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<document id="74B5ED65D84B352082912D15D556405C" ID-CLB-Dataset="45846" ID-DOI="10.5281/zenodo.279365" ID-GBIF-Dataset="4695ae82-fbd8-4fe7-839d-a8796a7324e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="279365" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460416647325" checkinUser="plazi" docAuthor="Menegon, Michele, Davenport, Tim R. B. &amp; Howell, Kim M." docDate="2011" docId="03B787851A477C5896E696B030E28CAC" docLanguage="en" docName="zt03120p054.pdf" docOrigin="Zootaxa 3120" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atheris matildae Menegon, Davenport &amp; Howell, sp. nov." docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="FF8EFFFD1A467C5F9671921A3355880E" masterDocTitle="Description of a new and critically endangered species of Atheris (Serpentes: Viperidae) from the Southern Highlands of Tanzania, with an overview of the country’s tree viper fauna" masterLastPageNumber="54" masterPageNumber="43" pageNumber="44" updateTime="1698328933845" updateUser="ExternalLinkService">
<mods:mods id="087AB122AD868050C718C541F45F8727" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="17A816535D2B3CF852F05AEF52336295">Description of a new and critically endangered species of Atheris (Serpentes: Viperidae) from the Southern Highlands of Tanzania, with an overview of the country’s tree viper fauna</mods:title>
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<mods:name id="7D39C8066754CD0258CDE3FCA6956FF8" type="personal">
<mods:role id="116B67EF6C459F0331FE46F73626D1F1">
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<mods:namePart id="C7403C5D22588E9FAF28BDEBD7C65C90">Menegon, Michele</mods:namePart>
</mods:name>
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<mods:namePart id="725CBDFBC86FB3964EAF1E734F104794">Davenport, Tim R. B.</mods:namePart>
</mods:name>
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<mods:namePart id="752A1F934AA8041B13D29EC960D187D9">Howell, Kim M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="38E58FB7216A426F7AAD6795F8442D86">text</mods:typeOfResource>
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<mods:title id="D50A77697423D36EC38EF4E3FE928726">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9B14EA4E6733399BA56C348CF51856D5">
<mods:date id="3B889135331F842D48232C404A49B255">2011</mods:date>
<mods:detail id="0B1B6C177248D5C89D26F50750A98B89" type="volume">
<mods:number id="32A7DEB030948436BF489A99D3EE6D58">3120</mods:number>
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<mods:classification id="D112D39D79ED75E12388FED824DD41DF">journal article</mods:classification>
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<mods:identifier id="F53DB3D18D28C3EB69190DBF426C9238" type="DOI">10.5281/zenodo.279365</mods:identifier>
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<mods:identifier id="31D90C7963054B06CBB237D798FAF645" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E60257D9953920D6C6E815020CA2353E" type="Zenodo-Dep">279365</mods:identifier>
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<treatment id="03B787851A477C5896E696B030E28CAC" ID-DOI="http://doi.org/10.5281/zenodo.5680229" ID-GBIF-Taxon="119425337" ID-Zenodo-Dep="5680229" LSID="urn:lsid:plazi:treatment:03B787851A477C5896E696B030E28CAC" httpUri="http://treatment.plazi.org/id/03B787851A477C5896E696B030E28CAC" lastPageId="7" lastPageNumber="50" pageId="1" pageNumber="44">
<subSubSection id="C30465181A477C5E96E696B032528CEB" pageId="1" pageNumber="44" type="nomenclature">
<paragraph id="8BA136931A477C5E96E696B030048CCA" blockId="1.[151,849,1194,1253]" box="[151,849,1194,1220]" pageId="1" pageNumber="44">
<heading id="D0E981FF1A477C5E96E696B030048CCA" bold="true" box="[151,849,1194,1220]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
<emphasis id="B96AEA811A477C5E96E696B030048CCA" bold="true" box="[151,849,1194,1220]" pageId="1" pageNumber="44">
<taxonomicName id="4C1E4D101A477C5E96E696B031B98CCA" authority="Menegon, Davenport &amp; Howell" authorityName="Menegon, Davenport &amp; Howell" box="[151,748,1194,1220]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae" status="sp. nov.">
<emphasis id="B96AEA811A477C5E96E696B0320B8CCA" bold="true" box="[151,350,1194,1220]" italics="true" pageId="1" pageNumber="44">Atheris matildae</emphasis>
Menegon, Davenport &amp; Howell
</taxonomicName>
<taxonomicNameLabel id="A25957FA1A477C5E948296B030048CCA" box="[755,849,1194,1220]" pageId="1" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA136931A477C5E96E696D732528CEB" blockId="1.[151,849,1194,1253]" box="[151,263,1229,1253]" pageId="1" pageNumber="44">
(
<figureCitation id="13252A161A477C5E96EE96D7338B8CEB" box="[159,222,1229,1253]" captionStart="FIGURE 1" captionStartId="2.[151,247,1971,1994]" captionTargetBox="[234,1331,304,1914]" captionTargetId="figure@2.[222,1341,285,1947]" captionTargetPageId="2" captionText="FIGURE 1. Comparison of general shape, size and body proportion between A. matildae holotype (left) and a fully grown specimen of A. ceratophora from Udzungwa (MTSN 7506, SVL = 54.2) (right) and an illustration of lateral head scalation." httpUri="https://zenodo.org/record/279366/files/figure.png" pageId="1" pageNumber="44">Fig. 1</figureCitation>
–3)
</paragraph>
</subSubSection>
<subSubSection id="C30465181A477C5E96E6970F30DE8E7F" pageId="1" pageNumber="44" type="materials_examined">
<paragraph id="8BA136931A477C5E96E6970F36788D5F" blockId="1.[151,1436,1301,2009]" pageId="1" pageNumber="44">
<emphasis id="B96AEA811A477C5E96E6970F325E8D23" bold="true" box="[151,267,1301,1325]" pageId="1" pageNumber="44">
<typeStatus id="54A588311A477C5E96E6970F32538D23" box="[151,262,1301,1325]" pageId="1" pageNumber="44" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult male,
<collectionCode id="ED0FAE561A477C5E97D9970F32A38D23" box="[424,502,1301,1325]" pageId="1" pageNumber="44">MTSN</collectionCode>
9344, collected in a forest fragment in Southern
<collectingCountry id="F30976031A477C5E924F970F37F08D23" box="[1086,1189,1301,1325]" name="Tanzania" pageId="1" pageNumber="44">Tanzania</collectingCountry>
, at about
<quantity id="4CE69B761A477C5E936C970F36208D23" box="[1309,1397,1301,1325]" metricMagnitude="3" metricUnit="m" metricValue="1.9949999999999999" pageId="1" pageNumber="44" unit="m" value="1995.0">1995 m</quantity>
by Omari Kibure and Obadia Mwaipungu in
<date id="FFA010531A477C5E941C972330418D5F" box="[621,788,1337,1361]" pageId="1" pageNumber="44" value="2009-02">February 2009</date>
; fixed in 70% EtOH, tissue fixed in 90% EtOH.
</paragraph>
<paragraph id="8BA136931A477C5E96B69747365D8D97" blockId="1.[151,1436,1301,2009]" pageId="1" pageNumber="44">
<emphasis id="B96AEA811A477C5E96B6974732138D7B" bold="true" box="[199,326,1373,1397]" pageId="1" pageNumber="44">
<typeStatus id="54A588311A477C5E96B6974732148D7B" box="[199,321,1373,1397]" pageId="1" pageNumber="44" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D18FD1A1A477C5E973C974732C98D7B" box="[333,412,1373,1397]" pageId="1" pageNumber="44" type="adult">2 adult</specimenCount>
males,
<collectionCode id="ED0FAE561A477C5E9785974731178D7B" box="[500,578,1373,1397]" pageId="1" pageNumber="44">MTSN</collectionCode>
9399 and
<collectionCode id="ED0FAE561A477C5E94CB9747305D8D7B" box="[698,776,1373,1397]" pageId="1" pageNumber="44">MTSN</collectionCode>
9418 and an immature
<collectionCode id="ED0FAE561A477C5E9266974737308D7B" box="[1047,1125,1373,1397]" pageId="1" pageNumber="44">MTSN</collectionCode>
9417 collected in
<date id="FFA010531A477C5E93469747339B8D97" pageId="1" pageNumber="44" value="2011-02">February 2011</date>
at the same locality as the
<typeStatus id="54A588311A477C5E978D979B31348D97" box="[508,609,1409,1433]" pageId="1" pageNumber="44" type="holotype">holotype</typeStatus>
, by Michele Menegon, Tim Davenport and Sophy Machaga.
</paragraph>
<paragraph id="8BA136931A477C5E96B697BF31028E0B" blockId="1.[151,1436,1301,2009]" pageId="1" pageNumber="44">
<emphasis id="B96AEA811A477C5E96B697BF32E98DB0" bold="true" box="[199,444,1445,1470]" pageId="1" pageNumber="44">Additional material.</emphasis>
<specimenCount id="9D18FD1A1A477C5E97B797BF31358DB3" box="[454,608,1445,1469]" pageId="1" pageNumber="44" type="generic">10 specimens</specimenCount>
collected at the
<typeStatus id="54A588311A477C5E955297BC30068DB0" box="[803,851,1446,1470]" pageId="1" pageNumber="44">type</typeStatus>
locality between
<date id="FFA010531A477C5E925097BF36778DB3" box="[1057,1314,1445,1469]" pageId="1" pageNumber="44" value="2011-03" valueMax="2011-04-31" valueMin="2011-03-00">March and April 2011</date>
and being kept alive for conservation purposes. 4 are males, 5 are females and 2 are immatures. Among them there is the adult female individual shown in Fig. 3.
</paragraph>
<paragraph id="8BA136931A477C5E96B6940830DE8E7F" blockId="1.[151,1436,1301,2009]" pageId="1" pageNumber="44">
<emphasis id="B96AEA811A477C5E96B6940832308E24" bold="true" box="[199,357,1553,1578]" pageId="1" pageNumber="44">
<typeStatus id="54A588311A477C5E96B6940832578E24" box="[199,258,1554,1578]" pageId="1" pageNumber="44">Type</typeStatus>
locality.
</emphasis>
Remote fragmented montane forest in Tanzania's Southern Highlands. Precise locality withheld until conservation insurance population secure. Additional information on the locality can be obtained for scientific purposes from the authors upon request. (www.atherismatildae.org)
</paragraph>
</subSubSection>
<subSubSection id="C30465181A477C5D96B69467379A88F9" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="diagnosis">
<paragraph id="8BA136931A477C5D96B69467379A88F9" blockId="1.[151,1436,1301,2009]" lastBlockId="2.[151,1436,151,247]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
<emphasis id="B96AEA811A477C5E96B6946732148E98" bold="true" box="[199,321,1661,1686]" pageId="1" pageNumber="44">Diagnosis.</emphasis>
<taxonomicName id="4C1E4D101A477C5E973B946731508E9B" box="[330,517,1661,1685]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae" status="sp. nov.">
<emphasis id="B96AEA811A477C5E973B946731508E9B" box="[330,517,1661,1685]" italics="true" pageId="1" pageNumber="44">Atheris matildae</emphasis>
</taxonomicName>
<emphasis id="B96AEA811A477C5E947F9467313F8E9B" bold="true" box="[526,618,1661,1685]" pageId="1" pageNumber="44">
<taxonomicNameLabel id="A25957FA1A477C5E947F9467313F8E9B" box="[526,618,1661,1685]" pageId="1" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from all other members of the genus except
<taxonomicName id="4C1E4D101A477C5E935B9467338F8EB7" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E935B9467338F8EB7" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
by the presence of two to three very enlarged erect, hornlike, supraciliary scales. It is distinguished from
<taxonomicName id="4C1E4D101A477C5E93F594BB32778ED3" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E93F594BB32778ED3" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
by the combination of the following morphological and molecular features, based on the data from
<specimenCount id="9D18FD1A1A477C5E93F194DF32598F0F" pageId="1" pageNumber="44" type="generic">69 specimens</specimenCount>
from all over the known range of
<taxonomicName id="4C1E4D101A477C5E94E494F330158F0F" box="[661,832,1769,1793]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E94E494F330158F0F" box="[661,832,1769,1793]" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
: (1) its larger size, TL of
<taxonomicName id="4C1E4D101A477C5E921894F337BE8F0F" box="[1129,1259,1769,1793]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A477C5E921894F337BE8F0F" box="[1129,1259,1769,1793]" italics="true" pageId="1" pageNumber="44">A. matildae</emphasis>
</taxonomicName>
<typeStatus id="54A588311A477C5E928294F036768F0C" box="[1267,1315,1770,1794]" pageId="1" pageNumber="44">type</typeStatus>
is
<quantity id="4CE69B761A477C5E933794F336C98F0F" box="[1350,1436,1769,1793]" metricMagnitude="-1" metricUnit="m" metricValue="6.43" pageId="1" pageNumber="44" unit="mm" value="643.0">643mm</quantity>
(the biggest male
<taxonomicName id="4C1E4D101A477C5E97119517315E8F2B" box="[352,523,1805,1829]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E97119517315E8F2B" box="[352,523,1805,1829]" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
ever recorded does not exceed
<quantity id="4CE69B761A477C5E951E951730908F2B" box="[879,965,1805,1829]" metricMagnitude="-1" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="44" unit="mm" value="510.0">510mm</quantity>
TL), (2) higher count of maximum transverse head scales (max.
<quantity id="4CE69B761A477C5E97D9952B32B78F47" box="[424,482,1841,1865]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="44" unit="in" value="20.0">20 in</quantity>
<taxonomicName id="4C1E4D101A477C5E9798952B31C68F47" box="[489,659,1841,1865]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E9798952B31C68F47" box="[489,659,1841,1865]" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
,
<quantity id="4CE69B761A477C5E94D3952B318E8F47" box="[674,731,1841,1865]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="1" pageNumber="44" unit="in" value="28.0">28 in</quantity>
<taxonomicName id="4C1E4D101A477C5E9492952B30318F47" box="[739,868,1841,1865]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A477C5E9492952B30318F47" box="[739,868,1841,1865]" italics="true" pageId="1" pageNumber="44">A. matildae</emphasis>
</taxonomicName>
), (3) four subequal suprarostral scales in
<emphasis id="B96AEA811A477C5E9348952B33938F63" italics="true" pageId="1" pageNumber="44">
<taxonomicName id="4C1E4D101A477C5E9348952B33978F63" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae">A. matildae</taxonomicName>
,
</emphasis>
the two central ones of the same size and the outer ones double in size in
<taxonomicName id="4C1E4D101A477C5E9228954F365D8F63" box="[1113,1288,1877,1901]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E9228954F365D8F63" box="[1113,1288,1877,1901]" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
, (4) marked difference in dorsal scale microdermatoglyphic pattern (irregular smooth surface in
<taxonomicName id="4C1E4D101A477C5E922D9563365C8F9F" box="[1116,1289,1913,1937]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A477C5E922D9563365C8F9F" box="[1116,1289,1913,1937]" italics="true" pageId="1" pageNumber="44">A. ceratophora</emphasis>
</taxonomicName>
, presence of papillae-like ridges in
<taxonomicName id="4C1E4D101A477C5E97E8958731498FBB" box="[409,540,1949,1973]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A477C5E97E8958731498FBB" box="[409,540,1949,1973]" italics="true" pageId="1" pageNumber="44">A. matildae</emphasis>
</taxonomicName>
), (5) in
<taxonomicName id="4C1E4D101A477C5E940E958730578FBB" box="[639,770,1949,1973]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A477C5E940E958730578FBB" box="[639,770,1949,1973]" italics="true" pageId="1" pageNumber="44">A. matildae</emphasis>
</taxonomicName>
, an extensive black marking across the frontal part of the mouth, including part of nasal, rostral, mental and few infralabial scales is often present, the above described colour feature has not been recorded in the examined
<taxonomicName id="4C1E4D101A447C5D948F928230FC88BE" box="[766,937,152,176]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="45" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A447C5D948F928230FC88BE" box="[766,937,152,176]" italics="true" pageId="2" pageNumber="45">A. ceratophora</emphasis>
</taxonomicName>
specimens and photographs. Genetic divergence of mitochondrial gene cytochrome b between
<taxonomicName id="4C1E4D101A447C5D948E92A630FB88DA" box="[767,942,188,212]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="45" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A447C5D948E92A630FB88DA" box="[767,942,188,212]" italics="true" pageId="2" pageNumber="45">A. ceratophora</emphasis>
</taxonomicName>
collected at
<typeStatus id="54A588311A447C5D923992A7372D88DB" box="[1096,1144,189,213]" pageId="2" pageNumber="45">type</typeStatus>
locality and
<taxonomicName id="4C1E4D101A447C5D936792A636C988DA" box="[1302,1436,188,212]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="45" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A447C5D936792A636C988DA" box="[1302,1436,188,212]" italics="true" pageId="2" pageNumber="45">A. matildae</emphasis>
</taxonomicName>
expressed as actual substitution difference is 3.18% based on uncorrected p-distance of 0.03180.
</paragraph>
</subSubSection>
<caption id="DF61661B1A447C5D96E695A9361E8FE6" httpUri="https://zenodo.org/record/279366/files/figure.png" pageId="2" pageNumber="45" targetBox="[234,1331,304,1914]" targetPageId="2">
<paragraph id="8BA136931A447C5D96E695A9361E8FE6" blockId="2.[151,1436,1971,2024]" pageId="2" pageNumber="45">
<emphasis id="B96AEA811A447C5D96E695A932458FC7" bold="true" box="[151,272,1971,1994]" pageId="2" pageNumber="45">FIGURE 1.</emphasis>
Comparison of general shape, size and body proportion between
<taxonomicName id="4C1E4D101A447C5D95D795AF374E8FC4" box="[934,1051,1972,1994]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="45" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A447C5D95D795AF374E8FC4" box="[934,1051,1972,1994]" italics="true" pageId="2" pageNumber="45">A. matildae</emphasis>
</taxonomicName>
holotype (left) and a fully grown specimen of
<taxonomicName id="4C1E4D101A447C5D968795C932C58FE6" box="[246,400,2002,2024]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="45" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A447C5D968795C932C58FE6" box="[246,400,2002,2024]" italics="true" pageId="2" pageNumber="45">A. ceratophora</emphasis>
</taxonomicName>
from Udzungwa (MTSN 7506, SVL=54.2) (right) and an illustration of lateral head scalation.
</paragraph>
</caption>
<subSubSection id="C30465181A457C5896B69282379E8AA9" lastPageId="7" lastPageNumber="50" pageId="3" pageNumber="46" type="materials_examined">
<paragraph id="8BA136931A457C5C96B69282317F8A32" blockId="3.[151,1437,151,572]" pageId="3" pageNumber="46">
<emphasis id="B96AEA811A457C5C96B6928231B288BE" bold="true" box="[199,743,151,176]" pageId="3" pageNumber="46">
<typeStatus id="54A588311A457C5C96B69282326A88BE" box="[199,319,152,176]" pageId="3" pageNumber="46" type="paratype">Paratypes</typeStatus>
and additional material variation.
</emphasis>
Details and meristics for the
<typeStatus id="54A588311A457C5C92339282372788BE" box="[1090,1138,152,176]" pageId="3" pageNumber="46">type</typeStatus>
series are summarized in
<tableCitation id="C69C03281A457C5C96E692A633BF88DA" box="[151,234,188,212]" captionStart="TABLE 1" captionStartId="4.[151,235,980,1001]" captionTargetBox="[151,1401,1113,1827]" captionTargetPageId="4" captionText="TABLE 1. Measurements and scalation of type-series of A. matildae, with data of A. ceratophora for comparison." httpUri="http://table.plazi.org/id/DF61661B1A427C5B96E691CE36498BE4" pageId="3" pageNumber="46" tableUuid="DF61661B1A427C5B96E691CE36498BE4">Table 1</tableCitation>
. A total of
<specimenCount id="9D18FD1A1A457C5C971B92A6315788DA" box="[362,514,188,212]" pageId="3" pageNumber="46" type="generic">13 specimens</specimenCount>
have been observed, three of them are
<typeStatus id="54A588311A457C5C95C892A7377D88DB" box="[953,1064,189,213]" pageId="3" pageNumber="46" type="paratype">paratypes</typeStatus>
. The most significant differences between
<typeStatus id="54A588311A457C5C968D92C5320B88F9" box="[252,350,223,247]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
and
<typeStatus id="54A588311A457C5C97E792FA32A288F6" box="[406,503,224,248]" pageId="3" pageNumber="46" type="paratype">paratype</typeStatus>
specimens are in body colouration (see Fig. 2). The young specimen
<collectionCode id="ED0FAE561A457C5C937F92C5360988F9" box="[1294,1372,223,247]" pageId="3" pageNumber="46">MTSN</collectionCode>
9417 tends to be more greenish, the zigzag ornamentation is more conspicuous and the top of the head is marbled in green/yellow. The two adult individuals, both males, are similar in colouration to
<typeStatus id="54A588311A457C5C9252933D37DD8931" box="[1059,1160,295,319]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
, with a back dorsum and a bright yellow zigzag dorsolateral pattern. Ten additional specimens have been recently collected and are being kept alive for conservation purposes. 5 are males, 5 are females, and 2 are young. A black patch around nasal, rostral, mental and first infralabials is present in most of the observed males but also in few females and immature individuals. Males in general tend to be darker with belly suffused with black. Adult females tend to be more yellow, in some cases with immaculate throat and belly; horn-like scales are yellow with black outer edges. Side of the head can be completely yellow or with black patches at the tip of the scales. In preservation the specimen retains the original colouration (see
<figureCitation id="13252A161A457C5C97A8903E314C8A32" box="[473,537,548,572]" captionStart="FIGURE 1" captionStartId="2.[151,247,1971,1994]" captionTargetBox="[234,1331,304,1914]" captionTargetId="figure@2.[222,1341,285,1947]" captionTargetPageId="2" captionText="FIGURE 1. Comparison of general shape, size and body proportion between A. matildae holotype (left) and a fully grown specimen of A. ceratophora from Udzungwa (MTSN 7506, SVL = 54.2) (right) and an illustration of lateral head scalation." httpUri="https://zenodo.org/record/279366/files/figure.png" pageId="3" pageNumber="46">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BA136931A457C5C96E695B5321D8FEA" blockId="3.[151,1436,1965,2021]" pageId="3" pageNumber="46">
<emphasis id="B96AEA811A457C5C96E695B532528FCD" bold="true" box="[151,263,1967,1987]" pageId="3" pageNumber="46">FIGURE 2.</emphasis>
<taxonomicName id="4C1E4D101A457C5C977F95B432D08FCA" box="[270,389,1966,1988]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A457C5C977F95B432D08FCA" box="[270,389,1966,1988]" italics="true" pageId="3" pageNumber="46">A. matildae</emphasis>
</taxonomicName>
<typeStatus id="54A588311A457C5C97FD95B432B88FCD" box="[396,493,1966,1987]" pageId="3" pageNumber="46" type="paratype">paratypes</typeStatus>
showing body shape, colouration and head details. Top - down:
<collectionCode id="ED0FAE561A457C5C92FA95B437848FCD" box="[1163,1233,1966,1987]" pageId="3" pageNumber="46">MTSN</collectionCode>
9399;
<collectionCode id="ED0FAE561A457C5C936A95B436348FCD" box="[1307,1377,1966,1987]" pageId="3" pageNumber="46">MTSN</collectionCode>
9418 and
<collectionCode id="ED0FAE561A457C5C96B395D5325D8FEA" box="[194,264,1999,2020]" pageId="3" pageNumber="46">MTSN</collectionCode>
9417.
</paragraph>
<paragraph id="8BA136931A427C5B96E69177324E8BAD" blockId="4.[151,1436,877,931]" pageId="4" pageNumber="47">
<emphasis id="B96AEA811A427C5B96E6917732538B8F" bold="true" box="[151,262,877,897]" pageId="4" pageNumber="47">FIGURE 3.</emphasis>
Illustration showing the variation in colour of the rostral and chin area, The black patches are present in many individuals of both sexes.
</paragraph>
<caption id="DF61661B1A427C5B96E691CE36498BE4" ID-Table-UUID="DF61661B1A427C5B96E691CE36498BE4" box="[151,1308,980,1002]" httpUri="http://table.plazi.org/id/DF61661B1A427C5B96E691CE36498BE4" pageId="4" pageNumber="47" targetBox="[151,1401,1113,1827]" targetIsTable="true" targetPageId="4">
<paragraph id="8BA136931A427C5B96E691CE36498BE4" blockId="4.[151,1401,980,1093]" box="[151,1308,980,1002]" pageId="4" pageNumber="47">
<emphasis id="B96AEA811A427C5B96E691CE32518BE7" bold="true" box="[151,260,980,1001]" pageId="4" pageNumber="47">TABLE 1.</emphasis>
Measurements and scalation of type-series of
<taxonomicName id="4C1E4D101A427C5B94AB91CF301A8BE4" box="[730,847,980,1002]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A427C5B94AB91CF301A8BE4" box="[730,847,980,1002]" italics="true" pageId="4" pageNumber="47">A. matildae</emphasis>
</taxonomicName>
, with data of
<taxonomicName id="4C1E4D101A427C5B95AA91CF37218BE4" box="[987,1140,980,1002]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A427C5B95AA91CF37218BE4" box="[987,1140,980,1002]" italics="true" pageId="4" pageNumber="47">A. ceratophora</emphasis>
</taxonomicName>
for comparison.
</paragraph>
</caption>
<paragraph id="8BA136931A427C5B9452961D362C8C12" blockId="4.[151,1401,980,1093]" box="[547,1401,1030,1052]" pageId="4" pageNumber="47">
<emphasis id="B96AEA811A427C5B9452961D362C8C12" box="[547,1401,1030,1052]" italics="true" pageId="4" pageNumber="47">
<taxonomicName id="4C1E4D101A427C5B9452961D37F98C12" box="[547,1196,1030,1052]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="matildae">A. matildae A. matildae A. matildae A. matildae</taxonomicName>
<taxonomicName id="4C1E4D101A427C5B9291961D362C8C12" box="[1248,1401,1030,1052]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="ceratophora">A. ceratophora</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA136931A427C5A96EE963536038A51" blockId="4.[151,1401,980,1093]" lastBlockId="5.[151,1437,151,607]" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
Specimen number
<collectionCode id="ED0FAE561A427C5B9452962A313C8C4B" box="[547,617,1072,1093]" pageId="4" pageNumber="47">MTSN</collectionCode>
9344
<collectionCode id="ED0FAE561A427C5B94AB962A30758C4B" box="[730,800,1072,1093]" pageId="4" pageNumber="47">MTSN</collectionCode>
9417
<collectionCode id="ED0FAE561A427C5B95FA962A30848C4B" box="[907,977,1072,1093]" pageId="4" pageNumber="47">MTSN</collectionCode>
9418
<collectionCode id="ED0FAE561A427C5B9246962A37288C4B" box="[1079,1149,1072,1093]" pageId="4" pageNumber="47">MTSN</collectionCode>
9399 n = 53
<emphasis id="B96AEA811A427C5B96B6957032BD8F8D" bold="true" box="[199,488,1898,1923]" pageId="4" pageNumber="47">
Description of
<typeStatus id="54A588311A427C5B970B957132B68F8D" box="[378,483,1899,1923]" pageId="4" pageNumber="47" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male preserved in 70% EtOH. Snout-vent length (SVL)
<quantity id="4CE69B761A427C5B92CA957036648F8C" box="[1211,1329,1898,1922]" metricMagnitude="-1" metricUnit="m" metricValue="5.407" pageId="4" pageNumber="47" unit="mm" value="540.7">540.7 mm</quantity>
, tail 96.0 mm, rostral width
<quantity id="4CE69B761A427C5B9719959432948FA8" box="[360,449,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="47" unit="mm" value="2.9">2.9 mm</quantity>
, rostral height
<quantity id="4CE69B761A427C5B941A959431908FA8" box="[619,709,1934,1958]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="47" unit="mm" value="0.8">0.8 mm</quantity>
; eye diameter (vertical)
<quantity id="4CE69B761A427C5B95A9959437678FA8" box="[984,1074,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="47" unit="mm" value="3.2">3.2 mm</quantity>
; snout to eye
<quantity id="4CE69B761A427C5B92BF9594367D8FA8" box="[1230,1320,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="47" unit="mm" value="3.3">3.3 mm</quantity>
. A heavybodied forest viper, sub-quadrangular in cross-section, with a rather thick prehensile tail (SVL/Tail approximately 5.7 times); head pear-shaped, with a very distinct neck, rounded snout and swollen supraorbital region that does bear two/three elongated, horn-like scales; eyes relatively large, laterally placed, and with a horizontal diameter approximately 3/4 of the snout length. Crown of head covered in small scales, slightly larger over the temporal region (maximum transverse head scales—28); they bear a prominent keel and become mucronate over the head; the rostral is flattened, rectangular, about 3.5 times broader than high, contacting first supralabials and four small; unkeeled, roundish, subequal suprarostrals, nasal wider than high, with raised, embossed posterior edge, nostril circular and approximately in the centre of the nasal; internasals 5, all strongly keeled; interorbitals 9, keeled; circumoculars 16–16, not keeled but terminating in black blunt knobs; 1 row of suboculars present; circumoculars separated from nasals by two to three rows of feebly knobbed scales; a row of three irregular scales, bordering supralabials between nasal and lower circumoculars; supralabials 10–10, the first three smaller, and 6–8 with a swollen lower edge; infralabials 10–10, posteriormost with swollen upper edge and first in contact at the midline behind the mental; mental triangular, approximately twice as wide as deep; gulars bordering chin shields feeblykeeled, but prominently keeled towards the rictus; 2 preventrals, first largest; 150 ventrals; 49 undivided subcaudals (including spine); anal entire; 25 rows dorsal scales anteriorly, 26 rows at midbody, 19 rows posteriorly.
</paragraph>
<paragraph id="8BA136931A427C5B96EE964036238F2D" pageId="4" pageNumber="47">
<table id="F91EC4331A4283A096E69643362C8F2D" box="[151,1401,1113,1827]" gridcols="6" gridrows="17" pageId="4" pageNumber="47">
<tr id="352E34D11A4283A096E69643362C8C61" box="[151,1401,1113,1135]" gridrow="0" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6964332BB8C61" box="[151,494,1113,1135]" gridcol="0" gridrow="0" pageId="4" pageNumber="47">Type</th>
<th id="76FF5DAD1A4283A09452964331FB8C61" box="[547,686,1113,1135]" gridcol="1" gridrow="0" pageId="4" pageNumber="47">Holytype</th>
<th id="76FF5DAD1A4283A094AB964330318C61" box="[730,868,1113,1135]" gridcol="2" gridrow="0" pageId="4" pageNumber="47">Paratype</th>
<th id="76FF5DAD1A4283A095FA964337438C61" box="[907,1046,1113,1135]" gridcol="3" gridrow="0" pageId="4" pageNumber="47">Paratype</th>
<th id="76FF5DAD1A4283A09246964337948C61" box="[1079,1217,1113,1135]" gridcol="4" gridrow="0" pageId="4" pageNumber="47">Paratype</th>
<th id="76FF5DAD1A4283A092AE9643362C8C61" box="[1247,1401,1113,1135]" gridcol="5" gridrow="0" pageId="4" pageNumber="47">N/A</th>
</tr>
<tr id="352E34D11A4283A096E69698362C8C96" box="[151,1401,1154,1176]" gridrow="1" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6969832BB8C96" box="[151,494,1154,1176]" gridcol="0" gridrow="1" pageId="4" pageNumber="47">Sex</th>
<td id="76FF5DAD1A4283A09452969831FB8C96" box="[547,686,1154,1176]" gridcol="1" gridrow="1" pageId="4" pageNumber="47">Male</td>
<td id="76FF5DAD1A4283A094AB969830318C96" box="[730,868,1154,1176]" gridcol="2" gridrow="1" pageId="4" pageNumber="47">Inmmature</td>
<td id="76FF5DAD1A4283A095FA969837438C96" box="[907,1046,1154,1176]" gridcol="3" gridrow="1" pageId="4" pageNumber="47">Male</td>
<td id="76FF5DAD1A4283A09246969837948C96" box="[1079,1217,1154,1176]" gridcol="4" gridrow="1" pageId="4" pageNumber="47">Male</td>
<td id="76FF5DAD1A4283A092AE9698362C8C96" box="[1247,1401,1154,1176]" gridcol="5" gridrow="1" pageId="4" pageNumber="47">Males only</td>
</tr>
<tr id="352E34D11A4283A096E696B1362C8CCF" box="[151,1401,1195,1217]" gridrow="2" pageId="4" pageNumber="47" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76FF5DAD1A4283A096E696B132BB8CCF" box="[151,494,1195,1217]" gridcol="0" gridrow="2" pageId="4" pageNumber="47">Head Lepidosis</th>
</tr>
<tr id="352E34D11A4283A096E696CF362C8CE5" box="[151,1401,1237,1259]" gridrow="3" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E696CF32BB8CE5" box="[151,494,1237,1259]" gridcol="0" gridrow="3" pageId="4" pageNumber="47">Suprarostrals (SRO)</th>
<td id="76FF5DAD1A4283A0945296CF31FB8CE5" box="[547,686,1237,1259]" gridcol="1" gridrow="3" pageId="4" pageNumber="47">4</td>
<td id="76FF5DAD1A4283A094AB96CF30318CE5" box="[730,868,1237,1259]" gridcol="2" gridrow="3" pageId="4" pageNumber="47">4</td>
<td id="76FF5DAD1A4283A095FA96CF37438CE5" box="[907,1046,1237,1259]" gridcol="3" gridrow="3" pageId="4" pageNumber="47">4</td>
<td id="76FF5DAD1A4283A0924696CF37948CE5" box="[1079,1217,1237,1259]" gridcol="4" gridrow="3" pageId="4" pageNumber="47">4</td>
<td id="76FF5DAD1A4283A092AE96CF362C8CE5" box="[1247,1401,1237,1259]" gridcol="5" gridrow="3" pageId="4" pageNumber="47">5–9</td>
</tr>
<tr id="352E34D11A4283A096E696E4362C8D1B" box="[151,1401,1278,1301]" gridrow="4" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E696E432BB8D1B" box="[151,494,1278,1301]" gridcol="0" gridrow="4" pageId="4" pageNumber="47">Interorbitals (IOS)</th>
<td id="76FF5DAD1A4283A0945296E431FB8D1B" box="[547,686,1278,1301]" gridcol="1" gridrow="4" pageId="4" pageNumber="47">12</td>
<td id="76FF5DAD1A4283A094AB96E430318D1B" box="[730,868,1278,1301]" gridcol="2" gridrow="4" pageId="4" pageNumber="47">12</td>
<td id="76FF5DAD1A4283A095FA96E437438D1B" box="[907,1046,1278,1301]" gridcol="3" gridrow="4" pageId="4" pageNumber="47">121</td>
<td id="76FF5DAD1A4283A0924696E437948D1B" box="[1079,1217,1278,1301]" gridcol="4" gridrow="4" pageId="4" pageNumber="47">12</td>
<td id="76FF5DAD1A4283A092AE96E4362C8D1B" box="[1247,1401,1278,1301]" gridcol="5" gridrow="4" pageId="4" pageNumber="47">7–11</td>
</tr>
<tr id="352E34D11A4283A096E6973D362C8D53" box="[151,1401,1319,1373]" gridrow="5" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6973D32BB8D53" box="[151,494,1319,1373]" gridcol="0" gridrow="5" pageId="4" pageNumber="47">Maximum transverse head scales (MTHS)</th>
<td id="76FF5DAD1A4283A09452973D31FB8D53" box="[547,686,1319,1373]" gridcol="1" gridrow="5" pageId="4" pageNumber="47">28</td>
<td id="76FF5DAD1A4283A094AB973D30318D53" box="[730,868,1319,1373]" gridcol="2" gridrow="5" pageId="4" pageNumber="47">28</td>
<td id="76FF5DAD1A4283A095FA973D37438D53" box="[907,1046,1319,1373]" gridcol="3" gridrow="5" pageId="4" pageNumber="47">27</td>
<td id="76FF5DAD1A4283A09246973D37948D53" box="[1079,1217,1319,1373]" gridcol="4" gridrow="5" pageId="4" pageNumber="47">28</td>
<td id="76FF5DAD1A4283A092AE973D362C8D53" box="[1247,1401,1319,1373]" gridcol="5" gridrow="5" pageId="4" pageNumber="47">19–20</td>
</tr>
<tr id="352E34D11A4283A096E69775362C8D88" box="[151,1401,1391,1414]" gridrow="6" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6977532BB8D88" box="[151,494,1391,1414]" gridcol="0" gridrow="6" pageId="4" pageNumber="47">Circumorbital scales (COS)</th>
<td id="76FF5DAD1A4283A09452977531FB8D88" box="[547,686,1391,1414]" gridcol="1" gridrow="6" pageId="4" pageNumber="47">16</td>
<td id="76FF5DAD1A4283A094AB977530318D88" box="[730,868,1391,1414]" gridcol="2" gridrow="6" pageId="4" pageNumber="47">16</td>
<td id="76FF5DAD1A4283A095FA977537438D88" box="[907,1046,1391,1414]" gridcol="3" gridrow="6" pageId="4" pageNumber="47">16</td>
<td id="76FF5DAD1A4283A09246977537948D88" box="[1079,1217,1391,1414]" gridcol="4" gridrow="6" pageId="4" pageNumber="47">162</td>
<td id="76FF5DAD1A4283A092AE9775362C8D88" box="[1247,1401,1391,1414]" gridcol="5" gridrow="6" pageId="4" pageNumber="47">13–19</td>
</tr>
<tr id="352E34D11A4283A096E69783362C8DA1" box="[151,1401,1433,1455]" gridrow="7" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6978332BB8DA1" box="[151,494,1433,1455]" gridcol="0" gridrow="7" pageId="4" pageNumber="47">Interoculabials (IOL)</th>
<td id="76FF5DAD1A4283A09452978331FB8DA1" box="[547,686,1433,1455]" gridcol="1" gridrow="7" pageId="4" pageNumber="47">1</td>
<td id="76FF5DAD1A4283A094AB978330318DA1" box="[730,868,1433,1455]" gridcol="2" gridrow="7" pageId="4" pageNumber="47">1–2</td>
<td id="76FF5DAD1A4283A095FA978337438DA1" box="[907,1046,1433,1455]" gridcol="3" gridrow="7" pageId="4" pageNumber="47">1</td>
<td id="76FF5DAD1A4283A09246978337948DA1" box="[1079,1217,1433,1455]" gridcol="4" gridrow="7" pageId="4" pageNumber="47">1–2</td>
<td id="76FF5DAD1A4283A092AE9783362C8DA1" box="[1247,1401,1433,1455]" gridcol="5" gridrow="7" pageId="4" pageNumber="47">0–1</td>
</tr>
<tr id="352E34D11A4283A096E697D8362C8DD7" box="[151,1401,1474,1497]" gridrow="8" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E697D832BB8DD7" box="[151,494,1474,1497]" gridcol="0" gridrow="8" pageId="4" pageNumber="47">Interocunasals (ION)</th>
<td id="76FF5DAD1A4283A0945297D831FB8DD7" box="[547,686,1474,1497]" gridcol="1" gridrow="8" pageId="4" pageNumber="47">3–4</td>
<td id="76FF5DAD1A4283A094AB97D830318DD7" box="[730,868,1474,1497]" gridcol="2" gridrow="8" pageId="4" pageNumber="47">4</td>
<td id="76FF5DAD1A4283A095FA97D837438DD7" box="[907,1046,1474,1497]" gridcol="3" gridrow="8" pageId="4" pageNumber="47">4</td>
<td id="76FF5DAD1A4283A0924697D837948DD7" box="[1079,1217,1474,1497]" gridcol="4" gridrow="8" pageId="4" pageNumber="47">3–4</td>
<td id="76FF5DAD1A4283A092AE97D8362C8DD7" box="[1247,1401,1474,1497]" gridcol="5" gridrow="8" pageId="4" pageNumber="47">2–4</td>
</tr>
<tr id="352E34D11A4283A096E697F1362C8E0C" box="[151,1401,1515,1538]" gridrow="9" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E697F132BB8E0C" box="[151,494,1515,1538]" gridcol="0" gridrow="9" pageId="4" pageNumber="47">Supralabials (SL)</th>
<td id="76FF5DAD1A4283A0945297F131FB8E0C" box="[547,686,1515,1538]" gridcol="1" gridrow="9" pageId="4" pageNumber="47">10</td>
<td id="76FF5DAD1A4283A094AB97F130318E0C" box="[730,868,1515,1538]" gridcol="2" gridrow="9" pageId="4" pageNumber="47">10</td>
<td id="76FF5DAD1A4283A095FA97F137438E0C" box="[907,1046,1515,1538]" gridcol="3" gridrow="9" pageId="4" pageNumber="47">10</td>
<td id="76FF5DAD1A4283A0924697F137948E0C" box="[1079,1217,1515,1538]" gridcol="4" gridrow="9" pageId="4" pageNumber="47">10</td>
<td id="76FF5DAD1A4283A092AE97F1362C8E0C" box="[1247,1401,1515,1538]" gridcol="5" gridrow="9" pageId="4" pageNumber="47">7–11</td>
</tr>
<tr id="352E34D11A4283A096E6940F362C8E25" box="[151,1401,1557,1579]" gridrow="10" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6940F32BB8E25" box="[151,494,1557,1579]" gridcol="0" gridrow="10" pageId="4" pageNumber="47">Infralabials (IL)</th>
<td id="76FF5DAD1A4283A09452940F31FB8E25" box="[547,686,1557,1579]" gridcol="1" gridrow="10" pageId="4" pageNumber="47">11</td>
<td id="76FF5DAD1A4283A094AB940F30318E25" box="[730,868,1557,1579]" gridcol="2" gridrow="10" pageId="4" pageNumber="47">11</td>
<td id="76FF5DAD1A4283A095FA940F37438E25" box="[907,1046,1557,1579]" gridcol="3" gridrow="10" pageId="4" pageNumber="47">11</td>
<td id="76FF5DAD1A4283A09246940F37948E25" box="[1079,1217,1557,1579]" gridcol="4" gridrow="10" pageId="4" pageNumber="47">11</td>
<td id="76FF5DAD1A4283A092AE940F362C8E25" box="[1247,1401,1557,1579]" gridcol="5" gridrow="10" pageId="4" pageNumber="47">8–12</td>
</tr>
<tr id="352E34D11A4283A096E69424362C8E5A" box="[151,1401,1598,1620]" gridrow="11" pageId="4" pageNumber="47" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76FF5DAD1A4283A096E6942432BB8E5A" box="[151,494,1598,1620]" gridcol="0" gridrow="11" pageId="4" pageNumber="47">Body scalation</th>
</tr>
<tr id="352E34D11A4283A096E6947D362C8E70" box="[151,1401,1639,1662]" gridrow="12" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6947D32BB8E70" box="[151,494,1639,1662]" gridcol="0" gridrow="12" pageId="4" pageNumber="47">Midbody Scale Rows (MSR)</th>
<td id="76FF5DAD1A4283A09452947D31FB8E70" box="[547,686,1639,1662]" gridcol="1" gridrow="12" pageId="4" pageNumber="47">26</td>
<td id="76FF5DAD1A4283A094AB947D30318E70" box="[730,868,1639,1662]" gridcol="2" gridrow="12" pageId="4" pageNumber="47">27</td>
<td id="76FF5DAD1A4283A095FA947D37438E70" box="[907,1046,1639,1662]" gridcol="3" gridrow="12" pageId="4" pageNumber="47">26</td>
<td id="76FF5DAD1A4283A09246947D37948E70" box="[1079,1217,1639,1662]" gridcol="4" gridrow="12" pageId="4" pageNumber="47">26</td>
<td id="76FF5DAD1A4283A092AE947D362C8E70" box="[1247,1401,1639,1662]" gridcol="5" gridrow="12" pageId="4" pageNumber="47">19–25</td>
</tr>
<tr id="352E34D11A4283A096E6948B362C8EA9" box="[151,1401,1681,1703]" gridrow="13" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6948B32BB8EA9" box="[151,494,1681,1703]" gridcol="0" gridrow="13" pageId="4" pageNumber="47">Ventrals</th>
<td id="76FF5DAD1A4283A09452948B31FB8EA9" box="[547,686,1681,1703]" gridcol="1" gridrow="13" pageId="4" pageNumber="47">150</td>
<td id="76FF5DAD1A4283A094AB948B30318EA9" box="[730,868,1681,1703]" gridcol="2" gridrow="13" pageId="4" pageNumber="47">142</td>
<td id="76FF5DAD1A4283A095FA948B37438EA9" box="[907,1046,1681,1703]" gridcol="3" gridrow="13" pageId="4" pageNumber="47">149</td>
<td id="76FF5DAD1A4283A09246948B37948EA9" box="[1079,1217,1681,1703]" gridcol="4" gridrow="13" pageId="4" pageNumber="47">150</td>
<td id="76FF5DAD1A4283A092AE948B362C8EA9" box="[1247,1401,1681,1703]" gridcol="5" gridrow="13" pageId="4" pageNumber="47">136–150</td>
</tr>
<tr id="352E34D11A4283A096E694A0362C8EDE" box="[151,1401,1722,1744]" gridrow="14" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E694A032BB8EDE" box="[151,494,1722,1744]" gridcol="0" gridrow="14" pageId="4" pageNumber="47">Subcaudals</th>
<td id="76FF5DAD1A4283A0945294A031FB8EDE" box="[547,686,1722,1744]" gridcol="1" gridrow="14" pageId="4" pageNumber="47">49</td>
<td id="76FF5DAD1A4283A094AB94A030318EDE" box="[730,868,1722,1744]" gridcol="2" gridrow="14" pageId="4" pageNumber="47">44</td>
<td id="76FF5DAD1A4283A095FA94A037438EDE" box="[907,1046,1722,1744]" gridcol="3" gridrow="14" pageId="4" pageNumber="47">50</td>
<td id="76FF5DAD1A4283A0924694A037948EDE" box="[1079,1217,1722,1744]" gridcol="4" gridrow="14" pageId="4" pageNumber="47">49</td>
<td id="76FF5DAD1A4283A092AE94A0362C8EDE" box="[1247,1401,1722,1744]" gridcol="5" gridrow="14" pageId="4" pageNumber="47">49–58</td>
</tr>
<tr id="352E34D11A4283A096E694F9362C8EF4" box="[151,1401,1763,1786]" gridrow="15" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E694F932BB8EF4" box="[151,494,1763,1786]" gridcol="0" gridrow="15" pageId="4" pageNumber="47">Max length (SVL+Tail) in mm</th>
<td id="76FF5DAD1A4283A0945294F931FB8EF4" box="[547,686,1763,1786]" gridcol="1" gridrow="15" pageId="4" pageNumber="47">643 (547+96)</td>
<td id="76FF5DAD1A4283A094AB94F930318EF4" box="[730,868,1763,1786]" gridcol="2" gridrow="15" pageId="4" pageNumber="47">391 (334+57)</td>
<td id="76FF5DAD1A4283A095FA94F937438EF4" box="[907,1046,1763,1786]" gridcol="3" gridrow="15" pageId="4" pageNumber="47">625 (532+93)</td>
<td id="76FF5DAD1A4283A0924694F937948EF4" box="[1079,1217,1763,1786]" gridcol="4" gridrow="15" pageId="4" pageNumber="47">631 (538+93)</td>
<td id="76FF5DAD1A4283A092AE94F9362C8EF4" box="[1247,1401,1763,1786]" gridcol="5" gridrow="15" pageId="4" pageNumber="47">510 (416+94)</td>
</tr>
<tr id="352E34D11A4283A096E69517362C8F2D" box="[151,1401,1805,1827]" gridrow="16" pageId="4" pageNumber="47">
<th id="76FF5DAD1A4283A096E6951732BB8F2D" box="[151,494,1805,1827]" gridcol="0" gridrow="16" pageId="4" pageNumber="47">SVL/Tail</th>
<td id="76FF5DAD1A4283A09452951731FB8F2D" box="[547,686,1805,1827]" gridcol="1" gridrow="16" pageId="4" pageNumber="47">5.7</td>
<td id="76FF5DAD1A4283A094AB951730318F2D" box="[730,868,1805,1827]" gridcol="2" gridrow="16" pageId="4" pageNumber="47">5.85</td>
<td id="76FF5DAD1A4283A095FA951737438F2D" box="[907,1046,1805,1827]" gridcol="3" gridrow="16" pageId="4" pageNumber="47">5.7</td>
<td id="76FF5DAD1A4283A09246951737948F2D" box="[1079,1217,1805,1827]" gridcol="4" gridrow="16" pageId="4" pageNumber="47">5.78</td>
<td id="76FF5DAD1A4283A092AE9517362C8F2D" box="[1247,1401,1805,1827]" gridcol="5" gridrow="16" pageId="4" pageNumber="47">5.4 – 5.8 range</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA136931A437C5A96E6949630658ECC" blockId="5.[151,1436,1674,1730]" pageId="5" pageNumber="48">
<emphasis id="B96AEA811A437C5A96E6949632538EAE" bold="true" box="[151,262,1676,1696]" pageId="5" pageNumber="48">FIGURE 4.</emphasis>
Dorsal scale microdermatoglyphic pattern of
<taxonomicName id="4C1E4D101A437C5A94A49491301E8EAE" box="[725,843,1674,1696]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A437C5A94A49491301E8EAE" box="[725,843,1674,1696]" italics="true" pageId="5" pageNumber="48">A. matildae</emphasis>
</taxonomicName>
(A–B) and
<taxonomicName id="4C1E4D101A437C5A95B39491370E8EAE" box="[962,1115,1674,1696]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A437C5A95B39491370E8EAE" box="[962,1115,1674,1696]" italics="true" pageId="5" pageNumber="48">A. ceratophora</emphasis>
</taxonomicName>
: (C–D). Note in B the papillaelike ridges covering the keratin layers on the scale of
<emphasis id="B96AEA811A437C5A94C594B630658ECC" box="[692,816,1708,1730]" italics="true" pageId="5" pageNumber="48">
<taxonomicName id="4C1E4D101A437C5A94C594B630798ECC" box="[692,812,1708,1730]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="matildae">A. matildae</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BA136931A437C5A96B694EB30BA8FB3" blockId="5.[151,1436,1777,1981]" pageId="5" pageNumber="48">
<emphasis id="B96AEA811A437C5A96B694EB32078F04" bold="true" box="[199,338,1777,1802]" pageId="5" pageNumber="48">Hemipenes.</emphasis>
Both hemipenes are only partially everted. They resemble the
<taxonomicName id="4C1E4D101A437C5A926194EB37EF8F07" box="[1040,1210,1777,1801]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A437C5A926194EB37EF8F07" box="[1040,1210,1777,1801]" italics="true" pageId="5" pageNumber="48">A. ceratophora</emphasis>
</taxonomicName>
one as described by Emmrich (1997). The sulcus is bifurcate on a typically divided organ. The extreme basal area is naked, followed by an area with enlarged proximal spines, most prominently on the outer side of each lobe. Towards the apex, on the inner side of the sulcus, there are a few smaller scattered spines, while the distal area seems to be characterized by the lack of clearly differentiated ornamentation. A more detailed description of hemipenial morphology will be possible when a fully everted hemipenis becomes available for examination.
</paragraph>
<caption id="DF61661B1A407C5996E6976E327C8DA2" httpUri="https://zenodo.org/record/279367/files/figure.png" pageId="6" pageNumber="49" targetBox="[229,1350,198,1362]" targetPageId="6">
<paragraph id="8BA136931A407C5996E6976E327C8DA2" blockId="6.[151,1436,1396,1452]" pageId="6" pageNumber="49">
<emphasis id="B96AEA811A407C5996E6976E325E8D84" bold="true" box="[151,267,1396,1419]" pageId="6" pageNumber="49">FIGURE 5.</emphasis>
Map of the
<taxonomicName id="4C1E4D101A407C5997FB976E32868D84" box="[394,467,1396,1418]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B96AEA811A407C5997FB976E32868D84" box="[394,467,1396,1418]" italics="true" pageId="6" pageNumber="49">Atheris</emphasis>
</taxonomicName>
species occurring in Tanzania (the occurrence of
<taxonomicName id="4C1E4D101A407C5995BE976F376E8D84" box="[975,1083,1396,1418]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="nitschei">
<emphasis id="B96AEA811A407C5995BE976F376E8D84" box="[975,1083,1396,1418]" italics="true" pageId="6" pageNumber="49">A. nitschei</emphasis>
</taxonomicName>
within Tanzania borders still need confirmation).
</paragraph>
</caption>
<paragraph id="8BA136931A407C5996E695AA325D8FE8" blockId="6.[151,1436,1967,2022]" pageId="6" pageNumber="49">
<emphasis id="B96AEA811A407C5996E695AA32538FCA" bold="true" box="[151,262,1968,1988]" pageId="6" pageNumber="49">FIGURE 6.</emphasis>
The forest fragment in the Southern Highlands of
<collectingCountry id="F30976031A407C59957495B530358FCB" box="[773,864,1967,1989]" name="Tanzania" pageId="6" pageNumber="49">Tanzania</collectingCountry>
where
<taxonomicName id="4C1E4D101A407C5995DB95AA374A8FCB" box="[938,1055,1967,1989]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A407C5995DB95AA374A8FCB" box="[938,1055,1967,1989]" italics="true" pageId="6" pageNumber="49">A. matildae</emphasis>
</taxonomicName>
was collected, and a detail of the forest canopy.
</paragraph>
<paragraph id="8BA136931A417C5896B6928D30368989" blockId="7.[151,1437,151,1187]" pageId="7" pageNumber="50">
<emphasis id="B96AEA811A417C5896B6928D320A88BE" bold="true" box="[199,351,151,176]" pageId="7" pageNumber="50">Colouration.</emphasis>
Dorsally it appears as a black snake with bright yellow dorso-lateral zig-zag lines. Flanks are marbled in yellow. Dorsum of the head is almost entirely black with scattered yellow scales or groups of scales, sides of the head are mainly yellow with an irregular longitudinal black marking. An extensive black marking across the frontal part of the mouth, including part of nasal, rostral, mental and few infralabial scales is present, by contrast it delimitates an inverted pale triangle. Eyes are light olive green (in life). Throat is yellow; belly is pale yellow to greyish-green, suffused by black speckling; horn-like scales are yellow with black outer edges. In preservation the specimen retains the original colouration (see
<figureCitation id="13252A161A417C589560937530078989" box="[785,850,367,391]" captionStart="FIGURE 1" captionStartId="2.[151,247,1971,1994]" captionTargetBox="[234,1331,304,1914]" captionTargetId="figure@2.[222,1341,285,1947]" captionTargetPageId="2" captionText="FIGURE 1. Comparison of general shape, size and body proportion between A. matildae holotype (left) and a fully grown specimen of A. ceratophora from Udzungwa (MTSN 7506, SVL = 54.2) (right) and an illustration of lateral head scalation." httpUri="https://zenodo.org/record/279366/files/figure.png" pageId="7" pageNumber="50">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BA136931A417C5896B6938E379E8AA9" blockId="7.[151,1437,151,1187]" pageId="7" pageNumber="50">
<emphasis id="B96AEA811A417C5896B6938E312689A2" bold="true" box="[199,627,403,428]" pageId="7" pageNumber="50">Dorsal scale microdermatoglyphics.</emphasis>
The surface microstructure of several scales from midbody and the last third of the body of two specimens of
<taxonomicName id="4C1E4D101A417C58944F93A231EA89DE" box="[574,703,440,464]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C58944F93A231EA89DE" box="[574,703,440,464]" italics="true" pageId="7" pageNumber="50">A. matildae</emphasis>
</taxonomicName>
(
<collectionCode id="ED0FAE561A417C5894BC93AD304889C1" box="[717,797,439,463]" pageId="7" pageNumber="50">MTSN</collectionCode>
9344, 9417) and three specimens of
<taxonomicName id="4C1E4D101A417C5892C793A2360B89DE" box="[1206,1374,440,464]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A417C5892C793A2360B89DE" box="[1206,1374,440,464]" italics="true" pageId="7" pageNumber="50">A. ceratophora</emphasis>
</taxonomicName>
from Usambara Mountains (
<collectionCode id="ED0FAE561A417C5897E493C632B089FA" box="[405,485,476,500]" pageId="7" pageNumber="50">MTSN</collectionCode>
5117, 5118, 5121) were examined by scanning electron microscopy, in order to evaluate the intra- and inter-specific differences. Two-dimensional classes of microdermatoglyphics were identified; the coarser one consists of juxtaposed or imbricated layers of keratin with a raised edge, forming a ‘scaly background’. This layer is shared by both
<taxonomicName id="4C1E4D101A417C589434905231BA8A6E" box="[581,751,584,608]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A417C589434905231BA8A6E" box="[581,751,584,608]" italics="true" pageId="7" pageNumber="50">A. ceratophora</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D101A417C589558905230FE8A6E" box="[809,939,584,608]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C589558905230FE8A6E" box="[809,939,584,608]" italics="true" pageId="7" pageNumber="50">A. matildae</emphasis>
</taxonomicName>
. At greater magnification (4000x) a further pattern of microdermatoglyphics is visible in
<taxonomicName id="4C1E4D101A417C5894E4907630438A8A" box="[661,790,620,644]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C5894E4907630438A8A" box="[661,790,620,644]" italics="true" pageId="7" pageNumber="50">A. matildae</emphasis>
</taxonomicName>
, where papillae-like ridges cover the entire surface of the keratin layer. The latter ornamentation is absent in
<taxonomicName id="4C1E4D101A417C5894A0908A302E8AA6" box="[721,891,656,680]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="ceratophora">
<emphasis id="B96AEA811A417C5894A0908A302E8AA6" box="[721,891,656,680]" italics="true" pageId="7" pageNumber="50">A. ceratophora</emphasis>
</taxonomicName>
specimens from
<typeStatus id="54A588311A417C58924B908A373F8AA6" box="[1082,1130,656,680]" pageId="7" pageNumber="50">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C30465181A417C5896B690A937F08C54" pageId="7" pageNumber="50" type="distribution">
<paragraph id="8BA136931A417C5896B690A937F08C54" blockId="7.[151,1437,151,1187]" pageId="7" pageNumber="50">
<emphasis id="B96AEA811A417C5896B690A931668AC2" bold="true" box="[199,563,691,716]" pageId="7" pageNumber="50">Distribution and conservation.</emphasis>
<taxonomicName id="4C1E4D101A417C58944B90AE31A18AC2" box="[570,756,692,716]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C58944B90AE31A18AC2" box="[570,756,692,716]" italics="true" pageId="7" pageNumber="50">Atheris matildae</emphasis>
</taxonomicName>
is currently known only from the
<typeStatus id="54A588311A417C58920090AF37F48AC3" box="[1137,1185,693,717]" pageId="7" pageNumber="50">type</typeStatus>
series and a few other individuals of both sexes, all collected in a remote montane forest fragment in the Southern Highlands. The site probably represents the remnants of a wider forested landscape, interspersed with plateau grasslands and possibly naturally isolated from other Southern Highland forest blocks. For this reason the forests are of great bio- logical value and now the focus of further exploration and conservation intervention. During the last decade the Southern Highlands have been the subject of extensive biological investigation by the Wildlife Conservation Society. However, this species has not been detected in any other areas. It is therefore probable that
<taxonomicName id="4C1E4D101A417C5892E3919636438BAA" box="[1170,1302,908,932]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C5892E3919636438BAA" box="[1170,1302,908,932]" italics="true" pageId="7" pageNumber="50">A. matildae</emphasis>
</taxonomicName>
is a rangerestricted forest species, now relying on just a few forest fragments.
<taxonomicName id="4C1E4D101A417C5895FC91AA37588BC6" box="[909,1037,944,968]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C5895FC91AA37588BC6" box="[909,1037,944,968]" italics="true" pageId="7" pageNumber="50">A. matildae</emphasis>
</taxonomicName>
has an extent of occurrence smaller than
<quantity id="4CE69B761A417C5896BF91CD32718BE1" box="[206,292,983,1007]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="50" unit="km" value="100.0">100 km</quantity>
2 with extent of occurrence, area of occupancy and quality of habitat in continuing decline. According to IUCN guidelines (IUCN 2010) therefore, we propose to list
<taxonomicName id="4C1E4D101A417C5895EC91E137708C1D" box="[925,1061,1019,1043]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C5895EC91E137708C1D" box="[925,1061,1019,1043]" italics="true" pageId="7" pageNumber="50">A. matildae</emphasis>
</taxonomicName>
as ‘Critically Endangered’ CR B1b(i,ii,iii). Further investigations are being carried out in order to collect more information on this magnificent snake, and a small breeding programme has been established (see www.atherismatildae.org).
</paragraph>
</subSubSection>
<subSubSection id="C30465181A417C5896B6967D30E28CAC" pageId="7" pageNumber="50" type="etymology">
<paragraph id="8BA136931A417C5896B6967D30E28CAC" blockId="7.[151,1437,151,1187]" pageId="7" pageNumber="50">
<emphasis id="B96AEA811A417C5896B6967D321B8C71" bold="true" box="[199,334,1127,1151]" pageId="7" pageNumber="50">Etymology.</emphasis>
<taxonomicName id="4C1E4D101A417C589725967D315A8C71" box="[340,527,1127,1151]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="50" phylum="Chordata" rank="species" species="matildae">
<emphasis id="B96AEA811A417C589725967D315A8C71" box="[340,527,1127,1151]" italics="true" pageId="7" pageNumber="50">Atheris matildae</emphasis>
</taxonomicName>
is named for TRBD's daughter Matilda Davenport, one of the next generation of herpetologists. We suggest the common name 'Matilda's Horned Viper'.
</paragraph>
</subSubSection>
</treatment>
</document>
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