 1969, Crassimurex (Pliocrassimurex) n. subgen., Nucellopsis Merle, 2005 and Pronucellopsis Merle, n. gen.: A, B, Flexopteron liancurtense n. sp.: A, holotype, MNHN.F.J02578 (Cossmann coll.), Liancourt-Saint-Pierre, France, Ypresian; B, paratype: MNHN.F.A77758 (Merle coll.), Liancourt-Saint-Pierre; C-E, Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp.: C, paratype (juvenile), MNHN.F.A77753, Molins del Rei, Spain, Zanclean; D, paratype, MNHN.F.A77752, Molins del Rei; E, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei; F, G, Nucellopsis parisiensis n. sp.: F, holotype, MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle), France, Bartonian; G, paratype, MNHN.F.A90963 (Faullummel coll.), Saint-Witz (Le Guépelle); H, I, Pronucellopsis pacaudi Merle, Ledon &amp; Goret, n. gen., n. sp.: H, holotype, MNHN.F.J02587 (Cossmann coll.), Châlons-sur-Vesle, France, Thanetian; I, paratype, MNHN.F.A90829 (Pacaud coll.), Jonchery-sur-Vesle, France, Thanetian. Scale bars: 5 mm. Scale bar: A, B, 3 mm; C, D, F-I, 5 mm; E, 10 mm. Credits: P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985833" httpUri="https://zenodo.org/record/13985833/files/figure.png" pageId="45" pageNumber="538">28G</figureCitation>
)
</materialsCitation>
<materialsCitation id="3B4F3C91FFCBE42BFC27F86FFC29F85F" ID-GBIF-Occurrence="4958671387" pageId="45" pageNumber="538">
1 spm; idem; MNHN.F.A90964 (Faullummel coll.) (
<figureCitation id="131C2A49FFCBE42BFC8DF805FC20F85F" box="[821,929,2003,2028]" captionStart="FIG" captionStartId="42.[132,143,1281,1298]" captionTargetBox="[132,1455,215,1223]" captionTargetId="figure-489@42.[143,795,242,771]" captionTargetPageId="42" captionText="FIG. 23. — Spiral cords and internal denticles of the outer lip in Nucellopsis parisiensis n. sp. (A-F) and N. dudariensis (Strauzs, 1966) (G, H): A, B, paratype, MNHN.F.A90964 (Faullummel coll.), Saint-Witz (Le Guépelle), Val dOise, Paris Basin, France, middle Eocene (Bartonian); C, D, paratype, MNHN.F.A90963 (Faullummel coll.),Saint-Witz (Le Guépelle);E, F, holotype,MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle);G, H, holotype of Cantharus (Pollia) dudariensis, MGSH E.467, Dudar, Hungary, middle Eocene (upper Lutetian). Scale bars: 5 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E, F, D. Serrette (MNHN); G, H, courtesy of Z. Kovács." figureDoi="http://doi.org/10.5281/zenodo.13985819" httpUri="https://zenodo.org/record/13985819/files/figure.png" pageId="45" pageNumber="538">Fig. 23A, B</figureCitation>
)
</materialsCitation>
<materialsCitation id="3B4F3C91FFCBE42BFC03F805FA31F858" ID-GBIF-Occurrence="4958671330" box="[955,1456,2003,2027]" pageId="45" pageNumber="538">6 spm; idem; MNHN.F.A90962 (Faullummel coll.).</materialsCitation>
<smallCapsWord id="8D7EA010FFC8E428FF3CF852FF7EF828" baselines="1942,1943" box="[132,255,1924,1948]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Additional" pageId="46" pageNumber="539">ADDITIONAL</smallCapsWord>
<smallCapsWord id="8D7EA010FFC8E428FEBDF85CFEE2F828" baselines="1943" box="[261,355,1930,1947]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="46" pageNumber="539">MATERIAL</smallCapsWord>
. —
<materialsCitation id="3B4F3C91FFC8E428FE34F852FDB4F862" ID-GBIF-Occurrence="4958671410" pageId="46" pageNumber="539">
<emphasis id="B953EADEFFC8E428FE34F852FE4FF82F" bold="true" box="[396,462,1924,1948]" pageId="46" pageNumber="539">
<collectingCountry id="F330765CFFC8E428FE34F852FE4FF82F" box="[396,462,1924,1948]" name="France" pageId="46" pageNumber="539">France</collectingCountry>
</emphasis>
• 1 spm; Paris Basin, Val dOise, Saint-Witz (Le Guépelle); Sable du Guépelle (biozone NP16); early Bartonian; MNHN.F.A27771 (Pacaud coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFC8E428FDE8F86FFC33F82F" ID-GBIF-Occurrence="4958671377" pageId="46" pageNumber="539">1 spm; Paris Basin, Val dOise, Auvers-sur-Oise (Bois-le-Roi); idem; MNHN.F.A27770 (Pacaud coll.)</materialsCitation>
<materialsCitation id="3B4F3C91FFC8E428FC70F852FB0CF805" ID-GBIF-Occurrence="4958671328" pageId="46" pageNumber="539">1 spm; Paris Basin, Aisne, Nogent-lArtaud; idem; MNHN.F.A90565 (Margerie coll.)</materialsCitation>
<materialsCitation id="3B4F3C91FFC8E428FB10F849FADEF862" ID-GBIF-Occurrence="4958671407" pageId="46" pageNumber="539">1 spm; Paris Basin, Aisne, Bézu-le-Guéry; idem;MNHN.F.A90825 (Faullummel coll.)</materialsCitation>
<materialsCitation id="3B4F3C91FFC8E429FAC9F86FFE8FFF53" ID-GBIF-Occurrence="4958671369" lastPageId="47" lastPageNumber="540" pageId="46" pageNumber="539">1 spm; Paris Basin, Aisne, Mont-Saint-Martin; idem; MNHN.F.J07787 (UMPC coll.)</materialsCitation>
<materialsCitation id="3B4F3C91FFC9E429FE9DFF1FFD35FF48" ID-GBIF-Occurrence="4958671421" pageId="47" pageNumber="540">1 spm; Paris Basin, Seine-et-Marne, Sainte-Aulde (Caumont); idem; MNHN.F.A90826 (Faullummel coll.)</materialsCitation>
<materialsCitation id="3B4F3C91FFC9E429FD7FFF35FE86FE83" ID-GBIF-Occurrence="4958671417" pageId="47" pageNumber="540">1 spm; Paris Basin, Seine-et-Marne, Luzancy; idem; MNHN.F.A90827 (Faullummel coll.)</materialsCitation>
<materialsCitation id="3B4F3C91FFC9E429FEA6FECEFEC1FEF9" ID-GBIF-Occurrence="4958671306" pageId="47" pageNumber="540">Paris Basin, Oise, Loisy; idem; MNHN.F.A90828 (Faullummel coll.).</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF586644FFCBE42BFF3CF980FE35F880" ID-DOI="http://doi.org/10.5281/zenodo.13985825" ID-Zenodo-Dep="13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="45" pageNumber="538" startId="45.[132,143,1622,1639]" targetBox="[132,1455,215,1562]" targetPageId="45" targetType="figure">
<paragraph id="8B9836CCFFCBE42BFF3CF980FE35F880" blockId="45.[132,1457,1622,1843]" pageId="45" pageNumber="538">
<smallCapsWord id="8D7EA010FFCBE42BFF3CF980FF1CF9D5" baselines="1635,1635" box="[132,157,1622,1639]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="45" pageNumber="538">FIG</smallCapsWord>
. 26. — New species of
<taxonomicName id="4C274D4FFFCBE42BFEDBF980FDD5F9D4" authority="De Gregorio, 1885" authorityName="de Gregorio" authorityYear="1885" box="[355,596,1622,1639]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCBE42BFEDBF980FE32F9D4" box="[355,435,1622,1639]" italics="true" pageId="45" pageNumber="538">Timbellus</emphasis>
De Gregorio, 1885
</taxonomicName>
:
<emphasis id="B953EADEFFCBE42BFDE2F980FDE6F9D4" bold="true" box="[602,615,1622,1639]" pageId="45" pageNumber="538">A</emphasis>
,
<emphasis id="B953EADEFFCBE42BFDC9F980FDFFF9D4" bold="true" box="[625,638,1622,1639]" pageId="45" pageNumber="538">B</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFD31F980FCC1F9D4" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[649,832,1622,1639]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="B953EADEFFCBE42BFD31F980FCC1F9D4" box="[649,832,1622,1639]" italics="true" pageId="45" pageNumber="538">Timbellus occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCBE42BFCFDF980FCF6F9D4" box="[837,887,1622,1639]" pageId="45" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFCBE42BFCC5F980FC0BF9D4" bold="true" box="[893,906,1622,1639]" pageId="45" pageNumber="538">A</emphasis>
, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;
<emphasis id="B953EADEFFCBE42BFEBCF9BAFE90F9CE" bold="true" box="[260,273,1644,1661]" pageId="45" pageNumber="538">B</emphasis>
, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;
<emphasis id="B953EADEFFCBE42BFC03F9BAFC48F9CE" bold="true" box="[955,969,1644,1661]" pageId="45" pageNumber="538">C</emphasis>
,
<emphasis id="B953EADEFFCBE42BFC6AF9BAFC61F9CE" bold="true" box="[978,992,1644,1661]" pageId="45" pageNumber="538">D</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFC51F9BAFBC8F9CE" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1001,1097,1644,1661]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="calciacus" status="n. sp.">
<emphasis id="B953EADEFFCBE42BFC51F9BAFBC8F9CE" box="[1001,1097,1644,1661]" italics="true" pageId="45" pageNumber="538">T.calciacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCBE42BFBF4F9BAFBFFF9CE" box="[1100,1150,1644,1661]" pageId="45" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFCBE42BFB3BF9BAFB10F9CE" bold="true" box="[1155,1169,1644,1661]" pageId="45" pageNumber="538">C</emphasis>
, holotype,MNHN.F.A90536 (
<taxonomicName id="4C274D4FFFCBE42BFA38F9BAFA2EF9CE" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1408,1455,1644,1661]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), Chaussy (Les Garennes), France, Lutetian;
<emphasis id="B953EADEFFCBE42BFD99F955FDAEF927" bold="true" box="[545,559,1667,1684]" pageId="45" pageNumber="538">D</emphasis>
, paratype, MNHN.F.A90537 (
<taxonomicName id="4C274D4FFFCBE42BFC9FF955FCD7F927" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[807,854,1667,1684]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), Chaussy (Les Garennes);
<emphasis id="B953EADEFFCBE42BFBDDF955FBF0F927" bold="true" box="[1125,1137,1667,1684]" pageId="45" pageNumber="538">E</emphasis>
,
<emphasis id="B953EADEFFCBE42BFBC5F955FB09F927" bold="true" box="[1149,1160,1667,1684]" pageId="45" pageNumber="538">F</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFB2BF955FA8BF927" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1171,1290,1667,1684]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="longicanalis" status="n. sp.">
<emphasis id="B953EADEFFCBE42BFB2BF955FA8BF927" box="[1171,1290,1667,1684]" italics="true" pageId="45" pageNumber="538">T. longicanalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCBE42BFAA9F955FAC2F927" box="[1297,1347,1667,1684]" pageId="45" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFCBE42BFAF2F955FAD7F927" bold="true" box="[1354,1366,1667,1684]" pageId="45" pageNumber="538">E</emphasis>
, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian;
<emphasis id="B953EADEFFCBE42BFD59F94CFD6DF918" bold="true" box="[737,748,1690,1707]" pageId="45" pageNumber="538">F</emphasis>
, paratype, MNHN.F.A24100, Fresville;
<emphasis id="B953EADEFFCBE42BFB8FF94CFBD3F918" bold="true" box="[1079,1106,1690,1707]" pageId="45" pageNumber="538">G -I</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFBE5F94CFB30F918" box="[1117,1201,1690,1707]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFCBE42BFBE5F94CFB30F918" box="[1117,1201,1690,1707]" italics="true" pageId="45" pageNumber="538">T. magnei</emphasis>
</taxonomicName>
n. name:
<emphasis id="B953EADEFFCBE42BFABFF94CFA94F918" bold="true" box="[1287,1301,1690,1707]" pageId="45" pageNumber="538">G</emphasis>
, MNHN.F.A32508 (
<taxonomicName id="4C274D4FFFCBE42BFF31F966FF39F972" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[137,184,1712,1729]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), juvenile specimen, Gan (Tuilerie), France;
<emphasis id="B953EADEFFCBE42BFDF7F966FDDCF972" bold="true" box="[591,605,1712,1729]" pageId="45" pageNumber="538">H</emphasis>
, MNHN.F.A90545 (
<taxonomicName id="4C274D4FFFCBE42BFCB8F966FCAEF972" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[768,815,1712,1729]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), juvenile specimen with protoconch, Gan (Tuilerie);
<emphasis id="B953EADEFFCBE42BFAA9F966FA96F972" bold="true" box="[1297,1303,1712,1729]" pageId="45" pageNumber="538">I</emphasis>
, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie);
<emphasis id="B953EADEFFCBE42BFEC9F911FE0FF96B" bold="true" box="[369,398,1735,1752]" pageId="45" pageNumber="538">J -L</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFE20F911FD80F96B" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[408,513,1735,1752]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="curvispina" status="n. sp.">
<emphasis id="B953EADEFFCBE42BFE20F911FD80F96B" box="[408,513,1735,1752]" italics="true" pageId="45" pageNumber="538">T. curvispina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCBE42BFDBEF911FDB9F96B" box="[518,568,1735,1752]" pageId="45" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFCBE42BFD87F911FDC8F96B" bold="true" box="[575,585,1735,1752]" pageId="45" pageNumber="538">J</emphasis>
, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian;
<emphasis id="B953EADEFFCBE42BFB39F911FB0FF96B" bold="true" box="[1153,1166,1735,1752]" pageId="45" pageNumber="538">K</emphasis>
, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian;
<emphasis id="B953EADEFFCBE42BFE55F908FE79F95C" bold="true" box="[493,504,1758,1775]" pageId="45" pageNumber="538">L</emphasis>
, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin;
<emphasis id="B953EADEFFCBE42BFC4DF908FB87F95C" bold="true" box="[1013,1030,1758,1775]" pageId="45" pageNumber="538">M</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFBA8F908FBBFF95C" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1040,1086,1758,1775]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="sixi" status="n. sp.">
<emphasis id="B953EADEFFCBE42BFBA8F908FBBFF95C" box="[1040,1086,1758,1775]" italics="true" pageId="45" pageNumber="538">T. sixi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCBE42BFBFBF908FBF4F95C" box="[1091,1141,1758,1775]" pageId="45" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
, holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian;
<emphasis id="B953EADEFFCBE42BFDA7F922FDC0F8B6" bold="true" box="[543,577,1780,1797]" pageId="45" pageNumber="538">N -P</emphasis>
,
<taxonomicName id="4C274D4FFFCBE42BFDF4F922FD41F8B6" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[588,704,1780,1797]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="538" phylum="Mollusca" rank="species" species="radulfiensis" status="n. sp.">
<emphasis id="B953EADEFFCBE42BFDF4F922FD41F8B6" box="[588,704,1780,1797]" italics="true" pageId="45" pageNumber="538">T. radulfiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCBE42BFD7DF922FD79F8B6" box="[709,760,1780,1797]" pageId="45" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFCBE42BFCB8F922FC8FF8B6" bold="true" box="[768,782,1780,1797]" pageId="45" pageNumber="538">N</emphasis>
, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian;
<emphasis id="B953EADEFFCBE42BFF3CF8DDFF12F8AF" bold="true" box="[132,147,1803,1820]" pageId="45" pageNumber="538">O</emphasis>
, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place;
<emphasis id="B953EADEFFCBE42BFD2AF8DDFD1FF8AF" bold="true" box="[658,670,1803,1820]" pageId="45" pageNumber="538">P</emphasis>
, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm.
</paragraph>
</caption>
<caption id="DF586644FFC8E428FF3CF990FEF7F889" ID-DOI="http://doi.org/10.5281/zenodo.13985831" ID-Zenodo-Dep="13985831" httpUri="https://zenodo.org/record/13985831/files/figure.png" pageId="46" pageNumber="539" startId="46.[132,143,1606,1623]" targetBox="[132,1455,215,1546]" targetPageId="46" targetType="figure">
<paragraph id="8B9836CCFFC8E428FF3CF990FEF7F889" blockId="46.[132,1457,1606,1850]" pageId="46" pageNumber="539">
<smallCapsWord id="8D7EA010FFC8E428FF3CF990FF1CF9E5" baselines="1619,1619" box="[132,157,1606,1623]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="46" pageNumber="539">FIG</smallCapsWord>
. 27. — New species of
<taxonomicName id="4C274D4FFFC8E428FEDEF990FDD9F9E4" authority="De Gregorio, 1885" authorityName="de Gregorio" authorityYear="1885" box="[358,600,1606,1623]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC8E428FEDEF990FE37F9E4" box="[358,438,1606,1623]" italics="true" pageId="46" pageNumber="539">Timbellus</emphasis>
De Gregorio, 1885
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC8E428FDD9F990FCC7F9E4" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" box="[609,838,1606,1623]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC8E428FDD9F990FD3AF9E4" box="[609,699,1606,1623]" italics="true" pageId="46" pageNumber="539">Pterynotus</emphasis>
Swainson, 1833
</taxonomicName>
(
<taxonomicName id="4C274D4FFFC8E428FCE8F990FBB2F9E4" authority="Houart, 1995" authorityName="Houart" authorityYear="1995" box="[848,1075,1606,1623]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC8E428FCE8F990FC3CF9E4" box="[848,957,1606,1623]" italics="true" pageId="46" pageNumber="539">Pterymarchia</emphasis>
<bibRefCitation id="EFB64B3DFFC8E428FC7BF990FBB2F9E4" author="HOUART R." box="[963,1075,1606,1623]" pageId="46" pageNumber="539" pagination="127 - 136" refId="ref44916" refString="HOUART R. 1995. - Pterymarchia n. gen. and Vaughtia n. gen., two new muricid genera (Gastropoda: Muricidae: Muricinae and Ocenebrinae). Apex 10 (4): 127 - 136. https: // www. biodiversitylibrary. org / page / 42257087" type="journal article" year="1995">Houart, 1995</bibRefCitation>
</taxonomicName>
),
<taxonomicName id="4C274D4FFFC8E428FB87F990FA98F9E4" authority="Merle, 2011" authorityName="Merle" authorityYear="2011" box="[1087,1305,1606,1623]" class="Gastropoda" family="Muricidae" genus="Textiliomurex" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC8E428FB87F990FB2CF9E4" box="[1087,1197,1606,1623]" italics="true" pageId="46" pageNumber="539">Textiliomurex</emphasis>
<bibRefCitation id="EFB64B3DFFC8E428FB0BF990FA98F9E4" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[1203,1305,1606,1623]" pageId="46" pageNumber="539" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">Merle, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC8E428FA99F990FF2FF9DD" authority="Houart, 1986" authorityName="Houart" authorityYear="1986" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC8E428FA99F990FAEAF9E4" box="[1313,1387,1606,1623]" italics="true" pageId="46" pageNumber="539">Ponderia</emphasis>
Houart, 1986
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC8E428FF60F98BFE2BF9DD" authority="Shuto, 1969" authorityName="Shuto" authorityYear="1969" box="[216,426,1629,1646]" class="Gastropoda" family="Muricidae" genus="Flexopteron" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC8E428FF60F98BFEBAF9DD" box="[216,315,1629,1646]" italics="true" pageId="46" pageNumber="539">Flexopteron</emphasis>
Shuto, 1969
</taxonomicName>
:
<emphasis id="B953EADEFFC8E428FE0AF98BFE55F9DD" bold="true" box="[434,468,1629,1646]" pageId="46" pageNumber="539">A -C</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FE66F98BFD34F9DD" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[478,693,1629,1646]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="submicropterus" status="sp. nov.">
<emphasis id="B953EADEFFC8E428FE66F98BFD34F9DD" box="[478,693,1629,1646]" italics="true" pageId="46" pageNumber="539">Timbellus submicropterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FD03F98BFD6FF9DD" box="[699,750,1629,1646]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFC8E428FD4CF98BFC80F9DD" bold="true" box="[756,769,1629,1646]" pageId="46" pageNumber="539">A</emphasis>
, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian;
<emphasis id="B953EADEFFC8E428FAF0F98BFAD4F9DD" bold="true" box="[1352,1365,1629,1646]" pageId="46" pageNumber="539">B</emphasis>
, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian;
<emphasis id="B953EADEFFC8E428FD58F9A5FD6FF937" bold="true" box="[736,750,1651,1668]" pageId="46" pageNumber="539">C</emphasis>
, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville;
<emphasis id="B953EADEFFC8E428FB15F9A5FB3AF937" bold="true" box="[1197,1211,1651,1668]" pageId="46" pageNumber="539">D</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FB7DF9A5FAF7F937" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1221,1398,1651,1668]" class="Gastropoda" family="Muricidae" genus="Timbellus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="submicropterus" status="n. sp.">
<emphasis id="B953EADEFFC8E428FB7DF9A5FB56F937" box="[1221,1239,1651,1668]" italics="true" pageId="46" pageNumber="539">T.</emphasis>
cf.
<emphasis id="B953EADEFFC8E428FB4CF9A5FAF7F937" box="[1268,1398,1651,1668]" italics="true" pageId="46" pageNumber="539">submicropterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FAC4F9A5FA2FF937" box="[1404,1454,1651,1668]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
, MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian;
<emphasis id="B953EADEFFC8E428FC27F95CFC2AF928" bold="true" box="[927,939,1674,1691]" pageId="46" pageNumber="539">E</emphasis>
,
<emphasis id="B953EADEFFC8E428FC0FF95CFC43F928" bold="true" box="[951,962,1674,1691]" pageId="46" pageNumber="539">F</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FC77F95CFA85F928" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[975,1284,1674,1691]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="gaasensis" status="sp. nov." subGenus="Pterymarchia">
<emphasis id="B953EADEFFC8E428FC77F95CFBABF928" box="[975,1066,1674,1691]" italics="true" pageId="46" pageNumber="539">Pterynotus</emphasis>
(
<emphasis id="B953EADEFFC8E428FB8EF95CFB24F928" box="[1078,1189,1674,1691]" italics="true" pageId="46" pageNumber="539">Pterymarchia</emphasis>
)
<emphasis id="B953EADEFFC8E428FB09F95CFA85F928" box="[1201,1284,1674,1691]" italics="true" pageId="46" pageNumber="539">gaasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FAB4F95CFABEF928" box="[1292,1343,1674,1691]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFC8E428FAF0F95CFAD5F928" bold="true" box="[1352,1364,1674,1691]" pageId="46" pageNumber="539">E</emphasis>
, holotype, MNHN.F.A32492 (
<taxonomicName id="4C274D4FFFC8E428FEA5F977FECDF901" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[285,332,1697,1714]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), Gaas (Espibos), France, Rupelian;
<emphasis id="B953EADEFFC8E428FD11F977FD35F901" bold="true" box="[681,692,1697,1714]" pageId="46" pageNumber="539">F</emphasis>
, paratype, MNHN.F.A90554 (
<taxonomicName id="4C274D4FFFC8E428FC15F977FC5DF901" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[941,988,1697,1714]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), Gaas (Espibos);
<emphasis id="B953EADEFFC8E428FB27F977FB2CF901" bold="true" box="[1183,1197,1697,1714]" pageId="46" pageNumber="539">G</emphasis>
,
<emphasis id="B953EADEFFC8E428FB01F977FB46F901" bold="true" box="[1209,1223,1697,1714]" pageId="46" pageNumber="539">H</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FB6BF977FAF5F901" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1235,1396,1697,1714]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="pelouatensis" status="n. sp." subGenus="Pterymarchia">
<emphasis id="B953EADEFFC8E428FB6BF977FB64F901" box="[1235,1253,1697,1714]" italics="true" pageId="46" pageNumber="539">P.</emphasis>
(
<emphasis id="B953EADEFFC8E428FB57F977FA83F901" box="[1263,1282,1697,1714]" italics="true" pageId="46" pageNumber="539">P.</emphasis>
)
<emphasis id="B953EADEFFC8E428FAB4F977FAF5F901" box="[1292,1396,1697,1714]" italics="true" pageId="46" pageNumber="539">pelouatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FAC2F977FA2FF901" box="[1402,1454,1697,1714]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFC8E428FF3CF961FF13F97B" bold="true" box="[132,146,1719,1736]" pageId="46" pageNumber="539">G</emphasis>
, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian;
<emphasis id="B953EADEFFC8E428FD72F961FD59F97B" bold="true" box="[714,728,1719,1736]" pageId="46" pageNumber="539">H</emphasis>
, paratype, MNHN.F.J17632, Saucats (Peloua);
<emphasis id="B953EADEFFC8E428FBDBF961FBE8F97B" bold="true" box="[1123,1129,1719,1736]" pageId="46" pageNumber="539">I</emphasis>
,
<emphasis id="B953EADEFFC8E428FBCBF961FBFCF97B" bold="true" box="[1139,1149,1719,1736]" pageId="46" pageNumber="539">J</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FB30F961FAF7F97B" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1160,1398,1719,1736]" class="Gastropoda" family="Muricidae" genus="Textiliomurex" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="chodmonensis" status="sp. nov.">
<emphasis id="B953EADEFFC8E428FB30F961FAF7F97B" box="[1160,1398,1719,1736]" italics="true" pageId="46" pageNumber="539">Textiliomurex chodmonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FAC4F961FA2DF97B" box="[1404,1452,1719,1736]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFC8E428FF3CF918FF0BF96C" bold="true" box="[132,138,1742,1759]" pageId="46" pageNumber="539">I</emphasis>
, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian;
<emphasis id="B953EADEFFC8E428FC75F918FC56F96C" bold="true" box="[973,983,1742,1759]" pageId="46" pageNumber="539">J</emphasis>
, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-enVexin (Carrière Darcy);
<emphasis id="B953EADEFFC8E428FEFBF933FED1F945" bold="true" box="[323,336,1765,1782]" pageId="46" pageNumber="539">K</emphasis>
,
<emphasis id="B953EADEFFC8E428FEE3F933FEE7F945" bold="true" box="[347,358,1765,1782]" pageId="46" pageNumber="539">L</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FEC8F933FE6CF945" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[368,493,1765,1782]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="remyi" status="sp. nov.">
<emphasis id="B953EADEFFC8E428FEC8F933FE6CF945" box="[368,493,1765,1782]" italics="true" pageId="46" pageNumber="539">Ponderia remyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FE4AF933FDA4F945" box="[498,549,1765,1782]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFC8E428FD93F933FDB9F945" bold="true" box="[555,568,1765,1782]" pageId="46" pageNumber="539">K</emphasis>
, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian;
<emphasis id="B953EADEFFC8E428FB88F933FBBAF945" bold="true" box="[1072,1083,1765,1782]" pageId="46" pageNumber="539">L</emphasis>
, paratype, MNHN.F.A90555, Fresville (Remy coll.);
<emphasis id="B953EADEFFC8E428FF00F92DFF48F8BF" bold="true" box="[184,201,1787,1804]" pageId="46" pageNumber="539">M</emphasis>
,
<emphasis id="B953EADEFFC8E428FF6CF92DFF63F8BF" bold="true" box="[212,226,1787,1804]" pageId="46" pageNumber="539">N</emphasis>
,
<taxonomicName id="4C274D4FFFC8E428FF56F92DFE56F8BF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[238,471,1787,1804]" class="Gastropoda" family="Muricidae" genus="Flexopteron" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="539" phylum="Mollusca" rank="species" species="constantinense" status="sp. nov.">
<emphasis id="B953EADEFFC8E428FF56F92DFE56F8BF" box="[238,471,1787,1804]" italics="true" pageId="46" pageNumber="539">Flexopteron constantinense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC8E428FE67F92DFD93F8BF" box="[479,530,1787,1804]" pageId="46" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B953EADEFFC8E428FDA2F92DFDAAF8BF" bold="true" box="[538,555,1787,1804]" pageId="46" pageNumber="539">M</emphasis>
, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian;
<emphasis id="B953EADEFFC8E428FAFFF92DFAD4F8BF" bold="true" box="[1351,1365,1787,1804]" pageId="46" pageNumber="539">N</emphasis>
, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL).
</paragraph>
</caption>
<subSubSection id="C33D6547FFC9E429FF3CFEB4FDA3FE27" pageId="47" pageNumber="540" type="etymology">
<paragraph id="8B9836CCFFC9E429FF3CFEB4FDA3FE27" blockId="47.[131,775,354,404]" pageId="47" pageNumber="540">
ETYMOLOGY. — From the Latin adjective
<taxonomicName id="4C274D4FFFC9E429FDA9FEB5FDEFFEC9" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[529,622,355,378]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFC9E429FDA9FEB5FDEFFEC9" box="[529,622,355,378]" italics="true" pageId="47" pageNumber="540">parisiensis</emphasis>
</taxonomicName>
, the geographic origin of this species being the Paris Basin.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC9E429FF3CFE7BFE25FD9A" pageId="47" pageNumber="540" type="materials_examined">
<paragraph id="8B9836CCFFC9E429FF3CFE7BFC89FE6D" blockId="47.[132,776,428,479]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FF3CFE7BFF34FE71" baselines="446,446" box="[132,181,429,452]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="47" pageNumber="540">
<typeStatus id="549C886EFFC9E429FF3CFE7BFF34FE71" box="[132,181,429,452]" pageId="47" pageNumber="540">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFC9E429FF04FE64FE96FE70" baselines="447" box="[188,279,434,451]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="47" pageNumber="540">HORIZON</smallCapsWord>
. — Sable du Guépelle (biozone NP16, see
<bibRefCitation id="EFB64B3DFFC9E429FD06FE7AFE89FE6C" author="GELY J. - P. &amp; LORENZ C." pageId="47" pageNumber="540" pagination="713 - 747" refId="ref44561" refString="GELY J. - P. &amp; LORENZ C. 1991. - Analyse sequentielle de l'Eocene et de l'Oligocene du Bassin parisien (France). Revue de l'institut francais du petrole 46 (6): 713 - 747." type="journal article" year="1991">Gély &amp; Lorenz (1991</bibRefCitation>
: pl. 1), middle Eocene (early Bartonian, Auversian).
</paragraph>
<paragraph id="8B9836CCFFC9E429FF3CFE21FE25FD9A" blockId="47.[132,775,502,553]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FF3CFE21FF36FDBF" baselines="520,520" box="[132,183,503,526]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="47" pageNumber="540">
<typeStatus id="549C886EFFC9E429FF3CFE21FF36FDBF" box="[132,183,503,526]" pageId="47" pageNumber="540">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFC9E429FF78FE2AFE9FFDBE" baselines="521" box="[192,286,508,525]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="47" pageNumber="540">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F330765CFFC9E429FEE8FE21FE17FDBD" box="[336,406,503,526]" name="France" pageId="47" pageNumber="540">France</collectingCountry>
, Paris Basin, Val dOise, Saint-Witz (Le Guépelle),
<bibRefCitation id="EFB64B3DFFC9E429FEAFFDC7FE21FD9A" author="INPN &amp; INVENTAIRE NATIONAL DU PATRIMOINE NATURELLE" box="[279,416,529,553]" pageId="47" pageNumber="540" refId="ref45200" refString="INPN (INVENTAIRE NATIONAL DU PATRIMOINE NATURELLE) 2024. - IDF 0034 - Le gisement bartonien de la carriere du Guepelle a Saint-Witz. Available at: https: // inpn. mnhn. fr / site / inpg / IDF 0034 / tab / interets (last access on 03 / 2024)." type="url" year="2024">INPN (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC9E429FF3CFD97FEC7FDC0" pageId="47" pageNumber="540" type="distribution">
<paragraph id="8B9836CCFFC9E429FF3CFD97FEC7FDC0" blockId="47.[132,775,576,627]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FF3CFD97FE93FDE4" baselines="594,595" box="[132,274,577,600]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="47" pageNumber="540">DISTRIBUTION</smallCapsWord>
. — Paris Basin (
<collectingCountry id="F330765CFFC9E429FE0BFD96FE78FDEB" box="[435,505,576,600]" name="France" pageId="47" pageNumber="540">France</collectingCountry>
), middle Eocene (early Bartonian, Auversian).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC9E429FF3CFD40FAEBFDC3" pageId="47" pageNumber="540" type="description">
<paragraph id="8B9836CCFFC9E429FF3CFD40FE94FD1D" blockId="47.[131,777,662,1453]" box="[132,277,662,688]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FF3CFD40FE94FD1D" baselines="681,681" box="[132,277,662,688]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="47" pageNumber="540">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFC9E429FF3CFD63FDF8FA1E" blockId="47.[131,777,662,1453]" pageId="47" pageNumber="540">
Smooth, multispiral protoconch of up to 2.5 whorls (see
<bibRefCitation id="EFB64B3DFFC9E429FF3EFD03FE86FD43" author="MERLE D." box="[134,263,725,752]" pageId="47" pageNumber="540" refId="ref46629" refString="MERLE D. 1999. - La radiation des Muricidae (Gastropoda: Neogastropoda) au Paleogene: approche phylogenetique et evolutive. Unpublished PhD thesis, Museum national d'Histoire naturelle, Paris, 499 p." type="book" year="1999">
<taxonomicName id="4C274D4FFFC9E429FF3EFD03FF47FD5C" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[134,198,725,751]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
1999
</bibRefCitation>
: pl. 23, fig. 5,
<typeStatus id="549C886EFFC9E429FE0BFD03FD98FD5C" box="[435,537,725,751]" pageId="47" pageNumber="540" type="holotype">holotype</typeStatus>
). Biconic teleoconch, up to
<quantity id="4CDF9B29FFC9E429FF73FD23FEB0FCA3" box="[203,305,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="47" pageNumber="540" unit="mm" value="21.5">21.5 mm</quantity>
in height, up to
<quantity id="4CDF9B29FFC9E429FE49FD23FDC3FCA3" box="[497,578,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="47" pageNumber="540" unit="mm" value="14.0">14 mm</quantity>
in width, biconic in profile and composed of six whorls. Moderately high spire with rounded or subcarinate whorls. Last whorl up to 73% of total length. Apical angle 64°. Spiral sculpture with strongly marked primary cords. First whorl: appearance of P1, s1, P2; P1 less marked than s1-P2; from second to third whorl, no addition of cord, P1 of equal strength to s1-P2; on fourth whorl, appearance of IP poorly marked. Fifth whorl: no addition of cord. Last whorl: IP (weak), P1 to P6, P6 not atrophied (on convex part of whorl), ADP, MP and ABP; s1 to s3 dominant, equal in strength to primary cords. First to second whorl: twelve varices. Third to fourth whorl: eleven varices. Fifth whorl: ten varices. Sixth whorl: nine varices. Varices more or less sublamellose. No true spine except angular point on P1. Suboval aperture up to 31% of diameter and up to 66% of length of last whorl (including siphonal canal). Columella slightly angular. Columellar lip narrow, adherent, smooth or with one tubercle. Outer lip with five or six denticles including D1 to D6. D1 appearing late in ontogeny. D6 occasional. ID missing. Siphonal canal, short, open, up to 31% of aperture length, slightly dorsally recurved. Pseudo-umbilicus deep, rather wide.
</paragraph>
<paragraph id="8B9836CCFFC9E429FF3CFA05FE9EFA58" blockId="47.[130,777,1491,2028]" box="[132,287,1491,1517]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FF3CFA05FE9EFA58" baselines="1510,1510" box="[132,287,1491,1517]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="47" pageNumber="540">COMPARISONS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFC9E429FF3CFA25FAEBFDC3" blockId="47.[130,777,1491,2028]" lastBlockId="47.[811,1457,215,624]" pageId="47" pageNumber="540">
<taxonomicName id="4C274D4FFFC9E429FF3CFA25FE3BF9BE" authority="Merle, 2005 c" authorityName="Merle" authorityYear="2005" box="[132,442,1522,1549]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="ponsi">
<emphasis id="B953EADEFFC9E429FF3CFA25FEAAF9BF" box="[132,299,1523,1549]" italics="true" pageId="47" pageNumber="540">Nucellopsis ponsi</emphasis>
<bibRefCitation id="EFB64B3DFFC9E429FE96FA25FE3BF9BE" author="MERLE D." box="[302,442,1522,1549]" pageId="47" pageNumber="540" pagination="177 - 189" refId="ref46903" refString="MERLE D. 2005 c. - Nucellopsis (Gastropoda, Muricidae), a new genus from the paralic domain of the European Palaeogene. Comptes Rendus Palevol 4 (1 - 2): 177 - 189. https: // doi. org / 10.1016 / j. crpv. 2004.11.005" type="journal article" year="2005">Merle, 2005c</bibRefCitation>
</taxonomicName>
differs from
<taxonomicName id="4C274D4FFFC9E429FD85FA25FD43F9BF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[573,706,1523,1548]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis" status="n. sp.">
<emphasis id="B953EADEFFC9E429FD85FA25FD43F9BF" box="[573,706,1523,1548]" italics="true" pageId="47" pageNumber="540">N. parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC9E429FD71FA25FC83F9BE" box="[713,770,1523,1549]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C274D4FFFC9E429FF3CF9C5FE4BF99E" ID-CoL="8748G" authority="(Deshayes, 1835)" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[132,458,1554,1581]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="plicatilis">
<emphasis id="B953EADEFFC9E429FF3CF9C5FE80F99F" box="[132,257,1554,1580]" italics="true" pageId="47" pageNumber="540">N. plicatilis</emphasis>
(
<bibRefCitation id="EFB64B3DFFC9E429FEAAF9C4FE42F99E" author="DESHAYES G. - P." box="[274,451,1554,1581]" pageId="47" pageNumber="540" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel &amp; Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC9E429FDBDF9C5FF44F9FF" ID-CoL="86ZBQ" authority="(Strauzs, 1966)" baseAuthorityName="Strauzs" baseAuthorityYear="1966" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="dudariensis">
<emphasis id="B953EADEFFC9E429FDBDF9C5FD21F99F" box="[517,672,1554,1580]" italics="true" pageId="47" pageNumber="540">N. dudariensis</emphasis>
(Strauzs, 1966)
</taxonomicName>
by having a deeper posterior sinus and weaker spiral cords. Its columellar lip is occasionally ornamented by a weak denticle mid-columella, whereas it is usually smooth in the other three species.
<taxonomicName id="4C274D4FFFC9E429FEE3F944FDBDF91F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[347,572,1682,1708]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B953EADEFFC9E429FEE3F944FDBDF91F" box="[347,572,1682,1708]" italics="true" pageId="47" pageNumber="540">Nucellopsis parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC9E429FDFBF945FD00F91E" box="[579,641,1683,1709]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C274D4FFFC9E429FD35F945FC86F91F" ID-CoL="8748G" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[653,775,1682,1708]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="plicatilis">
<emphasis id="B953EADEFFC9E429FD35F945FC86F91F" box="[653,775,1682,1708]" italics="true" pageId="47" pageNumber="540">N. plicatilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC9E429FF09F965FEC4F97F" ID-CoL="86ZBQ" baseAuthorityName="Strauzs" baseAuthorityYear="1966" box="[177,325,1714,1740]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="dudariensis">
<emphasis id="B953EADEFFC9E429FF09F965FEC4F97F" box="[177,325,1714,1740]" italics="true" pageId="47" pageNumber="540">N. dudariensis</emphasis>
</taxonomicName>
bear more numerous cords on the siphonal canal (MP and ABP) and more secondary cords than
<taxonomicName id="4C274D4FFFC9E429FD11F905FD7FF95F" authorityName="Merle" authorityYear="2005" box="[681,766,1747,1772]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="ponsi">
<emphasis id="B953EADEFFC9E429FD11F905FD7FF95F" box="[681,766,1747,1772]" italics="true" pageId="47" pageNumber="540">N. ponsi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C274D4FFFC9E429FF3CF924FEE7F8BF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[132,358,1778,1804]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B953EADEFFC9E429FF3CF924FEE7F8BF" box="[132,358,1778,1804]" italics="true" pageId="47" pageNumber="540">Nucellopsis parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC9E429FEC9F924FE2FF8BF" box="[369,430,1778,1804]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C274D4FFFC9E429FD87F925FD3AF8BF" ID-CoL="8748G" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[575,699,1778,1804]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="plicatilis">
<emphasis id="B953EADEFFC9E429FD87F925FD3AF8BF" box="[575,699,1778,1804]" italics="true" pageId="47" pageNumber="540">N. plicatilis</emphasis>
</taxonomicName>
by the appearance of P1 and s1 on the first teleoconch whorls, by having more angular teleoconch whorls, and by usually lacking D6. Both species share a multispiral protoconch.
<taxonomicName id="4C274D4FFFC9E429FD2DF887FF6DF838" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis" status="sp. nov.">
<emphasis id="B953EADEFFC9E429FD2DF887FF6DF838" italics="true" pageId="47" pageNumber="540">Nucellopsis parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC9E429FF4BF8A4FEAFF83F" box="[243,302,1906,1932]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
was erroneously identified as
<taxonomicName id="4C274D4FFFC9E429FDD7F8A7FE23F81F" ID-CoL="8YMTF" authority="(Deshayes, 1865)" baseAuthorityName="Deshayes" baseAuthorityYear="1865" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="depauperata">
<emphasis id="B953EADEFFC9E429FDD7F8A7FF5FF818" italics="true" pageId="47" pageNumber="540">Jsowerbya depauperata</emphasis>
(
<bibRefCitation id="EFB64B3DFFC9E429FF55F847FE1BF81F" author="DESHAYES G. - P." box="[237,410,1937,1964]" pageId="47" pageNumber="540" pagination="101 - 107" refId="ref44001" refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]." type="journal article" year="1865">Deshayes, 1865</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="EFB64B3DFFC9E429FE73F847FC86F818" author="COSSMANN M. &amp; PISSARRO G." box="[459,775,1937,1963]" pageId="47" pageNumber="540" pagination="26 - 45" refId="ref43883" refString="COSSMANN M. &amp; PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene du Bassin de Paris. Tome 2. Hermann, Paris: pl. 26 - 45. https: // www. biodiversitylibrary. org / page / 36319938" type="book chapter" year="1911">Cossmann &amp; Pissarro (1911)</bibRefCitation>
in spite of numerous differences, such as a narrower shell shape, IP on the sutural ramp and a deeper anal sulcus (
<bibRefCitation id="EFB64B3DFFC9E429FD70F807FCF2FF41" author="MERLE D." box="[712,883,215,2027]" pageId="47" pageNumber="540" pagination="505 - 517" refId="ref46835" refString="MERLE D. 2005 b. - Jsowerbya, new genus of Muricidae (Mollusca Gastropoda) from the Eocene of the Paris (France) and Hampshire basins (England), with a phylogenetic assessment of its ocenebrine versus ergalataxine affinities. Geobios 38 (4): 505 - 517. https: // doi. org / 10.1016 / j. geobios. 2004.02.002" type="journal article" year="2005">
<taxonomicName id="4C274D4FFFC9E429FD70F807FC86F858" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[712,775,2001,2027]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
2005b
</bibRefCitation>
).
<bibRefCitation id="EFB64B3DFFC9E429FC3FFF01FB8BFF42" author="MERLE D." box="[903,1034,215,241]" pageId="47" pageNumber="540" refId="ref46629" refString="MERLE D. 1999. - La radiation des Muricidae (Gastropoda: Neogastropoda) au Paleogene: approche phylogenetique et evolutive. Unpublished PhD thesis, Museum national d'Histoire naturelle, Paris, 499 p." type="book" year="1999">
<taxonomicName id="4C274D4FFFC9E429FC3FFF01FC47FF42" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[903,966,215,241]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1999
</bibRefCitation>
,
<bibRefCitation id="EFB64B3DFFC9E429FBAFFF01FBDBFF41" author="MERLE D." box="[1047,1114,215,242]" pageId="47" pageNumber="540" pagination="177 - 189" refId="ref46903" refString="MERLE D. 2005 c. - Nucellopsis (Gastropoda, Muricidae), a new genus from the paralic domain of the European Palaeogene. Comptes Rendus Palevol 4 (1 - 2): 177 - 189. https: // doi. org / 10.1016 / j. crpv. 2004.11.005" type="journal article" year="2005">2005c</bibRefCitation>
) identified
<taxonomicName id="4C274D4FFFC9E429FB68FF0EFAD6FF42" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1232,1367,216,241]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis" status="n. sp.">
<emphasis id="B953EADEFFC9E429FB68FF0EFAD6FF42" box="[1232,1367,216,241]" italics="true" pageId="47" pageNumber="540">N. parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC9E429FAE4FF0EFA14FF41" box="[1372,1429,216,242]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
as
<taxonomicName id="4C274D4FFFC9E429FC93FF2EFC41FEA2" ID-CoL="86ZBQ" baseAuthorityName="Strauzs" baseAuthorityYear="1966" box="[811,960,247,273]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="dudariensis">
<emphasis id="B953EADEFFC9E429FC93FF2EFC41FEA2" box="[811,960,247,273]" italics="true" pageId="47" pageNumber="540">N. dudariensis</emphasis>
</taxonomicName>
due to several similarities with the
<typeStatus id="549C886EFFC9E429FA8CFF21FA12FEA2" box="[1332,1427,247,273]" pageId="47" pageNumber="540" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D4FFFC9E429FC93FEC1FBE9FE82" box="[811,1128,279,305]" class="Gastropoda" family="Pisaniidae" genus="Cantharus" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="dudariensis" subGenus="Pollia">
<emphasis id="B953EADEFFC9E429FC93FEC1FC19FE82" box="[811,920,279,305]" italics="true" pageId="47" pageNumber="540">Cantharus</emphasis>
(
<emphasis id="B953EADEFFC9E429FC11FEC1FC64FE82" box="[937,997,279,305]" italics="true" pageId="47" pageNumber="540">Pollia</emphasis>
)
<emphasis id="B953EADEFFC9E429FC4CFEC1FBE9FE82" box="[1012,1128,279,305]" italics="true" pageId="47" pageNumber="540">dudariensis</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFC9E429FBC0FEC1FB7CFE82" box="[1144,1277,279,305]" captionStart="FIG" captionStartId="42.[132,143,1281,1298]" captionTargetBox="[132,1455,215,1223]" captionTargetId="figure-489@42.[143,795,242,771]" captionTargetPageId="42" captionText="FIG. 23. — Spiral cords and internal denticles of the outer lip in Nucellopsis parisiensis n. sp. (A-F) and N. dudariensis (Strauzs, 1966) (G, H): A, B, paratype, MNHN.F.A90964 (Faullummel coll.), Saint-Witz (Le Guépelle), Val dOise, Paris Basin, France, middle Eocene (Bartonian); C, D, paratype, MNHN.F.A90963 (Faullummel coll.),Saint-Witz (Le Guépelle);E, F, holotype,MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle);G, H, holotype of Cantharus (Pollia) dudariensis, MGSH E.467, Dudar, Hungary, middle Eocene (upper Lutetian). Scale bars: 5 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E, F, D. Serrette (MNHN); G, H, courtesy of Z. Kovács." figureDoi="http://doi.org/10.5281/zenodo.13985819" httpUri="https://zenodo.org/record/13985819/files/figure.png" pageId="47" pageNumber="540">Fig. 23G, H</figureCitation>
) from the upper Lutetian of
<collectingCountry id="F330765CFFC9E429FC1BFEE1FB80FEE2" box="[931,1025,311,337]" name="Hungary" pageId="47" pageNumber="540">Hungary</collectingCountry>
[apical angle near 64°, subangular whorls, thick varices, poorly developed shoulder sinus, five internal denticles (D1 to D5, D6 missing), angular columella with one small tubercle]. These similarities suggest that
<taxonomicName id="4C274D4FFFC9E429FA9DFE41FA31FE03" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1317,1456,407,432]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="parisiensis" status="n. sp.">
<emphasis id="B953EADEFFC9E429FA9DFE41FA31FE03" box="[1317,1456,407,432]" italics="true" pageId="47" pageNumber="540">N. parisiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC9E429FC96FE61FCE8FE62" box="[814,873,439,465]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFC9E429FC19FE61FBB9FE63" box="[929,1080,438,464]" class="Gastropoda" family="Pisaniidae" genus="Cantharus" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="dudariensis">
<emphasis id="B953EADEFFC9E429FC19FE61FBB9FE63" box="[929,1080,438,464]" italics="true" pageId="47" pageNumber="540">N. dudariensis</emphasis>
</taxonomicName>
are closely related. However, close examinaton of the
<typeStatus id="549C886EFFC9E429FC41FE00FBD9FE43" box="[1017,1112,470,496]" pageId="47" pageNumber="540" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D4FFFC9E429FBC2FE01FA90FE43" box="[1146,1297,470,496]" class="Gastropoda" family="Pisaniidae" genus="Cantharus" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="dudariensis">
<emphasis id="B953EADEFFC9E429FBC2FE01FA90FE43" box="[1146,1297,470,496]" italics="true" pageId="47" pageNumber="540">N. dudariensis</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFC9E429FA99FE00FA26FE43" box="[1313,1447,470,496]" captionStart="FIG" captionStartId="42.[132,143,1281,1298]" captionTargetBox="[132,1455,215,1223]" captionTargetId="figure-489@42.[143,795,242,771]" captionTargetPageId="42" captionText="FIG. 23. — Spiral cords and internal denticles of the outer lip in Nucellopsis parisiensis n. sp. (A-F) and N. dudariensis (Strauzs, 1966) (G, H): A, B, paratype, MNHN.F.A90964 (Faullummel coll.), Saint-Witz (Le Guépelle), Val dOise, Paris Basin, France, middle Eocene (Bartonian); C, D, paratype, MNHN.F.A90963 (Faullummel coll.),Saint-Witz (Le Guépelle);E, F, holotype,MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle);G, H, holotype of Cantharus (Pollia) dudariensis, MGSH E.467, Dudar, Hungary, middle Eocene (upper Lutetian). Scale bars: 5 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E, F, D. Serrette (MNHN); G, H, courtesy of Z. Kovács." figureDoi="http://doi.org/10.5281/zenodo.13985819" httpUri="https://zenodo.org/record/13985819/files/figure.png" pageId="47" pageNumber="540">Fig. 23G, H</figureCitation>
) reveals several differences with the Parisian species, such as the late appearance of s1 on the last whorl, s1 less coarse than the primary cords P1 and P2, fewer secondary cords (only s1 instead s1 to s3), and the stronger development of IP.
</paragraph>
</subSubSection>
</treatment>
</document>