267 lines
32 KiB
XML
267 lines
32 KiB
XML
<document id="052CC5672125A7D6859056DBCEFFBAF4" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFA13C42FC06FBCAFE90699A" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada epiphallica Köhler 2011, n.sp." docType="treatment" docVersion="3" lastPageNumber="195" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="193" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="6FC13E84B32ABCF484E49218AA3CC430" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B97E6FABE389AFA77D96D4072C87B040">
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<mods:title id="7FF779F1FED3D17D63794C90580B4703">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="D07D27300C258EBF3E842DC5A09D01C3" type="personal">
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<mods:role id="029D69E14208F043918E9A2B53ABDB94">
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<mods:roleTerm id="5FCE3C3BCAF84C3678DEE511AC3365B4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A6D78C3D39DFF011939E15DD4763823">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="8D3B8CD9460B5589A354656DD459EADB">text</mods:typeOfResource>
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<mods:title id="E15C5781F1E6C9B1198A271616B4DFA1">Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part id="EF47E161EAAB64CE6727B5D8DC1475D6">
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<mods:date id="F24F74C132E2BFBECA513714BBC0A121">2011</mods:date>
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<mods:number id="540AE6A52C67B46BC39350E5CE7E506A">2011-11-30</mods:number>
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</mods:detail>
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<mods:detail id="7042C14C26AEAFFB2B5F808FA593A944" type="volume">
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<mods:number id="978602DA3AA83D773938A96AF919E9C0">63</mods:number>
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</mods:detail>
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<mods:detail id="506ED85F0905FB33CDB8ED08869ECB0D" type="issue">
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<mods:number id="FA2CB9E99BB050F2F2E13F4600A8644E">2</mods:number>
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<mods:extent id="FE17C7673237C4E14CC1E77964BFC961" unit="page">
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<mods:start id="AE4916E7889699E68913299A4E939EC3">167</mods:start>
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<mods:end id="780B742AE5414C14D76555AB48C972AE">202</mods:end>
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<mods:url id="B6836289A0720C478DA1FCC402DCCFBF">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
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</mods:location>
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<mods:classification id="8D0DD76606747517E1086C7AFFF07916">journal article</mods:classification>
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<mods:identifier id="C6B6A5AC238A610A60EA9F2B3DE9060B" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
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<mods:identifier id="80812208C31973EF43D9F91D91D2C4D1" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="2F9FB4538436BB7E1D0B8BC3B2C2A401" type="Zenodo-Dep">5239547</mods:identifier>
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<mods:identifier id="0DD10DB85B22AAA65844E928B05C2309" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
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</mods:mods>
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<treatment id="0384503AFFA13C42FC06FBCAFE90699A" LSID="urn:lsid:plazi:treatment:0384503AFFA13C42FC06FBCAFE90699A" httpUri="http://treatment.plazi.org/id/0384503AFFA13C42FC06FBCAFE90699A" lastPageId="28" lastPageNumber="195" pageId="26" pageNumber="193">
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<subSubSection id="C337B2A7FFA13C44FC06FBCAFAA96A78" box="[958,1348,1048,1074]" pageId="26" pageNumber="193" type="nomenclature">
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<paragraph id="8B92E12CFFA13C44FC06FBCAFAA96A78" blockId="26.[958,1348,1048,1074]" box="[958,1348,1048,1074]" pageId="26" pageNumber="193">
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<heading id="D0DA5640FFA13C44FC06FBCAFAA96A78" box="[958,1348,1048,1074]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="193" reason="2">
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<emphasis id="B9593D3EFFA13C44FC06FBCAFAA96A78" bold="true" box="[958,1348,1048,1074]" pageId="26" pageNumber="193">
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<taxonomicName id="4C2D9AAFFFA13C44FC06FBCAFAED6A78" ID-CoL="D7QV" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[958,1280,1048,1074]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="193" phylum="Mollusca" rank="species" species="epiphallica" status="sp. nov.">
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<emphasis id="B9593D3EFFA13C44FC06FBCAFAED6A78" bold="true" box="[958,1280,1048,1074]" italics="true" pageId="26" pageNumber="193">Amplirhagada epiphallica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26A8045FFA13C44FAB0FBCAFAA96A78" box="[1288,1348,1048,1074]" pageId="26" pageNumber="193" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C337B2A7FFA13C44FCFFFB97FA7D6B7D" pageId="26" pageNumber="193" type="materials_examined">
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<paragraph id="8B92E12CFFA13C44FCFFFB97FACC6AFE" blockId="26.[839,1467,1093,1204]" pageId="26" pageNumber="193">
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<materialsCitation id="3B45EB71FFA13C44FCFFFB97FAF16AFE" collectingDate="1987-06-14" collectionCode="V" collectorName="NW Kimberley, N" latitude="-14.952778" location="Prince Frederick Harbour" longLatPrecision="21" longitude="125.275" pageId="26" pageNumber="193" specimenCode="RFS 14-1" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B9593D3EFFA13C44FCFFFB97FC316A17" bold="true" box="[839,988,1093,1117]" pageId="26" pageNumber="193">
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<typeStatus id="54965F8EFFA13C44FCFFFB97FC6D6A17" box="[839,896,1093,1117]" pageId="26" pageNumber="193">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingRegion id="49E92FCEFFA13C44FC54FB97FB5D6A16" box="[1004,1200,1093,1116]" country="Australia" name="Western Australia" pageId="26" pageNumber="193">Western Australia</collectingRegion>
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,
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<collectorName id="26D884FAFFA13C44FB05FB97FA6B6A16" box="[1213,1414,1093,1116]" pageId="26" pageNumber="193">NW Kimberley, N</collectorName>
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side of
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<location id="8EF2B7F7FFA13C44FCDDFBB0FB946A33" LSID="urn:lsid:plazi:treatment:0384503AFFA13C42FC06FBCAFE90699A:8EF2B7F7FFA13C44FCDDFBB0FB946A33" box="[869,1145,1122,1145]" latitude="-14.952778" longLatPrecision="21" longitude="125.275" name="Prince Frederick Harbour" pageId="26" pageNumber="193" stateProvince="Western Australia">Prince Frederick Harbour</location>
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, near mouth,
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<locationDeviation id="9A20F80EFFA13C44FAB4FBB0FC5F6ADD" location="Mt. Anderdon" pageId="26" pageNumber="193">
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<quantity id="4CD54CC9FFA13C44FAB4FBB0FABD6A33" box="[1292,1360,1122,1145]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="193" unit="km" value="16.0">16 km</quantity>
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W of Mt. Anderdon
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</locationDeviation>
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,
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<quantity id="4CD54CC9FFA13C44FC02FB52FBEC6ADD" box="[954,1025,1152,1175]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="193" unit="km" value="0.3">0.3 km</quantity>
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from coast;
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<geoCoordinate id="EE1987EBFFA13C44FBC6FB52FB196ADD" box="[1150,1268,1152,1175]" degrees="14" direction="south" minutes="57" orientation="latitude" pageId="26" pageNumber="193" precision="15" seconds="10" value="-14.952778">14°57'10"S</geoCoordinate>
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<geoCoordinate id="EE1987EBFFA13C44FB41FB52FA936ADD" box="[1273,1406,1152,1175]" degrees="125" direction="east" minutes="16" orientation="longitude" pageId="26" pageNumber="193" precision="15" seconds="30" value="125.275">125°16'30"E</geoCoordinate>
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||
(
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<specimenCode id="DB8B4957FFA13C44FA32FB52FC976AFE" collectionCode="RFS" pageId="26" pageNumber="193">RFS 14-1</specimenCode>
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||
; coll.
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<collectionCode id="ED3C79E9FFA13C44FC02FB4FFC206AFE" box="[954,973,1181,1204]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="193" type="Museum">V</collectionCode>
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||
. Kessner,
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<date id="FF93C7ECFFA13C44FB81FB4FFB506AFE" box="[1081,1213,1181,1204]" pageId="26" pageNumber="193" value="1987-06-14">
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<collectingDate id="EFD73E04FFA13C44FB81FB4FFB506AFE" box="[1081,1213,1181,1204]" pageId="26" pageNumber="193" value="1987-06-14">14 Jun 1987</collectingDate>
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</date>
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) (
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<figureCitation id="1316FDA9FFA13C44FB6AFB4FFAFC6AFE" box="[1234,1297,1181,1204]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="26" pageNumber="193">Fig. 1</figureCitation>
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)
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</materialsCitation>
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.
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||
</paragraph>
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<paragraph id="8B92E12CFFA13C44FCFFFB15FA7D6B7D" blockId="26.[839,1467,1223,1335]" pageId="26" pageNumber="193">
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<emphasis id="B9593D3EFFA13C44FCFFFB15FC336A95" bold="true" box="[839,990,1223,1247]" pageId="26" pageNumber="193">Type material</emphasis>
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||
.
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<materialsCitation id="3B45EB71FFA13C44FC51FB1AFBFC6AB6" accessionNumber="S34745" collectionCode="WAM" pageId="26" pageNumber="193" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54965F8EFFA13C44FC51FB1AFBA56A95" box="[1001,1096,1224,1247]" pageId="26" pageNumber="193" type="holotype">Holotype</typeStatus>
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||
<collectionCode id="ED3C79E9FFA13C44FBF3FB1AFB676A95" box="[1099,1162,1224,1247]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="26" pageNumber="193" type="Museum">WAM</collectionCode>
|
||
<accessionNumber id="947E7CCFFFA13C44FB37FB1AFB336A95" box="[1167,1246,1224,1247]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34745" pageId="26" pageNumber="193" type="EnaNcbi">S34745</accessionNumber>
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(preserved specimen) (Pl. 1.15;
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<tableCitation id="C6AFD497FFA13C44FC0DFB37FBE86AB6" box="[949,1029,1253,1276]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximum–minimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="26" pageNumber="193">Table 1</tableCitation>
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||
)
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</materialsCitation>
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||
.
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<materialsCitation id="3B45EB71FFA13C44FB98FB37FC526B53" accessionNumber="S34746" collectionCode="WAM" pageId="26" pageNumber="193" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54965F8EFFA13C44FB98FB37FB6B6AB6" box="[1056,1158,1253,1276]" pageId="26" pageNumber="193" type="paratype">Paratypes</typeStatus>
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||
<collectionCode id="ED3C79E9FFA13C44FB37FB37FB3D6AB6" box="[1167,1232,1253,1276]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="26" pageNumber="193" type="Museum">WAM</collectionCode>
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<accessionNumber id="947E7CCFFFA13C44FB62FB37FAC76AB6" box="[1242,1322,1253,1276]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34746" pageId="26" pageNumber="193" type="EnaNcbi">S34746</accessionNumber>
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(3 preserved specimens)
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</materialsCitation>
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,
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<materialsCitation id="3B45EB71FFA13C44FC71FAD0FA9D6B53" box="[969,1392,1282,1305]" collectionCode="FMNH" pageId="26" pageNumber="193" specimenCode="FMNH 200539" specimenCount="1" typeStatus="paratype">
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||
<specimenCode id="DB8B4957FFA13C44FC71FAD0FB876B53" box="[969,1130,1282,1305]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="26" pageNumber="193" type="Museum">FMNH 200539</specimenCode>
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||
(4 preserved specimens)
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B45EB71FFA13C44FAC2FAD0FBD16B7D" accessionNumber="S34747" collectionCode="WAM" pageId="26" pageNumber="193" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED3C79E9FFA13C44FAC2FAD0FA576B53" box="[1402,1466,1282,1305]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="26" pageNumber="193" type="Museum">WAM</collectionCode>
|
||
<accessionNumber id="947E7CCFFFA13C44FCFFFAF2FC7A6B7D" box="[839,919,1312,1335]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34747" pageId="26" pageNumber="193" type="EnaNcbi">S34747</accessionNumber>
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||
(2 dried shells)
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B45EB71FFA13C44FBFEFAF2FA616B7D" box="[1094,1420,1312,1335]" collectionCode="FMNH" pageId="26" pageNumber="193" specimenCode="FMNH 220541" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB8B4957FFA13C44FBFEFAF2FB0B6B7D" box="[1094,1254,1312,1335]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="26" pageNumber="193" type="Museum">FMNH 220541</specimenCode>
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||
(2 dried shells)
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337B2A7FFA13C44FCFFFA98FBE86BD6" pageId="26" pageNumber="193" type="etymology">
|
||
<paragraph id="8B92E12CFFA13C44FCFFFA98FBE86BD6" blockId="26.[839,1467,1354,1474]" pageId="26" pageNumber="193">
|
||
<emphasis id="B9593D3EFFA13C44FCFFFA98FC2F6B28" bold="true" box="[839,962,1354,1378]" pageId="26" pageNumber="193">Etymology</emphasis>
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||
. The species epithet refers to the presence of an epiphallus, which is a very distinctive feature among species of
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||
<taxonomicName id="4C2D9AAFFFA13C44FCDCFA57FC136BD6" box="[868,1022,1413,1436]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="193" phylum="Mollusca" rank="genus">
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||
<emphasis id="B9593D3EFFA13C44FCDCFA57FC136BD6" box="[868,1022,1413,1436]" italics="true" pageId="26" pageNumber="193">Amplirhagada</emphasis>
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||
</taxonomicName>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337B2A7FFA13C45FB8FFA7AFB7E69F9" lastPageId="27" lastPageNumber="194" pageId="26" pageNumber="193" type="description">
|
||
<paragraph id="8B92E12CFFA13C44FB8FFA7AFB266B88" blockId="26.[839,1467,1354,1474]" box="[1079,1227,1448,1474]" pageId="26" pageNumber="193">
|
||
<emphasis id="B9593D3EFFA13C44FB8FFA7AFB266B88" bold="true" box="[1079,1227,1448,1474]" pageId="26" pageNumber="193">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFA13C44FCFFFA07FA6A69CD" blockId="26.[839,1467,1493,2015]" pageId="26" pageNumber="193">
|
||
<emphasis id="B9593D3EFFA13C44FCFFFA07FC906BA6" box="[839,893,1493,1516]" italics="true" pageId="26" pageNumber="193">Shell</emphasis>
|
||
(Pl. 1.15;
|
||
<figureCitation id="1316FDA9FFA13C44FC41FA07FB646BA6" box="[1017,1161,1493,1516]" captionStart="Figure 43" captionStartId="29.[121,183,1522,1543]" captionTargetBox="[121,1421,151,1510]" captionTargetId="figure-9@29.[121,1421,151,1510]" captionTargetPageId="29" captionText="Figure 43.Amplirhagada epiphallica n.sp.SEM of shell, radula and jaw (WAM S34746).(A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up of sculpture on last whorl, lateral view. (D) Jaw. (E) Central and inner lateral teeth viewed from above. (F) Marginal teeth viewed from above.Scale bars: A–C, 1 mm; D, 100 µm; E–F, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239635" httpUri="https://zenodo.org/record/5239635/files/figure.png" pageId="26" pageNumber="193">Fig. 43 A–C</figureCitation>
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||
). Broadly conical with low spire, almost discoid. Thin to delicate, translucent. Periphery rounded to angulate; upper and basal sectors of whorls well rounded. Umbilicus 10–50% concealed by columellar reflection. Background colour pale yellowish brown; peripheral band well marked, moderately thick, brown, visible on most whorls; sub-sutural well marked to diffuse, moderately thick, visible on most whorls; outer and inner lip colour whitish. Protoconch
|
||
<quantity id="4CD54CC9FFA13C44FBD2F912FB57689D" box="[1130,1210,1728,1751]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="193" unit="mm" value="2.5">2.5 mm</quantity>
|
||
in diameter, comprising about 1.2 whorls, essentially smooth. Teleoconch with regular axial growth lines; evenly distributed across shell surface. Angle of aperture about 45 degrees; outer lip thin, well rounded, expanded, not reflected; basal node absent or weakly developed. Parietal wall of inner lip inconspicuous. Average shell size 13.7±0.8 × 20.1±
|
||
<quantity id="4CD54CC9FFA13C44FB7FF8A2FAF469CD" box="[1223,1305,1904,1927]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="193" unit="mm" value="0.7">0.7 mm</quantity>
|
||
(
|
||
<tableCitation id="C6AFD497FFA13C44FA9FF8A2FA9A69CD" box="[1319,1399,1904,1927]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximum–minimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="26" pageNumber="193">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFA13C44FCDDF85FFB876995" blockId="26.[839,1467,1493,2015]" pageId="26" pageNumber="193">
|
||
<emphasis id="B9593D3EFFA13C44FCDDF85FFB5669EE" box="[869,1211,1933,1956]" italics="true" pageId="26" pageNumber="193">Radular and jaw morphology</emphasis>
|
||
(
|
||
<figureCitation id="1316FDA9FFA13C44FB76F85FFA8969EE" box="[1230,1380,1933,1956]" captionStart="Figure 43" captionStartId="29.[121,183,1522,1543]" captionTargetBox="[121,1421,151,1510]" captionTargetId="figure-9@29.[121,1421,151,1510]" captionTargetPageId="29" captionText="Figure 43.Amplirhagada epiphallica n.sp.SEM of shell, radula and jaw (WAM S34746).(A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Close-up of sculpture on last whorl, lateral view. (D) Jaw. (E) Central and inner lateral teeth viewed from above. (F) Marginal teeth viewed from above.Scale bars: A–C, 1 mm; D, 100 µm; E–F, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239635" httpUri="https://zenodo.org/record/5239635/files/figure.png" pageId="26" pageNumber="193">Fig. 43 D–F</figureCitation>
|
||
). Tooth formula C + 13–14 + 4 + 18–20, with 150 rows of teeth 130 (n = 1). Jaw with 11 plates.
|
||
</paragraph>
|
||
<caption id="DF52B1A4FFA03C45FFC1F9DEFE46683B" ID-DOI="http://doi.org/10.5281/zenodo.5239629" ID-Zenodo-Dep="5239629" httpUri="https://zenodo.org/record/5239629/files/figure.png" pageId="27" pageNumber="194" startId="27.[121,184,1548,1569]" targetBox="[121,1421,151,1530]" targetPageId="27">
|
||
<paragraph id="8B92E12CFFA03C45FFC1F9DEFE46683B" blockId="27.[121,1421,1548,1649]" pageId="27" pageNumber="194">
|
||
Figure 40.
|
||
<taxonomicName id="4C2D9AAFFFA03C45FF59F9DEFE15686B" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[225,504,1548,1569]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="194" phylum="Mollusca" rank="species" species="angustocauda" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFA03C45FF59F9DEFE15686B" box="[225,504,1548,1569]" italics="true" pageId="27" pageNumber="194">Amplirhagada angustocauda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFA03C45FE46F9DEFDC1686B" box="[510,556,1548,1569]" pageId="27" pageNumber="194" rank="species">n.sp.</taxonomicNameLabel>
|
||
SEM of shell, radula and jaw (WAM
|
||
<accessionNumber id="947E7CCFFFA03C45FC2EF9DEFC0E686B" box="[918,995,1548,1569]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34743" pageId="27" pageNumber="194" type="EnaNcbi">S34743</accessionNumber>
|
||
).
|
||
<emphasis id="B9593D3EFFA03C45FC4BF9DEFBFC686B" box="[1011,1041,1548,1569]" italics="true" pageId="27" pageNumber="194">(A)</emphasis>
|
||
Protoconch and first teleoconch whorls viewed from above.
|
||
<emphasis id="B9593D3EFFA03C45FE87F9F5FEB06876" box="[319,349,1575,1596]" italics="true" pageId="27" pageNumber="194">(B)</emphasis>
|
||
Detail of first teleoconch whorls viewed from above.
|
||
<emphasis id="B9593D3EFFA03C45FCD0F9F5FC656876" box="[872,904,1575,1596]" italics="true" pageId="27" pageNumber="194">(C)</emphasis>
|
||
Close-up of sculpture on last whorl, lateral view.
|
||
<emphasis id="B9593D3EFFA03C45FAD4F9F5FA606876" box="[1388,1421,1575,1596]" italics="true" pageId="27" pageNumber="194">(D)</emphasis>
|
||
Jaw.
|
||
<emphasis id="B9593D3EFFA03C45FF11F990FF25681D" box="[169,200,1602,1623]" italics="true" pageId="27" pageNumber="194">(E)</emphasis>
|
||
Central and inner lateral teeth viewed from above.
|
||
<emphasis id="B9593D3EFFA03C45FD04F990FD37681D" box="[700,730,1602,1623]" italics="true" pageId="27" pageNumber="194">(F)</emphasis>
|
||
Marginal teeth viewed from above. Scale bars:
|
||
<emphasis id="B9593D3EFFA03C45FB13F990FB57681D" box="[1195,1210,1602,1623]" italics="true" pageId="27" pageNumber="194">A</emphasis>
|
||
, 200 µm;
|
||
<emphasis id="B9593D3EFFA03C45FAA6F990FAA2681D" box="[1310,1359,1602,1623]" italics="true" pageId="27" pageNumber="194">B–C,</emphasis>
|
||
1 mm
|
||
<emphasis id="B9593D3EFFA03C45FFC1F98EFF67683B" box="[121,138,1628,1649]" italics="true" pageId="27" pageNumber="194">D</emphasis>
|
||
, 100 µm;
|
||
<emphasis id="B9593D3EFFA03C45FF54F98EFF16683B" box="[236,251,1628,1649]" italics="true" pageId="27" pageNumber="194">E</emphasis>
|
||
, 20 µm;
|
||
<emphasis id="B9593D3EFFA03C45FEE8F98EFE8F683B" box="[336,354,1628,1649]" italics="true" pageId="27" pageNumber="194">F,</emphasis>
|
||
10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B92E12CFFA03C45FF2FF946FAD869DC" blockId="27.[121,749,1684,2000]" lastBlockId="27.[794,1421,1684,1971]" pageId="27" pageNumber="194">
|
||
<emphasis id="B9593D3EFFA03C45FF2FF946FE6B68E1" box="[151,390,1684,1707]" italics="true" pageId="27" pageNumber="194">Genital morphology</emphasis>
|
||
(
|
||
<figureCitation id="1316FDA9FFA03C45FE22F946FDC568E1" box="[410,552,1684,1707]" captionStart-0="Figure 44" captionStart-1="Figure 45" captionStartId-0="30.[166,229,562,583]" captionStartId-1="30.[166,230,1583,1604]" captionTargetBox-0="[174,787,163,530]" captionTargetBox-1="[175,786,661,1564]" captionTargetId-0="figure-9@30.[166,793,151,538]" captionTargetId-1="figure-32@30.[166,793,653,1576]" captionTargetPageId-0="30" captionTargetPageId-1="30" captionText-0="Figure 44. Genitalia of Amplirhagada epiphallica n.sp. (WAM S34746). For labelling of structures see Fig. 3. Scale bar = 5 mm." captionText-1="Figure 45. Penial anatomy of Amplirhagada epiphallica n.sp. (WAM S34746). For labelling of structures see Fig. 4. (A) Interior anatomy of the penial wall. (B) Schematic view of the penis with the penial sheath being removed. Scale bars = 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5239637" figureDoi-1="http://doi.org/10.5281/zenodo.5239639" httpUri-0="https://zenodo.org/record/5239637/files/figure.png" httpUri-1="https://zenodo.org/record/5239639/files/figure.png" pageId="27" pageNumber="194">Figs 44–45</figureCitation>
|
||
). Penis complex elongated, tubular, coiled; actual penis extending proximal two thirds of penial complex. Inner penial wall completely lined with regular, fine pustulation arranged in corrugated pilasters; no main stimulatory pilaster differentiated. Penial sheath thick. Penial retractor muscle stubby, no longer than 1/10 of penial complex. Vas deferens moderately thick, entering penial sheath at apical end of penial complex, forming an extended loop, reflecting as an elongate, thin, tubular epiphallus, which gives rise to the short penis. Inner wall of epiphallus smooth, ciliated. Penial verge tiny, slender, forming a loop with pointed tip. Vas deferens elongated, winding before entering the uterus. Vagina long, tubular; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct, with bulbous head. Free oviduct elongated, coiled before entering the uterus, comprising more than half of length of anterior part of oviduct. Spermoviduct no longer than anterior part of oviduct. Albumen gland as long as spermoviduct.
|
||
</paragraph>
|
||
<paragraph id="8B92E12CFFA03C45FC80F84EFB7E69F9" blockId="27.[794,1421,1684,1971]" box="[824,1171,1948,1971]" pageId="27" pageNumber="194">
|
||
<emphasis id="B9593D3EFFA03C45FC80F84EFBE669F9" box="[824,1035,1948,1971]" italics="true" pageId="27" pageNumber="194">Aestivation strategy</emphasis>
|
||
. Free sealer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF52B1A4FFA73C42FF1EFD5AFCF66CF2" ID-DOI="http://doi.org/10.5281/zenodo.5239631" ID-Zenodo-Dep="5239631" httpUri="https://zenodo.org/record/5239631/files/figure.png" pageId="28" pageNumber="195" startId="28.[166,229,648,669]" targetBox="[181,788,155,631]" targetPageId="28">
|
||
<paragraph id="8B92E12CFFA73C42FF1EFD5AFCF66CF2" blockId="28.[166,795,648,696]" pageId="28" pageNumber="195">
|
||
Figure 41. Genitalia of
|
||
<taxonomicName id="4C2D9AAFFFA73C42FE3FFD5AFD736CD7" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[391,670,648,669]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="species" species="angustocauda" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFA73C42FE3FFD5AFD736CD7" box="[391,670,648,669]" italics="true" pageId="28" pageNumber="195">Amplirhagada angustocauda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFA73C42FD1CFD5AFD3F6CD7" box="[676,722,648,669]" pageId="28" pageNumber="195" rank="species">n.sp.</taxonomicNameLabel>
|
||
(WAM
|
||
<accessionNumber id="947E7CCFFFA73C42FF1EFD71FF196CF2" box="[166,244,675,696]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34743" pageId="28" pageNumber="195" type="EnaNcbi">S34743</accessionNumber>
|
||
). For labelling of structures see Fig. 3. Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF52B1A4FFA73C42FF1EFA23FD4F681C" ID-DOI="http://doi.org/10.5281/zenodo.5239633" ID-Zenodo-Dep="5239633" httpUri="https://zenodo.org/record/5239633/files/figure.png" pageId="28" pageNumber="195" startId="28.[166,229,1521,1542]" targetBox="[172,791,759,1481]" targetPageId="28">
|
||
<paragraph id="8B92E12CFFA73C42FF1EFA23FD4F681C" blockId="28.[166,794,1521,1622]" pageId="28" pageNumber="195">
|
||
Figure 42. Penial anatomy of
|
||
<taxonomicName id="4C2D9AAFFFA73C42FE74FA23FD08684C" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[460,741,1521,1542]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="species" species="angustocauda" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFA73C42FE74FA23FD08684C" box="[460,741,1521,1542]" italics="true" pageId="28" pageNumber="195">Amplirhagada angustocauda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFA73C42FD54FA23FCF7684C" box="[748,794,1521,1542]" pageId="28" pageNumber="195" rank="species">n.sp.</taxonomicNameLabel>
|
||
(WAM
|
||
<accessionNumber id="947E7CCFFFA73C42FF56F9D9FED6686A" box="[238,315,1547,1568]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34743" pageId="28" pageNumber="195" type="EnaNcbi">S34743</accessionNumber>
|
||
). For labelling of structures see Fig. 4.
|
||
<emphasis id="B9593D3EFFA73C42FD16F9DEFD21686B" box="[686,716,1548,1569]" italics="true" pageId="28" pageNumber="195">(A)</emphasis>
|
||
Interior anatomy of the penial wall.
|
||
<emphasis id="B9593D3EFFA73C42FE0CF9F4FE3F6871" box="[436,466,1574,1595]" italics="true" pageId="28" pageNumber="195">(B)</emphasis>
|
||
Schematic view of the penis with the penial sheath being removed. Scale bars = 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C337B2A7FFA73C42FF1EF941FE90699A" pageId="28" pageNumber="195" type="discussion">
|
||
<paragraph id="8B92E12CFFA73C42FF1EF941FE90699A" blockId="28.[166,794,1683,2000]" pageId="28" pageNumber="195">
|
||
<emphasis id="B9593D3EFFA73C42FF1EF941FEE268E1" bold="true" box="[166,271,1683,1707]" pageId="28" pageNumber="195">Remarks</emphasis>
|
||
. Anatomical description based on dissection of the
|
||
<typeStatus id="54965F8EFFA73C42FF69F963FED96882" box="[209,308,1713,1736]" pageId="28" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
. Listed by Solem (1991) as “
|
||
<taxonomicName id="4C2D9AAFFFA73C42FDC5F963FF3668AF" authority="NSP" authorityName="NSP" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9593D3EFFA73C42FDC5F963FCF76882" box="[637,794,1713,1736]" italics="true" pageId="28" pageNumber="195">Amplirhagada</emphasis>
|
||
NSP
|
||
</taxonomicName>
|
||
35” and found in sympatry with
|
||
<taxonomicName id="4C2D9AAFFFA73C42FD8EF91CFD5F68AF" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[566,690,1742,1765]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="species" species="inusitata" status="sp. nov.">
|
||
<emphasis id="B9593D3EFFA73C42FD8EF91CFD5F68AF" box="[566,690,1742,1765]" italics="true" pageId="28" pageNumber="195">A. inusitata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26A8045FFA73C42FD01F91CFD0068AF" box="[697,749,1742,1765]" pageId="28" pageNumber="195" rank="species">n.sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C2D9AAFFFA73C42FF1EF939FE4B6948" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" box="[166,422,1771,1794]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="species" species="pusilla">
|
||
<emphasis id="B9593D3EFFA73C42FF1EF939FEE26948" box="[166,271,1771,1794]" italics="true" pageId="28" pageNumber="195">A. pusilla</emphasis>
|
||
<bibRefCitation id="EFBC9CDDFFA73C42FEA0F939FE4B6948" author="Solem, A" box="[280,422,1771,1794]" pageId="28" pageNumber="195" pagination="147 - 320" refId="ref20498" refString="Solem, A. 1981. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Suppl. 11: 147 - 320." type="journal article" year="1981">Solem, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
. The flat shell is similar to that of
|
||
<taxonomicName id="4C2D9AAFFFA73C42FF1EF8DBFEC1696A" authorityName="Köhler" authorityYear="2011" box="[166,300,1801,1824]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="species" species="discoidea">
|
||
<emphasis id="B9593D3EFFA73C42FF1EF8DBFEC1696A" box="[166,300,1801,1824]" italics="true" pageId="28" pageNumber="195">A. discoidea</emphasis>
|
||
</taxonomicName>
|
||
. However, the genital anatomy of this species is very unusual for
|
||
<taxonomicName id="4C2D9AAFFFA73C42FEC1F8F4FDFB6977" box="[377,534,1830,1853]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9593D3EFFA73C42FEC1F8F4FDFB6977" box="[377,534,1830,1853]" italics="true" pageId="28" pageNumber="195">Amplirhagada</emphasis>
|
||
</taxonomicName>
|
||
showing a combination of exclusive features, such as a very long and winding vas deferens, a long tubular vagina, very long tubular penis and vas deferens reflecting as an epiphallus within the strongly developed penial sheath, and a short, thin and coiled penial verge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |