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<document id="DBB810AF21A4D98972FB8A73117DED92" ID-CLB-Dataset="36709" ID-DOI="10.11646/zootaxa.4227.4.4" ID-GBIF-Dataset="69b17c45-2ce2-42ba-b023-0b17f3b83fb4" ID-ISSN="1175-5326" ID-Zenodo-Dep="293613" ID-ZooBank="D71F8124-BD24-4FDC-A659-9ECC496150BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1486670234657" checkinUser="plazi" docAuthor="Okanishi, Masanori" docDate="2017" docId="038E87A04E78D36AFF7FFE4AE229F8EE" docLanguage="en" docName="data.D8284B1967D88C5941EF86CB439E2B88.cached" docOrigin="Zootaxa 4227 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asteroporpa (Astrohelix) Doderlein 1930" docType="treatment" docVersion="9" lastPageNumber="544" masterDocId="FFB7FFD84E79D36BFFE8FFFFE11FFF8C" masterDocTitle="A taxonomic review of the genus Astrohelix Döderlein, 1930 including the synonymy of the subgenus Asteroporpa (Astromoana) Baker, 1980 to Astrohelix" masterLastPageNumber="553" masterPageNumber="543" pageNumber="544" updateTime="1698436741822" updateUser="ExternalLinkService">
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<mods:title id="A3088DDD074C449C48CCA9638FEBB37D">A taxonomic review of the genus Astrohelix Döderlein, 1930 including the synonymy of the subgenus Asteroporpa (Astromoana) Baker, 1980 to Astrohelix</mods:title>
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<mods:namePart id="02014BD4F59CDB1DBDA3BB689125758D">Okanishi, Masanori</mods:namePart>
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<mods:date id="F9A854C223B173AA645FFB4D23E00A2A">2017</mods:date>
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<mods:identifier id="2E834C7C918A8592F5216190F1A81105" type="DOI">10.11646/zootaxa.4227.4.4</mods:identifier>
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<treatment id="038E87A04E78D36AFF7FFE4AE229F8EE" ID-DOI="http://doi.org/10.5281/zenodo.5688476" ID-GBIF-Taxon="127678115" ID-Zenodo-Dep="5688476" LSID="urn:lsid:plazi:treatment:038E87A04E78D36AFF7FFE4AE229F8EE" httpUri="http://treatment.plazi.org/id/038E87A04E78D36AFF7FFE4AE229F8EE" lastPageNumber="544" pageId="1" pageNumber="544">
<subSubSection id="C33D653D4E78D36AFF7FFE4AE38FFE43" box="[151,656,437,463]" pageId="1" pageNumber="544" type="nomenclature">
<paragraph id="8B9836B64E78D36AFF7FFE4AE38FFE43" blockId="1.[151,656,437,463]" box="[151,656,437,463]" pageId="1" pageNumber="544">
<heading id="D0D081DA4E78D36AFF7FFE4AE38FFE43" bold="true" box="[151,656,437,463]" fontSize="11" level="1" pageId="1" pageNumber="544" reason="1">
<taxonomicName id="4C274D354E78D36AFF7FFE4AE38FFE43" authority="Doderlein, 1930" authorityName="Doderlein" authorityYear="1930" box="[151,656,437,463]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subGenus" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFF7FFE4AE38FFE43" bold="true" box="[151,656,437,463]" pageId="1" pageNumber="544">
<emphasis id="B953EAA44E78D36AFF7FFE4AE034FE43" bold="true" box="[151,299,437,463]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFED4FE4AE0A6FE43" bold="true" box="[316,441,437,463]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<bibRefCitation id="EFB64B474E78D36AFE23FE4AE38FFE43" author="Doderlein" box="[459,656,437,463]" pageId="1" pageNumber="544" refString="Doderlein, L. (1930) Die Ophiuroiden der deutschen Tiefsee-Expedition. 2. Euryalae. Deutsche Tiefsee-Expedition 1898 - 1899, 22, 347 - 396." type="journal article" year="1930">Döderlein, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D653D4E78D36AFF7FFE03E399FDBC" pageId="1" pageNumber="544" type="reference_group">
<paragraph id="8B9836B64E78D36AFF7FFE03E399FDBC" blockId="1.[151,1151,508,560]" pageId="1" pageNumber="544">
<treatmentCitationGroup id="AB3711984E78D36AFF7FFE03E399FDBC" pageId="1" pageNumber="544">
<taxonomicName id="4C274D354E78D36AFF7FFE03E351FD9E" authority="Baker, 1980: 36" authorityName="Baker" authorityPageNumber="36" authorityYear="1980" box="[151,590,508,530]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subGenus" subGenus="Astromoana">
<emphasis id="B953EAA44E78D36AFF7FFE03E00AFD9D" box="[151,277,508,529]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFECBFE03E081FD9D" box="[291,414,508,529]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
<treatmentCitation id="0A8610A74E78D36AFE44FE03E351FD9E" author="Baker" box="[428,590,508,530]" page="36" pageId="1" pageNumber="544" year="1980">
<bibRefCitation id="EFB64B474E78D36AFE44FE03E335FD9E" author="Baker" box="[428,554,508,530]" pageId="1" pageNumber="544" refString="Baker, A. N. (1980) Euryalinid Ophiuroidea (Echinodermata) from Australia, New Zealand, and the south-west Pacific Ocean. New Zealand Journal of Zoology, 7, 11 - 83." type="journal article" year="1980">Baker, 1980</bibRefCitation>
: 36
</treatmentCitation>
</taxonomicName>
–38;
<treatmentCitation id="0A8610A74E78D36AFD69FE03E24DFD9E" author="McKnight" box="[641,850,508,530]" page="57" pageId="1" pageNumber="544" year="2000">
<bibRefCitation id="EFB64B474E78D36AFD69FE03E232FD9E" author="McKnight" box="[641,813,508,530]" pageId="1" pageNumber="544" refString="McKnight, D. G. (2000) The Marine Fauna of New Zealand: Basket-stars and snake-stars (Echinodermata: Ophiuroidea: Euryalinida). National Institute of Water and Atmospheric Research Biodiversity Memoir, 115, 1 - 79." type="journal article" year="2000">McKnight, 2000</bibRefCitation>
: 57
</treatmentCitation>
; Okanishi &amp; Fujita, 2011: 27.
<emphasis id="B953EAA44E78D36AFF7FFDE5E003FDBC" bold="true" box="[151,284,538,560]" pageId="1" pageNumber="544">Type species</emphasis>
:
<taxonomicName id="4C274D354E78D36AFEC2FDE4E39DFDBC" ID-CoL="ec1013fa-9148-4a5b-ae61-f8c5b3359542" authority="Koehler, 1904" authorityName="Koehler" authorityYear="1904" box="[298,642,538,560]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrotoma" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="bellator">
<emphasis id="B953EAA44E78D36AFEC2FDE4E0F6FDBC" box="[298,489,538,560]" italics="true" pageId="1" pageNumber="544">Astrotoma bellator</emphasis>
<bibRefCitation id="EFB64B474E78D36AFE07FDE5E39DFDBC" author="Koehler" box="[495,642,538,560]" pageId="1" pageNumber="544" refString="Koehler, R. (1904) Ophiures de l'Expedition du Siboga. Part I. Ophiures de Mer profonde. Siboga-Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33D653D4E78D36AFF7FFDA2E571FC89" pageId="1" pageNumber="544" type="diagnosis">
<paragraph id="8B9836B64E78D36AFF7FFDA2E571FC89" blockId="1.[151,1437,605,1890]" pageId="1" pageNumber="544">
<emphasis id="B953EAA44E78D36AFF7FFDA2E015FDFA" bold="true" box="[151,266,605,630]" pageId="1" pageNumber="544">Diagnosis</emphasis>
. No concentric ridges or furrows exist on aboral disc. The hooklet-bearing plates occur on the aboral periphery of the disc. Each hooklet bears one to three secondary teeth. Except for the hooklet-bearing plates, body is entirely covered by round granule-shaped epidermal ossicles, round or polygonal plate-shaped epidermal ossicles, and cone-shaped epidermal ossicles with one to five terminal projections. Teeth are acute, spiniform. Three to six arm spines are adjacent to each tentacle pore in middle portion of the arms.
</paragraph>
</subSubSection>
<subSubSection id="C33D653D4E78D36AFF2FFCEEE075FA02" pageId="1" pageNumber="544" type="discussion">
<paragraph id="8B9836B64E78D36AFF2FFCEEE075FA02" blockId="1.[151,1437,605,1890]" pageId="1" pageNumber="544">
<emphasis id="B953EAA44E78D36AFF2FFCEEE024FCA6" bold="true" box="[199,315,785,810]" pageId="1" pageNumber="544">Remarks.</emphasis>
The genus
<taxonomicName id="4C274D354E78D36AFE23FCEEE322FCA5" box="[459,573,785,809]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFE23FCEEE322FCA5" box="[459,573,785,809]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
is morphologically similar to the subgenus
<taxonomicName id="4C274D354E78D36AFBA1FCEDE465FCA6" authority="(Astromoana)" baseAuthorityName="Astromoana" box="[1097,1402,786,810]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFBA1FCEDE5CBFCA5" box="[1097,1236,786,809]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFB01FCEDE46EFCA5" box="[1257,1393,786,809]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
</taxonomicName>
in having no conspicuous ridges on the aboral disc; epidermal ossicles with terminal projections on disc and basal arms; five arm spines in basal portion of arms; hooklets with one or two inner teeth; and small genital slits (
<bibRefCitation id="EFB64B474E78D36AFF76FC82E05DFC1A" author="Koehler" box="[158,322,893,918]" pageId="1" pageNumber="544" refString="Koehler, R. (1904) Ophiures de l'Expedition du Siboga. Part I. Ophiures de Mer profonde. Siboga-Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
;
<bibRefCitation id="EFB64B474E78D36AFEA5FC82E318FC1A" author="Doderlein" box="[333,519,893,918]" pageId="1" pageNumber="544" refString="Doderlein, L. (1930) Die Ophiuroiden der deutschen Tiefsee-Expedition. 2. Euryalae. Deutsche Tiefsee-Expedition 1898 - 1899, 22, 347 - 396." type="journal article" year="1930">Döderlein, 1930</bibRefCitation>
;
<bibRefCitation id="EFB64B474E78D36AFDFAFC81E3CCFC1A" author="Mortensen" box="[530,723,894,918]" pageId="1" pageNumber="544" refString="Mortensen, T. (1933) Studies of Indo-Pacific euryalids. Fidenskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn, 96, 1 - 75." type="journal article" year="1933">Mortensen, 1933</bibRefCitation>
). However, the presence of hooklet-bearing plates on the aboral disc, which is a clear and distinctive diagnostic character of the genus
<taxonomicName id="4C274D354E78D36AFC38FC5DE487FC36" authority="Baker, 1980" authorityName="Baker" authorityYear="1980" box="[976,1432,929,954]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subGenus" subGenus="Astromoana">
<emphasis id="B953EAA44E78D36AFC38FC5DE544FC35" box="[976,1115,930,953]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFB87FC5DE5E8FC35" box="[1135,1271,930,953]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
<bibRefCitation id="EFB64B474E78D36AFAE2FC5EE487FC36" author="Baker" box="[1290,1432,929,954]" pageId="1" pageNumber="544" refString="Baker, A. N. (1980) Euryalinid Ophiuroidea (Echinodermata) from Australia, New Zealand, and the south-west Pacific Ocean. New Zealand Journal of Zoology, 7, 11 - 83." type="journal article" year="1980">Baker, 1980</bibRefCitation>
</taxonomicName>
, have not been previously described in
<taxonomicName id="4C274D354E78D36AFD90FC3AE5A1FC52" authority="Mortensen, 1933" authorityName="Mortensen" authorityYear="1933" box="[632,1214,965,990]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subSpecies" species="bellator" subSpecies="paucidens">
<emphasis id="B953EAA44E78D36AFD90FC3AE2CBFC51" box="[632,980,965,989]" italics="true" pageId="1" pageNumber="544">Astrohelix bellator paucidens</emphasis>
(
<bibRefCitation id="EFB64B474E78D36AFC02FC39E5A9FC52" author="Mortensen" box="[1002,1206,966,990]" pageId="1" pageNumber="544" refString="Mortensen, T. (1933) Studies of Indo-Pacific euryalids. Fidenskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn, 96, 1 - 75." type="journal article" year="1933">Mortensen, 1933</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="EFB64B474E78D36AFB3AFC3AE483FC52" author="Doderlein" box="[1234,1436,965,990]" pageId="1" pageNumber="544" refString="Doderlein, L. (1930) Die Ophiuroiden der deutschen Tiefsee-Expedition. 2. Euryalae. Deutsche Tiefsee-Expedition 1898 - 1899, 22, 347 - 396." type="journal article" year="1930">Döderlein (1930)</bibRefCitation>
redescribed
<typeStatus id="549C88144E78D36AFEC8FC16E09DFB8E" box="[288,386,1001,1026]" pageId="1" pageNumber="544" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D354E78D36AFE43FC15E39FFB8D" box="[427,640,1001,1025]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrotoma" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="bellator">
<emphasis id="B953EAA44E78D36AFE43FC15E39FFB8D" box="[427,640,1001,1025]" italics="true" pageId="1" pageNumber="544">Astrotoma bellator</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C274D354E78D36AFD49FC16E2C9FB8D" box="[673,982,1001,1025]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subSpecies" species="bellator" subSpecies="bellator">
<emphasis id="B953EAA44E78D36AFD49FC16E2C9FB8D" box="[673,982,1001,1025]" italics="true" pageId="1" pageNumber="544">Astrohelix bellator bellator</emphasis>
</taxonomicName>
) and noted the presence of the hookletbearing plates on disc, but he did not include it as a diagnostic character of his erected genus
<emphasis id="B953EAA44E78D36AFB19FBF2E474FBA9" box="[1265,1387,1037,1061]" italics="true" pageId="1" pageNumber="544">
<taxonomicName id="4C274D354E78D36AFB19FBF2E478FBA9" box="[1265,1383,1037,1061]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">Astrohelix</taxonomicName>
.
</emphasis>
My examination of all known specimens of
<taxonomicName id="4C274D354E78D36AFD8BFBCEE3CAFBC5" box="[611,725,1073,1097]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFD8BFBCEE3CAFBC5" box="[611,725,1073,1097]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
revealed that they certainly possess hooklet-bearing plates on the aboral periphery of the disc (
<figureCitation id="131C2A334E78D36AFDF9FBAAE349FBE2" box="[529,598,1109,1134]" captionStart="FIGURE 1" captionStartId="3.[151,250,1877,1899]" captionTargetBox="[226,1359,193,1854]" captionTargetId="figure@3.[226,1360,193,1855]" captionTargetPageId="3" captionText="FIGURE 1. Holotype (ZMA E 2044) of Astrotoma bellator Koehler, 1904 (= Asteroporpa (Astrohelix) bellator (Koehler, 1904 )). A, aboral disc; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, cone-shaped epidermal ossicles on aboral center part of the disc; E, aboral periphery of the disc; F, aboral basal portion of the arm; G, jaws; H, oral peripheral part of the disc. Arrow heads indicate hooklets (E), transverse rows of hooklet-bearing plates (F) and cone-shaped epidermal ossicles (G, H). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/306380/files/figure.png" pageId="1" pageNumber="544">Figs 1</figureCitation>
A, E; 3C, E). Therefore, there are no morphological differences between
<taxonomicName id="4C274D354E78D36AFF7FFB86E0D9FB1E" authority="Doderlein, 1930" authorityName="Doderlein" authorityYear="1930" box="[151,454,1145,1170]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFF7FFB86E016FB1D" box="[151,265,1145,1169]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
<bibRefCitation id="EFB64B474E78D36AFEF8FB86E0D9FB1E" author="Doderlein" box="[272,454,1145,1170]" pageId="1" pageNumber="544" refString="Doderlein, L. (1930) Die Ophiuroiden der deutschen Tiefsee-Expedition. 2. Euryalae. Deutsche Tiefsee-Expedition 1898 - 1899, 22, 347 - 396." type="journal article" year="1930">Döderlein, 1930</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C274D354E78D36AFE15FB85E2A3FB1E" authority="Baker, 1980" authorityName="Baker" authorityYear="1980" box="[509,956,1145,1170]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subGenus" subGenus="Astromoana">
<emphasis id="B953EAA44E78D36AFE15FB85E397FB1D" box="[509,648,1146,1169]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFD71FB85E23EFB1D" box="[665,801,1146,1169]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
<bibRefCitation id="EFB64B474E78D36AFCD9FB86E2A3FB1E" author="Baker" box="[817,956,1145,1170]" pageId="1" pageNumber="544" refString="Baker, A. N. (1980) Euryalinid Ophiuroidea (Echinodermata) from Australia, New Zealand, and the south-west Pacific Ocean. New Zealand Journal of Zoology, 7, 11 - 83." type="journal article" year="1980">Baker, 1980</bibRefCitation>
</taxonomicName>
, and I here propose the synonymy of these two taxa. On the molecular tree in
<bibRefCitation id="EFB64B474E78D36AFDD6FB62E275FB3A" author="Okanishi" box="[574,874,1181,1206]" pageId="1" pageNumber="544" refString="Okanishi, M. &amp; Fujita, T. (2013) Molecular phylogeny based on increased number of species and genes revealed more robust family-level systematics of the order Euryalida (Echinodermata: Ophiuroidea). Molecular Phylogenetics and Evolution, 69, 566 - 580." type="journal article" year="2013">Okanishi &amp; Fujita (2013)</bibRefCitation>
,
<taxonomicName id="4C274D354E78D36AFC93FB61E455FB39" box="[891,1354,1181,1206]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="australiensis" subGenus="Asteroporpa">
<emphasis id="B953EAA44E78D36AFC93FB61E519FB39" box="[891,1030,1182,1205]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFBF3FB61E5B9FB39" box="[1051,1190,1182,1205]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
)
<emphasis id="B953EAA44E78D36AFB53FB62E455FB39" box="[1211,1354,1181,1205]" italics="true" pageId="1" pageNumber="544">australiensis</emphasis>
</taxonomicName>
sitting between
<taxonomicName id="4C274D354E78D36AFF14FB3DE334FB55" box="[252,555,1217,1242]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="reticulata" subGenus="Astromoana">
<emphasis id="B953EAA44E78D36AFF14FB3DE00BFB55" box="[252,276,1218,1241]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFECEFB3DE0B1FB55" box="[294,430,1218,1241]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
<emphasis id="B953EAA44E78D36AFE57FB3EE334FB55" box="[447,555,1217,1241]" italics="true" pageId="1" pageNumber="544">reticulata</emphasis>
</taxonomicName>
(or
<taxonomicName id="4C274D354E78D36AFDB4FB3EE25DFB55" box="[604,834,1217,1241]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B953EAA44E78D36AFDB4FB3EE25DFB55" box="[604,834,1217,1241]" italics="true" pageId="1" pageNumber="544">Astrohelix reticulata</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C274D354E78D36AFC6BFB3DE5E6FB55" box="[899,1273,1217,1242]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="muricatopatella" subGenus="Astromoana">
<emphasis id="B953EAA44E78D36AFC6BFB3DE283FB55" box="[899,924,1218,1241]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFC45FB3DE52AFB55" box="[941,1077,1218,1241]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
<emphasis id="B953EAA44E78D36AFBAEFB3EE5E6FB55" box="[1094,1273,1217,1241]" italics="true" pageId="1" pageNumber="544">muricatopatella</emphasis>
</taxonomicName>
(or
<taxonomicName id="4C274D354E78D36AFAC2FB3EE055FB71" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="muricatopatella">
<emphasis id="B953EAA44E78D36AFAC2FB3EE055FB71" italics="true" pageId="1" pageNumber="544">Astrohelix muricatopatella</emphasis>
</taxonomicName>
). Hence,
<taxonomicName id="4C274D354E78D36AFE52FB19E3F7FB72" authority="(Astromoana)" baseAuthorityName="Astromoana" box="[442,744,1254,1278]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFE52FB19E35AFB71" box="[442,581,1254,1277]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFDBFFB19E3C0FB71" box="[599,735,1254,1277]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
</taxonomicName>
(or
<taxonomicName id="4C274D354E78D36AFCF3FB1AE292FB71" box="[795,909,1253,1277]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFCF3FB1AE292FB71" box="[795,909,1253,1277]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
) is not monophyletic. However, I here refrain from synonymising
<taxonomicName id="4C274D354E78D36AFE90FAF6E0F4FAAD" box="[376,491,1289,1313]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFE90FAF6E0F4FAAD" box="[376,491,1289,1313]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
with
<emphasis id="B953EAA44E78D36AFDC1FAF5E3AEFAAD" box="[553,689,1290,1313]" italics="true" pageId="1" pageNumber="544">Asteropotpa</emphasis>
at the genus level because I did not take into account all species of
<taxonomicName id="4C274D354E78D36AFF7FFAD1E03DFAC9" box="[151,290,1326,1349]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFF7FFAD1E03DFAC9" box="[151,290,1326,1349]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
</taxonomicName>
in this study. Therefore, I here simply combine
<taxonomicName id="4C274D354E78D36AFC88FAD2E2CCFAC9" box="[864,979,1325,1349]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFC88FAD2E2CCFAC9" box="[864,979,1325,1349]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
as
<taxonomicName id="4C274D354E78D36AFC17FAD1E1CCFAE6" authority="Doderlein, 1930" authorityName="Doderlein" authorityYear="1930" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subGenus" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFC17FAD1E595FAC9" box="[1023,1162,1326,1349]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFB76FAD2E40FFAC9" box="[1182,1296,1325,1349]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<bibRefCitation id="EFB64B474E78D36AFACDFAD2E1CCFAE6" author="Doderlein" pageId="1" pageNumber="544" refString="Doderlein, L. (1930) Die Ophiuroiden der deutschen Tiefsee-Expedition. 2. Euryalae. Deutsche Tiefsee-Expedition 1898 - 1899, 22, 347 - 396." type="journal article" year="1930">Döderlein, 1930</bibRefCitation>
</taxonomicName>
. Molecular analysis with all species of
<taxonomicName id="4C274D354E78D36AFD7AFAADE202FAE5" box="[658,797,1362,1385]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFD7AFAADE202FAE5" box="[658,797,1362,1385]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C274D354E78D36AFCBDFAAEE2D8FAE5" box="[853,967,1361,1385]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFCBDFAAEE2D8FAE5" box="[853,967,1361,1385]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
are required for the taxonomic review of these (sub) genera.
</paragraph>
</subSubSection>
<subSubSection id="C33D653D4E78D36AFF2FFA65E229F8EE" pageId="1" pageNumber="544" type="reference_group">
<paragraph id="8B9836B64E78D36AFF2FFA65E215F9EA" blockId="1.[151,1437,605,1890]" pageId="1" pageNumber="544">
<taxonomicName id="4C274D354E78D36AFF2FFA65E35FFA3D" ID-CoL="02b59fb1-b2e3-4523-bb4b-6dbd99acce9c" box="[199,576,1433,1458]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="bellator" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFF2FFA65E04DFA3D" box="[199,338,1434,1457]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFE8BFA66E0CAFA3D" box="[355,469,1433,1457]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFE0FFA66E35FFA3D" box="[487,576,1433,1457]" italics="true" pageId="1" pageNumber="544">bellator</emphasis>
</taxonomicName>
comprises two subspecies,
<taxonomicName id="4C274D354E78D36AFC92FA65E483FA3E" authority="Koehler, 1904" authorityName="Koehler" authorityYear="1904" box="[890,1436,1433,1458]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subSpecies" species="bellator" subGenus="Astrohelix" subSpecies="bellator">
<emphasis id="B953EAA44E78D36AFC92FA65E28CFA3D" box="[890,915,1434,1457]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFC4CFA66E509FA3D" box="[932,1046,1433,1457]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFBCFFA66E5FEFA3D" box="[1063,1249,1433,1457]" italics="true" pageId="1" pageNumber="544">bellator bellator</emphasis>
(
<bibRefCitation id="EFB64B474E78D36AFB19FA66E48BFA3E" author="Koehler" box="[1265,1428,1433,1458]" pageId="1" pageNumber="544" refString="Koehler, R. (1904) Ophiures de l'Expedition du Siboga. Part I. Ophiures de Mer profonde. Siboga-Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C274D354E78D36AFF2FFA41E215FA5A" authority="Mortensen, 1933" authorityName="Mortensen" authorityYear="1933" box="[199,778,1469,1494]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="subSpecies" species="bellator" subGenus="Astrohelix" subSpecies="paucidens">
<emphasis id="B953EAA44E78D36AFF2FFA41E1C0FA59" box="[199,223,1470,1493]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFF07FA42E07EFA59" box="[239,353,1469,1493]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFE9BFA42E35AFA59" box="[371,581,1469,1493]" italics="true" pageId="1" pageNumber="544">bellator paucidens</emphasis>
<bibRefCitation id="EFB64B474E78D36AFDA4FA41E215FA5A" author="Mortensen" box="[588,778,1470,1494]" pageId="1" pageNumber="544" refString="Mortensen, T. (1933) Studies of Indo-Pacific euryalids. Fidenskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn, 96, 1 - 75." type="journal article" year="1933">Mortensen, 1933</bibRefCitation>
</taxonomicName>
. Okanishi &amp; Fujita (2011) provided a diagnostic character key of
<taxonomicName id="4C274D354E78D36AFF1CFA1DE338FA76" ID-CoL="fd11a361-e370-4dab-8a0d-a25757a49051" authority="(Astromoana)" baseAuthorityName="Astromoana" box="[244,551,1506,1530]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFF1CFA1DE060FA75" box="[244,383,1506,1529]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFE7EFA1DE301FA75" box="[406,542,1506,1529]" italics="true" pageId="1" pageNumber="544">Astromoana</emphasis>
)
</taxonomicName>
(=
<taxonomicName id="4C274D354E78D36AFDA7FA1DE3C5FA75" ID-CoL="fd11a361-e370-4dab-8a0d-a25757a49051" box="[591,730,1506,1529]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFDA7FA1DE3C5FA75" box="[591,730,1506,1529]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
</taxonomicName>
(
<taxonomicName id="4C274D354E78D36AFD1AFA1EE27BFA75" ID-CoL="5c82e761-adda-4ccb-9d5c-a9a133b784ed" box="[754,868,1505,1529]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="genus">
<emphasis id="B953EAA44E78D36AFD1AFA1EE27BFA75" box="[754,868,1505,1529]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
</taxonomicName>
)) and the holotypes of two subspecies can be distinguished from each other by the combination of these characters (see Discussion below). Therefore, I here propose that the two subspecies should be elevated to species-level as
<taxonomicName id="4C274D354E78D36AFC5BF9D5E1C7F9EA" ID-CoL="02b59fb1-b2e3-4523-bb4b-6dbd99acce9c" authority="Koehler, 1904" authorityName="Koehler" authorityYear="1904" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="bellator" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFC5BF9D5E521F9CD" box="[947,1086,1578,1601]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFBA7F9D6E5DEF9CD" box="[1103,1217,1577,1601]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFB3AF9D6E434F9CD" box="[1234,1323,1577,1601]" italics="true" pageId="1" pageNumber="544">bellator</emphasis>
(
<bibRefCitation id="EFB64B474E78D36AFAD2F9D6E1CFF9EA" author="Koehler" pageId="1" pageNumber="544" refString="Koehler, R. (1904) Ophiures de l'Expedition du Siboga. Part I. Ophiures de Mer profonde. Siboga-Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C274D354E78D36AFEE6F9B1E21AF9EA" ID-CoL="01e4bbf4-6361-48f9-a2b1-c43189fd9394" authority="Mortensen, 1933" authorityName="Mortensen" authorityYear="1933" box="[270,773,1613,1638]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="paucidens" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFEE6F9B1E038F9E9" box="[270,295,1614,1637]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFEDFF9B2E0B6F9E9" box="[311,425,1613,1637]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFE51F9B2E334F9E9" box="[441,555,1613,1637]" italics="true" pageId="1" pageNumber="544">paucidens</emphasis>
(
<bibRefCitation id="EFB64B474E78D36AFDD2F9B1E3E2F9EA" author="Mortensen" box="[570,765,1614,1638]" pageId="1" pageNumber="544" refString="Mortensen, T. (1933) Studies of Indo-Pacific euryalids. Fidenskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn, 96, 1 - 75." type="journal article" year="1933">Mortensen, 1933</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B9836B64E78D36AFF2FF98DE229F8EE" blockId="1.[151,1437,605,1890]" pageId="1" pageNumber="544">
<materialsCitation id="3B4F3CEB4E78D36AFF2FF98DE5F5F95D" ID-GBIF-Occurrence="1427602968" country="Fiji" county="northwestern Australia" elevation="275" location="In" municipality="western Japan" pageId="1" pageNumber="544" specimenCount="1">
<location id="8EF8606D4E78D36AFF2FF98DE1C1F905" LSID="urn:lsid:plazi:treatment:038E87A04E78D36AFF7FFE4AE229F8EE:8EF8606D4E78D36AFF2FF98DE1C1F905" box="[199,222,1650,1673]" country="Fiji" county="northwestern Australia" municipality="western Japan" name="In" pageId="1" pageNumber="544">In</location>
addition to
<taxonomicName id="4C274D354E78D36AFE80F98DE283F906" ID-CoL="02b59fb1-b2e3-4523-bb4b-6dbd99acce9c" authority="Koehler, 1904" authorityName="Koehler" authorityYear="1904" box="[360,924,1649,1674]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="bellator" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFE80F98DE0ECF905" box="[360,499,1650,1673]" italics="true" pageId="1" pageNumber="544">Asteroporpa</emphasis>
(
<emphasis id="B953EAA44E78D36AFDECF98EE369F905" box="[516,630,1649,1673]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFD60F98EE3FEF905" box="[648,737,1649,1673]" italics="true" pageId="1" pageNumber="544">bellator</emphasis>
(
<bibRefCitation id="EFB64B474E78D36AFD19F98EE28BF906" author="Koehler" box="[753,916,1649,1674]" pageId="1" pageNumber="544" refString="Koehler, R. (1904) Ophiures de l'Expedition du Siboga. Part I. Ophiures de Mer profonde. Siboga-Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
)
</taxonomicName>
from the
<location id="8EF8606D4E78D36AFBE5F98EE5E8F906" LSID="urn:lsid:plazi:treatment:038E87A04E78D36AFF7FFE4AE229F8EE:8EF8606D4E78D36AFBE5F98EE5E8F906" box="[1037,1271,1649,1674]" country="Fiji" county="northwestern Australia" municipality="western Japan" name="southern Philippines" pageId="1" pageNumber="544">southern Philippines</location>
,
<quantity id="4CDF9B534E78D36AFAEBF98EE454F905" box="[1283,1355,1649,1674]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="544" unit="m" value="275.0">
<elevation id="000AD1854E78D36AFAEBF98EE454F905" box="[1283,1355,1649,1674]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="544" unit="m" value="275.0">275 m</elevation>
</quantity>
and
<taxonomicName id="4C274D354E78D36AFA6CF98DE364F922" ID-CoL="01e4bbf4-6361-48f9-a2b1-c43189fd9394" authority="Mortensen, 1933" authorityName="Mortensen" authorityYear="1933" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="paucidens" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFA6CF98DE483F905" box="[1412,1436,1650,1673]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFF48F96AE00DF921" box="[160,274,1685,1709]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFEC0F96AE086F921" box="[296,409,1685,1709]" italics="true" pageId="1" pageNumber="544">paucidens</emphasis>
(
<bibRefCitation id="EFB64B474E78D36AFE45F969E36CF922" author="Mortensen" box="[429,627,1686,1710]" pageId="1" pageNumber="544" refString="Mortensen, T. (1933) Studies of Indo-Pacific euryalids. Fidenskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn, 96, 1 - 75." type="journal article" year="1933">Mortensen, 1933</bibRefCitation>
)
</taxonomicName>
from
<collectingMunicipality id="6BFCACCC4E78D36AFD23F969E26EF921" box="[715,881,1686,1709]" pageId="1" pageNumber="544">western Japan</collectingMunicipality>
,
<elevation id="000AD1854E78D36AFC69F969E2D2F921" box="[897,973,1686,1710]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="544" unit="m" value="190.0">
<quantity id="4CDF9B534E78D36AFC69F969E2D2F921" box="[897,973,1686,1710]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="544" unit="m" value="190.0">190 m</quantity>
</elevation>
, this subgenus currently comprises the following species:
<taxonomicName id="4C274D354E78D36AFE84F945E3E3F95E" ID-CoL="7715938a-5435-48da-b8ba-f18f94227a56" authority="Baker, 1980" authorityName="Baker" authorityYear="1980" box="[364,764,1721,1746]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="indicus" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFE84F945E09AF95D" box="[364,389,1722,1745]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFE7EF946E317F95D" box="[406,520,1721,1745]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFDF2F946E374F95D" box="[538,619,1721,1745]" italics="true" pageId="1" pageNumber="544">indicus</emphasis>
<bibRefCitation id="EFB64B474E78D36AFD9BF946E3E3F95E" author="Baker" box="[627,764,1721,1746]" pageId="1" pageNumber="544" refString="Baker, A. N. (1980) Euryalinid Ophiuroidea (Echinodermata) from Australia, New Zealand, and the south-west Pacific Ocean. New Zealand Journal of Zoology, 7, 11 - 83." type="journal article" year="1980">Baker, 1980</bibRefCitation>
</taxonomicName>
from
<collectingCounty id="62F94E3A4E78D36AFCAAF946E55BF95E" box="[834,1092,1721,1746]" pageId="1" pageNumber="544">northwestern Australia</collectingCounty>
to
<collectingCountry id="F33076264E78D36AFB81F946E58CF95E" box="[1129,1171,1721,1746]" name="Fiji" pageId="1" pageNumber="544">Fiji</collectingCountry>
,
<elevation id="000AD1854E78D36AFB77F945E5F5F95D" box="[1183,1258,1722,1746]" metricMagnitude="2" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="544" unit="m" value="141.0">
<quantity id="4CDF9B534E78D36AFB77F945E5F5F95D" box="[1183,1258,1722,1746]" metricMagnitude="2" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="544" unit="m" value="141.0">141 m</quantity>
</elevation>
</materialsCitation>
;
<materialsCitation id="3B4F3CEB4E78D36AFB1FF945E5F9F979" ID-GBIF-Occurrence="1427602972" country="New Zealand" elevation="186" location="southern New Caleconia" pageId="1" pageNumber="544" specimenCount="1">
<taxonomicName id="4C274D354E78D36AFB1FF945E087F97A" ID-CoL="b4be0d90-3559-49ed-b893-74c74a5052f5" authority="Baker, 1980" authorityName="Baker" authorityYear="1980" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="reticulata" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFB1FF945E410F95D" box="[1271,1295,1722,1745]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFAC8F946E48DF95D" box="[1312,1426,1721,1745]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFF7FF922E01CF979" box="[151,259,1757,1781]" italics="true" pageId="1" pageNumber="544">reticulata</emphasis>
<bibRefCitation id="EFB64B474E78D36AFEE5F922E087F97A" author="Baker" box="[269,408,1757,1782]" pageId="1" pageNumber="544" refString="Baker, A. N. (1980) Euryalinid Ophiuroidea (Echinodermata) from Australia, New Zealand, and the south-west Pacific Ocean. New Zealand Journal of Zoology, 7, 11 - 83." type="journal article" year="1980">Baker, 1980</bibRefCitation>
</taxonomicName>
from northwestern
<collectingCountry id="F33076264E78D36AFD68F921E206F97A" box="[640,793,1757,1782]" name="New Zealand" pageId="1" pageNumber="544">New Zealand</collectingCountry>
to
<location id="8EF8606D4E78D36AFCABF922E57FF97A" LSID="urn:lsid:plazi:treatment:038E87A04E78D36AFF7FFE4AE229F8EE:8EF8606D4E78D36AFCABF922E57FF97A" box="[835,1120,1757,1782]" country="New Zealand" name="southern New Caleconia" pageId="1" pageNumber="544">southern New Caleconia</location>
,
<quantity id="4CDF9B534E78D36AFB87F921E5F9F979" box="[1135,1254,1758,1782]" metricMagnitude="2" metricUnit="m" metricValue="1.86" metricValueMax="3.01" metricValueMin="0.71" pageId="1" pageNumber="544" unit="m" value="186.0" valueMax="301.0" valueMin="71.0">
<elevation id="000AD1854E78D36AFB87F921E5F9F979" box="[1135,1254,1758,1782]" metricMagnitude="2" metricUnit="m" metricValue="1.86" metricValueMax="3.01" metricValueMin="0.71" pageId="1" pageNumber="544" unit="m" value="186.0" valueMax="301.0" valueMin="71.0">71–301 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="3B4F3CEB4E78D36AFB1DF921E553F895" ID-GBIF-Occurrence="1427602970" country="Japan" elevation="275" location="Japan" pageId="1" pageNumber="544" specimenCount="1">
<taxonomicName id="4C274D354E78D36AFB1DF921E364F896" ID-CoL="40e58f42-a745-49da-98a0-19ef183b4fd3" authority="Okanishi &amp; Fujita, 2011" authorityName="Okanishi &amp; Fujita" authorityYear="2011" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="muricatopatella" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFB1DF921E412F979" box="[1269,1293,1758,1781]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFAC8F922E48DF979" box="[1312,1426,1757,1781]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFF7FF8FEE055F895" box="[151,330,1793,1817]" italics="true" pageId="1" pageNumber="544">muricatopatella</emphasis>
Okanishi &amp; Fujita, 2011
</taxonomicName>
from southwestern
<collectingCountry id="F33076264E78D36AFC86F8FDE2AEF895" box="[878,945,1794,1817]" name="Japan" pageId="1" pageNumber="544">Japan</collectingCountry>
,
<elevation id="000AD1854E78D36AFC2BF8FDE553F895" box="[963,1100,1794,1818]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="4.1" metricValueMin="1.4" pageId="1" pageNumber="544" unit="m" value="275.0" valueMax="410.0" valueMin="140.0">
<quantity id="4CDF9B534E78D36AFC2BF8FDE553F895" box="[963,1100,1794,1818]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="4.1" metricValueMin="1.4" pageId="1" pageNumber="544" unit="m" value="275.0" valueMax="410.0" valueMin="140.0">140–410 m</quantity>
</elevation>
</materialsCitation>
;
<materialsCitation id="3B4F3CEB4E78D36AFBB6F8FEE229F8EE" ID-GBIF-Occurrence="1427602973" collectorName="Baker &amp; Okanishi &amp; Fujita &amp; T. D. O'Hara &amp; Table" country="Japan" elevation="743" location="Japan" pageId="1" pageNumber="544" specimenCount="1">
and
<taxonomicName id="4C274D354E78D36AFB7CF8FDE0B8F8B2" ID-CoL="c430010f-e124-4690-b354-049fb536402a" authority="Okanishi &amp; Fujita, 2011" authorityName="Okanishi &amp; Fujita" authorityYear="2011" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrohelix" kingdom="Animalia" order="Euryalida" pageId="1" pageNumber="544" phylum="Echinodermata" rank="species" species="koyoae" subGenus="Astrohelix">
<emphasis id="B953EAA44E78D36AFB7CF8FDE5B2F895" box="[1172,1197,1794,1817]" italics="true" pageId="1" pageNumber="544">A.</emphasis>
(
<emphasis id="B953EAA44E78D36AFB2CF8FEE429F895" box="[1220,1334,1793,1817]" italics="true" pageId="1" pageNumber="544">Astrohelix</emphasis>
)
<emphasis id="B953EAA44E78D36AFAA5F8FEE483F895" box="[1357,1436,1793,1817]" italics="true" pageId="1" pageNumber="544">koyoae</emphasis>
Okanishi &amp; Fujita, 2011
</taxonomicName>
from southern
<collectingCountry id="F33076264E78D36AFDBCF8D9E388F8B1" box="[596,663,1830,1853]" name="Japan" pageId="1" pageNumber="544">Japan</collectingCountry>
,
<elevation id="000AD1854E78D36AFD4AF8D9E3E1F8B1" box="[674,766,1830,1854]" metricMagnitude="2" metricUnit="m" metricValue="7.431" pageId="1" pageNumber="544" unit="m" value="743.1">
<quantity id="4CDF9B534E78D36AFD4AF8D9E3E1F8B1" box="[674,766,1830,1854]" metricMagnitude="2" metricUnit="m" metricValue="7.431" pageId="1" pageNumber="544" unit="m" value="743.1">743.1 m</quantity>
</elevation>
(
<bibRefCitation id="EFB64B474E78D36AFCE5F8DAE286F8B2" author="Baker" box="[781,921,1829,1854]" pageId="1" pageNumber="544" refString="Baker, A. N. (1980) Euryalinid Ophiuroidea (Echinodermata) from Australia, New Zealand, and the south-west Pacific Ocean. New Zealand Journal of Zoology, 7, 11 - 83." type="journal article" year="1980">
<collectorName id="26D253604E78D36AFCE5F8DAE24CF8B2" box="[781,851,1829,1854]" pageId="1" pageNumber="544">Baker</collectorName>
, 1980
</bibRefCitation>
;
<collectorName id="26D253604E78D36AFC4DF8DAE515F8B2" box="[933,1034,1829,1854]" pageId="1" pageNumber="544">Okanishi</collectorName>
&amp;
<collectorName id="26D253604E78D36AFBC7F8DAE56CF8B2" box="[1071,1139,1829,1854]" pageId="1" pageNumber="544">Fujita</collectorName>
, 2011;
<collectorName id="26D253604E78D36AFB2CF8D9E444F8B2" box="[1220,1371,1830,1854]" pageId="1" pageNumber="544">T. D. O’Hara</collectorName>
, pers. comm.). A tabular key to the six species is given in
<tableCitation id="C6A5030D4E78D36AFD34F8B6E22FF8EE" box="[732,816,1865,1890]" captionStart="TABLE 1" captionStartId="5.[125,202,715,736]" captionTargetBox="[138,1897,888,1485]" captionTargetPageId="5" captionText="TABLE 1. A tabular characters key to the species of Asteroporpa (Astrohelix)." httpUri="http://table.plazi.org/id/DF58663E4E7CD36EFF95FD34E242FD6C" pageId="1" pageNumber="544" tableUuid="DF58663E4E7CD36EFF95FD34E242FD6C">
<collectorName id="26D253604E78D36AFD34F8B6E205F8EE" box="[732,794,1865,1890]" pageId="1" pageNumber="544">Table</collectorName>
1
</tableCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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