400 lines
56 KiB
XML
400 lines
56 KiB
XML
<document id="EB646095B92E7FDA11B6A04BB2D7EED7" ID-CLB-Dataset="28158" ID-DOI="10.11646/zootaxa.4563.1.1" ID-GBIF-Dataset="3e3068c1-c158-4d97-9aec-31fc80551edb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2600858" ID-ZooBank="A44D4C11-ADA1-48A8-9F00-B46BDF7B2A49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553170725873" checkinUser="plazi" docAuthor="Seeman, Owen D." docDate="2019" docId="03B63A508D5D015BFF60108203A44625" docLanguage="en" docName="zootaxa.4563.1.1.pdf" docOrigin="Zootaxa 4563 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megisthanus zachariei Seeman 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FF8F42288D7D017FFFF7166307564009" masterDocTitle="The megisthanid mites (Mesostigmata: Megisthanidae) of Australia" masterLastPageNumber="40" masterPageNumber="1" pageNumber="33" updateTime="1698702915705" updateUser="ExternalLinkService">
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<mods:titleInfo id="223E015027F793A612DF65B76ED6DE45">
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<mods:title id="07892816D363D7EC9A4F5E92AA0C6BEC">The megisthanid mites (Mesostigmata: Megisthanidae) of Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="65775EC911D4594AE995A1056F5C348B" type="personal">
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<mods:role id="58B9ADF8B696082852960B0857A56A24">
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<mods:roleTerm id="E6CCE8BF036E1046226C7435DE73F9CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0034920DA9DED1BB36C455500C9AE013">Seeman, Owen D.</mods:namePart>
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<mods:typeOfResource id="386339336E1EA6BADC77682ACB6390E6">text</mods:typeOfResource>
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<mods:title id="7CBA5F82AC6EDAA0587DD299DD16DEB2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E453D0CC09B1D76A0FB9AD4FC3D7BCED">
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<mods:date id="2D5F26CD3ABDABB0AAA076BDC2B1D53B">2019</mods:date>
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<mods:detail id="FBDB0444D7AF45FE21E4CF25B8D5E040" type="pubDate">
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<mods:number id="1C5DC0F76B1B66AB31BFD70F06D7B655">2019-02-28</mods:number>
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<mods:detail id="93944856E2B58D11E3060E06C0BA1CDF" type="volume">
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<mods:number id="02BE57193486AD8776689A753DEA2144">4563</mods:number>
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<mods:detail id="013405C28DC415E1EA88A4AA3605D04E" type="issue">
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<mods:number id="267BA8140DEF2A63E50AED4ADA8A6118">1</mods:number>
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<mods:start id="B5064C552F3537A04B575EA8B08214AE">1</mods:start>
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<mods:end id="4C091EEAB841356194B399C4BDB310B5">40</mods:end>
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</mods:part>
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<mods:classification id="053BBE0A48B4FCF8C067E458438517F5">journal article</mods:classification>
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<mods:identifier id="AA8011DF1EFC2C2BEA2A9DEAFDEA7207" type="CLB-Dataset">28158</mods:identifier>
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<mods:identifier id="85D006AC7E674AA10C82F6637EBC8774" type="DOI">10.11646/zootaxa.4563.1.1</mods:identifier>
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<mods:identifier id="7F77D000ADDF7D65507D88982D8F802F" type="GBIF-Dataset">3e3068c1-c158-4d97-9aec-31fc80551edb</mods:identifier>
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<mods:identifier id="5D3232F7B3A5BB70E5ACF344B22BF71A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DABF33EC2FCE3B68BBB1F0F64D1D8BF6" type="Zenodo-Dep">2600858</mods:identifier>
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<mods:identifier id="FFDBEE437469E70195D63DCAF2ED354A" type="ZooBank">A44D4C11-ADA1-48A8-9F00-B46BDF7B2A49</mods:identifier>
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</mods:mods>
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<treatment id="03B63A508D5D015BFF60108203A44625" ID-DOI="http://doi.org/10.5281/zenodo.5924277" ID-GBIF-Taxon="154474404" ID-Zenodo-Dep="5924277" LSID="urn:lsid:plazi:treatment:03B63A508D5D015BFF60108203A44625" httpUri="http://treatment.plazi.org/id/03B63A508D5D015BFF60108203A44625" lastPageId="36" lastPageNumber="37" pageId="32" pageNumber="33">
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<subSubSection id="C305D8CD8D5D015FFF601082055846F2" box="[151,526,1761,1787]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BA08B468D5D015FFF601082055846F2" blockId="32.[151,526,1761,1821]" box="[151,526,1761,1787]" pageId="32" pageNumber="33">
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<heading id="D0E83C2A8D5D015FFF601082055846F2" bold="true" box="[151,526,1761,1787]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
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<emphasis id="B96B57548D5D015FFF601082055846F2" bold="true" box="[151,526,1761,1787]" pageId="32" pageNumber="33">
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<taxonomicName id="4C1FF0C58D5D015FFF60108206FF46F2" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[151,425,1761,1787]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="zachariei" status="sp. nov.">
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<emphasis id="B96B57548D5D015FFF60108206FF46F2" bold="true" box="[151,425,1761,1787]" italics="true" pageId="32" pageNumber="33">Megisthanus zachariei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A258EA2F8D5D015FFE471082055846F2" box="[432,526,1761,1787]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C305D8CD8D5D015FFF601167066E4715" box="[151,312,1796,1821]" pageId="32" pageNumber="33" type="description">
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||
<paragraph id="8BA08B468D5D015FFF601167066E4715" blockId="32.[151,526,1761,1821]" box="[151,312,1796,1821]" pageId="32" pageNumber="33">
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<figureCitation id="132497C38D5D015FFF601167066E4715" box="[151,312,1796,1821]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="33.[151,250,1957,1979]" captionStartId-1="34.[151,250,1933,1955]" captionStartId-2="35.[151,250,1023,1045]" captionTargetBox-0="[170,1410,633,1925]" captionTargetBox-1="[199,1380,210,1889]" captionTargetBox-2="[176,1416,226,970]" captionTargetId-0="figure@33.[161,1425,625,1936]" captionTargetId-1="figure@34.[186,1410,193,1915]" captionTargetId-2="figure@35.[161,1426,193,1001]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionText-0="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." captionText-1="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." captionText-2="FIGURE 23. Megisthanus zachariei sp. nov. (A) female anterior subcapitulum; (B) female palp, trochanter–genu; (C) major male anterior subcapitulum; (D) major male palp, trochanter–genu; (E) palp trochantal spur, aspect from right palp. Arrows indicate spurs." httpUri-0="https://zenodo.org/record/2600900/files/figure.png" httpUri-1="https://zenodo.org/record/2600902/files/figure.png" httpUri-2="https://zenodo.org/record/2600904/files/figure.png" pageId="32" pageNumber="33">Figures 21–23</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C305D8CD8D5D015FFF60112F027F47A5" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BA08B468D5D015FFF60112F027F47A5" blockId="32.[151,1437,1868,2037]" pageId="32" pageNumber="33">
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<emphasis id="B96B57548D5D015FFF60112F05E8476C" bold="true" box="[151,702,1868,1893]" pageId="32" pageNumber="33">
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Material examined.
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<materialsCitation id="3B77811B8D5D015FFE57112F05E8476C" ID-GBIF-Occurrence="2013950918" box="[416,702,1868,1893]" country="Australia" location="Queensland" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Queensland">
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<collectingCountry id="F308CBD68D5D015FFE57112F0544476C" box="[416,530,1868,1893]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
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,
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<collectingRegion id="49DB45A48D5D015FFDDD112F05EF476C" box="[554,697,1868,1893]" country="Australia" name="Queensland" pageId="32" pageNumber="33">Queensland</collectingRegion>
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,
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</materialsCitation>
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</emphasis>
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ex
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<taxonomicName id="4C1FF0C58D5D015FFCF7112F03E0476D" authority="Macleay, 1871" authorityName="Macleay" authorityYear="1871" box="[768,1206,1868,1893]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kaupii">
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<emphasis id="B96B57548D5D015FFCF7112F04B8476D" box="[768,1006,1868,1892]" italics="true" pageId="32" pageNumber="33">Aulacocyclus kaupii</emphasis>
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<bibRefCitation id="EF8EF6B78D5D015FFBF5112F03E0476D" author="Macleay, W. J." box="[1026,1206,1868,1893]" pageId="32" pageNumber="33" pagination="159 - 205" refId="ref22370" refString="Macleay, W. J. (1871) Notes on a collection of insects from Gayndah (continued). Transactions of the Entomological Society of New South Wales, 2, 159 - 205." type="journal article" year="1871">Macleay, 1871</bibRefCitation>
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</taxonomicName>
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.
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<materialsCitation id="3B77811B8D5D015FFB3A112F02CF4781" ID-GBIF-Occurrence="2013950912" collectingDate="2012-04-18" collectionCode="QMS" collectorName="O. Seeman & H. Urbina & Bartlett" country="Australia" latitude="-27.966667" location="Goomburra State Forest" longLatPrecision="1234" longitude="152.31667" pageId="32" pageNumber="33" specimenCode="QMS 109500" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<typeStatus id="54A435E48D5D015FFB3A112F0263476C" box="[1229,1333,1868,1893]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
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female,
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||
<location id="8EC0DD9D8D5D015FFF60110C06E4478E" LSID="urn:lsid:plazi:treatment:03B63A508D5D015BFF60108203A44625:8EC0DD9D8D5D015FFF60110C06E4478E" box="[151,434,1903,1928]" country="Australia" latitude="-27.966667" longLatPrecision="1234" longitude="152.31667" name="Goomburra State Forest" pageId="32" pageNumber="33" stateProvince="Queensland">Goomburra State Forest</location>
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,
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<geoCoordinate id="EE2BED818D5D015FFE35110C054D4781" box="[450,539,1903,1928]" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-27.966667">27°58′S</geoCoordinate>
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||
<geoCoordinate id="EE2BED818D5D015FFDDF110C05C54781" box="[552,659,1903,1928]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="152.31667">152°19′E</geoCoordinate>
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,
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||
<date id="FFA1AD868D5D015FFD53110C044A4781" box="[676,796,1903,1928]" pageId="32" pageNumber="33" value="2012-04-18">
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||
<collectingDate id="EFE5546E8D5D015FFD53110C044A4781" box="[676,796,1903,1928]" pageId="32" pageNumber="33" value="2012-04-18">18.iv.2012</collectingDate>
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</date>
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,
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<collectorName id="26EAEE908D5D015FFCDA111304E74781" box="[813,945,1904,1928]" pageId="32" pageNumber="33">O. Seeman</collectorName>
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,
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<collectorName id="26EAEE908D5D015FFC36111303614781" box="[961,1079,1903,1928]" pageId="32" pageNumber="33">H. Urbina</collectorName>
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& J.
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<collectorName id="26EAEE908D5D015FFB72110C038F4781" box="[1157,1241,1903,1928]" pageId="32" pageNumber="33">Bartlett</collectorName>
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(
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<specimenCode id="DBB9233D8D5D015FFB1A111302C74781" box="[1261,1425,1903,1928]" collectionCode="QMS" pageId="32" pageNumber="33">QMS 109500</specimenCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B77811B8D5D015FFF6011F6040347A4" ID-GBIF-Occurrence="2013950917" box="[151,853,1940,1965]" collectingDate="2012-04-18" collectionCode="QMS" collectorName="O. Seeman & H. Urbina & Bartlett" country="Australia" latitude="-27.966667" location="Goomburra State Forest" longLatPrecision="1234" longitude="152.31667" pageId="32" pageNumber="33" specimenCode="QMS 109501" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
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<typeStatus id="54A435E48D5D015FFF6011F607A847A4" box="[151,254,1941,1965]" pageId="32" pageNumber="33" type="paratype">Paratype</typeStatus>
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: 1 major male, same data as holotype (
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<specimenCode id="DBB9233D8D5D015FFD5811F7041A47A4" box="[687,844,1940,1965]" collectionCode="QMS" pageId="32" pageNumber="33">QMS 109501</specimenCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B77811B8D5D015FFCA811F7027047A5" ID-GBIF-Occurrence="2013950903" box="[863,1318,1940,1965]" collectionCode="QM" country="Australia" pageId="32" pageNumber="33" specimenCount="2" stateProvince="Queensland" typeStatus="holotype">
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<typeStatus id="54A435E48D5D015FFCA811F7049147A4" box="[863,967,1940,1965]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
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and
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<typeStatus id="54A435E48D5D015FFC0911F6030847A4" box="[1022,1118,1941,1965]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
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deposited in
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<collectionCode id="ED0E13838D5D015FFB0211F7027047A5" box="[1269,1318,1940,1964]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="32" pageNumber="33" type="Museum">QM</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C305D8CD8D5D015EFF3011DB0405416C" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="diagnosis">
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<paragraph id="8BA08B468D5D015EFF3011DB0405416C" blockId="32.[151,1437,1868,2037]" lastBlockId="33.[151,1437,151,573]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<emphasis id="B96B57548D5D015FFF3011DB061747D8" bold="true" box="[199,321,1976,2001]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
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<emphasis id="B96B57548D5D015FFEBF11DB055C47D9" box="[328,522,1976,2000]" italics="true" pageId="32" pageNumber="33">Female and male</emphasis>
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. Sternal setae st4 absent. Seta pd2 of femur IV on rudimentary projection. Dorsal shield finely reticulate. Ventrianal shield widest at mid-level of anus.
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<emphasis id="B96B57548D5D015FFC6911BF04A447FD" box="[926,1010,2012,2036]" italics="true" pageId="32" pageNumber="33">Female</emphasis>
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||
. Dorsal shield ovate, length
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<quantity id="4CE726A38D5D015FFAC511BF02C147FD" box="[1330,1431,2012,2036]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="32" pageNumber="33" unit="mm" value="2.2">2.20 mm</quantity>
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||
, maximum width
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||
<quantity id="4CE726A38D5C015EFEA016F406EB40A6" box="[343,445,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="34" unit="mm" value="1.15">1.15 mm</quantity>
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, not including secondary sclerotisation. Dorsal setae long (160–380), smooth anteriorly, sparsely barbed posteriorly; podonotum more setose than opisthonotum. Sternogynal shields separate, each bearing 3–4 setae; internal genitalia with rudimentary latigynal element, small mesogynal element. Ventrianal shield weakly reniform.
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||
<emphasis id="B96B57548D5C015EFE93176606BC4115" box="[356,490,260,284]" italics="true" pageId="33" pageNumber="34">Major male</emphasis>
|
||
. Larger than female, dorsal shield length
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||
<quantity id="4CE726A38D5C015EFC261767036C4115" box="[977,1082,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="33" pageNumber="34" unit="mm" value="2.45">2.45 mm</quantity>
|
||
, width
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||
<quantity id="4CE726A38D5C015EFB65176703AA4115" box="[1170,1276,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="33" pageNumber="34" unit="mm" value="1.7">1.70 mm</quantity>
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; setae on soft cuticle laterad dorsal shield short to moderate length, 40–100; 25 setae around genital opening; corniculi with small medial inner tooth; setae h1–3 smooth, palp coxal seta barbed.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D5C015BFF30170C04D744A9" lastPageId="36" lastPageNumber="37" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="8BA08B468D5C015EFF30170C06CF4234" blockId="33.[151,1437,151,573]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96B57548D5C015EFF30170C06E04181" bold="true" box="[199,438,367,392]" pageId="33" pageNumber="34">
|
||
Description.
|
||
<emphasis id="B96B57548D5C015EFEA8170C06E04181" box="[351,438,367,392]" italics="true" pageId="33" pageNumber="34">Female</emphasis>
|
||
</emphasis>
|
||
(n = 1). Lightly sclerotised specimen. Dorsum (
|
||
<figureCitation id="132497C38D5C015EFC20170C03324181" box="[983,1124,367,392]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="33" pageNumber="34">Figs 21A, C</figureCitation>
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,
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<figureCitation id="132497C38D5C015EFB87171303F24181" box="[1136,1188,368,392]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="33" pageNumber="34">22A</figureCitation>
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). Idiosomal length ca.
|
||
<quantity id="4CE726A38D5C015EFF6017F7065641A5" box="[151,256,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="33" pageNumber="34" unit="mm" value="2.35">2.35 mm</quantity>
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||
, width
|
||
<quantity id="4CE726A38D5C015EFEAF17F7069441A5" box="[344,450,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="33" pageNumber="34" unit="mm" value="1.6">1.60 mm</quantity>
|
||
, body length (including
|
||
<taxonomicName id="4C1FF0C58D5C015EFD1717F7043C41A4" authorityName="Krefft" authorityYear="1871" box="[736,874,404,429]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="33" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
, to tip of gnathotectum)
|
||
<quantity id="4CE726A38D5C015EFB7A17F703A141A5" box="[1165,1271,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="33" pageNumber="34" unit="mm" value="3.05">3.05 mm</quantity>
|
||
. Dorsal shield ovate, length
|
||
<quantity id="4CE726A38D5C015EFECF17DB06F541C6" box="[312,419,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="33" pageNumber="34" unit="mm" value="2.2">2.20 mm</quantity>
|
||
, width
|
||
<quantity id="4CE726A38D5C015EFE0817D4053D41C6" box="[511,619,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="34" unit="mm" value="1.15">1.15 mm</quantity>
|
||
; secondary sclerotisation absent. Dorsal shield hypertrichous, densely setose anteromedially; all setae long (160–380), longest laterally, setae smooth anteriorly and sparsely barbed posteriorly. Dorsal shield cuticle finely reticulated. Setae in soft cuticle barbed, on small platelets, length 180–550, longer posterolaterally.
|
||
</paragraph>
|
||
<caption id="DF60DBCE8D5C015EFF6011C603D847D3" httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1957,1979]" targetBox="[170,1410,633,1925]" targetPageId="33">
|
||
<paragraph id="8BA08B468D5C015EFF6011C603D847D3" blockId="33.[151,1436,1957,2010]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96B57548D5C015EFF6011C6067247B3" bold="true" box="[151,292,1957,1979]" pageId="33" pageNumber="34">FIGURE 21.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D5C015EFED911C6054E47B2" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[302,536,1957,1979]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zachariei" status="sp. nov.">
|
||
<emphasis id="B96B57548D5C015EFED911C6054E47B2" box="[302,536,1957,1979]" italics="true" pageId="33" pageNumber="34">Megisthanus zachariei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D5C015EFDD511C5052147B2" bold="true" box="[546,631,1958,1979]" pageId="33" pageNumber="34">
|
||
<taxonomicNameLabel id="A258EA2F8D5C015EFDD511C5052147B2" box="[546,631,1958,1979]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D)
|
||
<taxonomicName id="4C1FF0C58D5C015EFE5811A706BC47D3" box="[431,490,1988,2010]" pageId="33" pageNumber="34" rank="species" species="major">major</taxonomicName>
|
||
male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF60DBCE8D5F015DFF6011EE048147E9" httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1933,1955]" targetBox="[199,1380,210,1889]" targetPageId="34">
|
||
<paragraph id="8BA08B468D5F015DFF6011EE048147E9" blockId="34.[151,1436,1933,2016]" pageId="34" pageNumber="35">
|
||
<emphasis id="B96B57548D5F015DFF6011EE067647AB" bold="true" box="[151,288,1933,1955]" pageId="34" pageNumber="35">FIGURE 22.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D5F015DFED111EE055D47AA" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[294,523,1933,1955]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="zachariei" status="sp. nov.">
|
||
<emphasis id="B96B57548D5F015DFED111EE055D47AA" box="[294,523,1933,1955]" italics="true" pageId="34" pageNumber="35">Megisthanus zachariei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EA2F8D5F015DFDE411ED053547AA" box="[531,611,1934,1955]" pageId="34" pageNumber="35" rank="species">
|
||
<emphasis id="B96B57548D5F015DFDE411ED053547AA" bold="true" box="[531,611,1934,1955]" pageId="34" pageNumber="35">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E)
|
||
<taxonomicName id="4C1FF0C58D5F015DFDF811CF051C47CB" box="[527,586,1964,1986]" pageId="34" pageNumber="35" rank="species" species="major">major</taxonomicName>
|
||
male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF60DBCE8D5E015CFF60159C067C445B" httpUri="https://zenodo.org/record/2600904/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1023,1045]" targetBox="[176,1416,226,970]" targetPageId="35">
|
||
<paragraph id="8BA08B468D5E015CFF60159C067C445B" blockId="35.[151,1436,1023,1106]" pageId="35" pageNumber="36">
|
||
<emphasis id="B96B57548D5E015CFF60159C0677441D" bold="true" box="[151,289,1023,1045]" pageId="35" pageNumber="36">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D5E015CFEDF159C0559441C" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[296,527,1023,1045]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zachariei" status="sp. nov.">
|
||
<emphasis id="B96B57548D5E015CFEDF159C0559441C" box="[296,527,1023,1045]" italics="true" pageId="35" pageNumber="36">Megisthanus zachariei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D5E015CFDE0159C053F441D" bold="true" box="[535,617,1023,1044]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="A258EA2F8D5E015CFDE0159C053F441D" box="[535,617,1023,1044]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A) female anterior subcapitulum; (B) female palp, trochanter–genu; (C) major male anterior subcapitulum; (D) major male palp, trochanter–genu; (E) palp trochantal spur, aspect from right palp. Arrows indicate spurs.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA08B468D5E015CFF3012E3050C44D6" blockId="35.[151,1437,1150,2004]" pageId="35" pageNumber="36">
|
||
<emphasis id="B96B57548D5E015CFF3012E3065A449E" box="[199,268,1152,1175]" italics="true" pageId="35" pageNumber="36">Venter</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D5E015CFEE9121D06DF449E" box="[286,393,1150,1175]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="35" pageNumber="36">Figs 21B</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D5E015CFE61121C06E5449E" box="[406,435,1151,1175]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="35" pageNumber="36">22</figureCitation>
|
||
B–C). Setae st1 135, sparsely barbed. Sternal shield (
|
||
<figureCitation id="132497C38D5E015CFBFA121D0324449E" box="[1037,1138,1150,1175]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="35" pageNumber="36">Fig. 22B</figureCitation>
|
||
) reticulate, with two pairs of setae, 3–4 pairs of pores opening ventrally and one pair of lyrifissures; setae st2–3 sparsely barbed, st2 100, st3 80, st4 absent; shield medial length 100.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5E015CFF301288032D459D" blockId="35.[151,1437,1150,2004]" pageId="35" pageNumber="36">
|
||
Sternogynal shields (
|
||
<figureCitation id="132497C38D5E015CFE4512880540450A" box="[434,534,1259,1284]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="35" pageNumber="36">Fig. 22B</figureCitation>
|
||
) free posteriorly, genital opening length 305, width 340, each sternogynal shield length 270, lineate-reticulate, each bearing 3–4 smooth setae and one lyrifissure; anterior setae (35) slightly thicker and longer than posterior setae (20–30). Internal genitalia comprises three obvious elements (
|
||
<emphasis id="B96B57548D5E015CFB14135702534542" box="[1251,1285,1332,1355]" italics="true" pageId="35" pageNumber="36">mg</emphasis>
|
||
,
|
||
<emphasis id="B96B57548D5E015CFAE0135702674542" box="[1303,1329,1332,1355]" italics="true" pageId="35" pageNumber="36">va</emphasis>
|
||
,
|
||
<emphasis id="B96B57548D5E015CFAB3135002094542" box="[1348,1375,1331,1355]" italics="true" pageId="35" pageNumber="36">vh</emphasis>
|
||
) and rudimentary latigynal (
|
||
<emphasis id="B96B57548D5E015CFE57133406E04566" box="[416,438,1367,1391]" italics="true" pageId="35" pageNumber="36">lg</emphasis>
|
||
) elements (
|
||
<figureCitation id="132497C38D5E015CFDB7133505FD4566" box="[576,683,1366,1391]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="35" pageNumber="36">Fig. 22C</figureCitation>
|
||
); rudimentary latigynal elements fused with thickened margin of sternogynal shield, with small porose area near attachment of
|
||
<emphasis id="B96B57548D5E015CFCBB131F0430459A" box="[844,870,1404,1427]" italics="true" pageId="35" pageNumber="36">va</emphasis>
|
||
with sternogynal shield.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5E015CFF3013FD04F7462D" blockId="35.[151,1437,1150,2004]" pageId="35" pageNumber="36">
|
||
Ventral shield (
|
||
<figureCitation id="132497C38D5E015CFE8213FD068C45BE" box="[373,474,1438,1463]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="35" pageNumber="36">Fig. 21B</figureCitation>
|
||
) extends 880 posteriorly from posterior margin of genital opening; maximum width behind coxa IV 420; shield with 12–13 pairs of setae around shield margin, 1 pair posteriorly within shield (
|
||
<typeStatus id="54A435E48D5E015CFF691385065245F6" box="[158,260,1510,1535]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
, just within margin), and with 1–2 pairs of setae off shield margin close to sternogynal shield; setae smooth, length 45–90; shield rugose due to incomplete sclerotisation.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5E015CFF30104D022A4665" blockId="35.[151,1437,1150,2004]" pageId="35" pageNumber="36">
|
||
Ventrianal shield slightly reniform (
|
||
<figureCitation id="132497C38D5E015CFDAD104D0596464E" box="[602,704,1582,1607]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="35" pageNumber="36">Fig. 21B</figureCitation>
|
||
), with anus in centre of shield; anterior width ca. 440, maximum width at mid-level of anus, 620, length 380; shield reticulate; 14–16 loosely paired, smooth setae, length 55–80.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5E015CFF301015056646DE" blockId="35.[151,1437,1150,2004]" pageId="35" pageNumber="36">
|
||
Peritrematal shield hypertrichous, post-peritrematal region extends 530 behind coxa IV into metapodal region (
|
||
<figureCitation id="132497C38D5E015CFF6810F8065046BA" box="[159,262,1691,1716]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="35" pageNumber="36">Fig. 21B</figureCitation>
|
||
); setae laterad legs smooth to sparsely barbed, length 30–55; three pairs of long setae on anterior margin of peritrematal shield, length to 250.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5E015CFF301080063947DD" blockId="35.[151,1437,1150,2004]" pageId="35" pageNumber="36">
|
||
<taxonomicName id="4C1FF0C58D5E015CFF301080060046F2" authorityName="Krefft" authorityYear="1871" box="[199,342,1763,1787]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="35" pageNumber="36" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B96B57548D5E015CFF301080060046F2" box="[199,342,1763,1787]" italics="true" pageId="35" pageNumber="36">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132497C38D5E015CFE91108006EB46F2" box="[358,445,1763,1788]" captionStart="FIGURE 23" captionStartId="35.[151,250,1023,1045]" captionTargetBox="[176,1416,226,970]" captionTargetId="figure@35.[161,1426,193,1001]" captionTargetPageId="35" captionText="FIGURE 23. Megisthanus zachariei sp. nov. (A) female anterior subcapitulum; (B) female palp, trochanter–genu; (C) major male anterior subcapitulum; (D) major male palp, trochanter–genu; (E) palp trochantal spur, aspect from right palp. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600904/files/figure.png" pageId="35" pageNumber="36">Figs 23</figureCitation>
|
||
A–B). Gnathotectum reticulate, medial length 690. Subcapitulum with setae h1 260, h2 145 (detached), h3 240 (detached), pc 185, all setae sparsely barbed; subcapitulum poorly sclerotised, with some fine striae, subcapitular gutter with five folds where rows of denticles should be. Corniculi length 250 with medial tooth. Palp (
|
||
<figureCitation id="132497C38D5E015CFEDB112D06C1476E" box="[300,407,1870,1895]" captionStart="FIGURE 23" captionStartId="35.[151,250,1023,1045]" captionTargetBox="[176,1416,226,970]" captionTargetId="figure@35.[161,1426,193,1001]" captionTargetPageId="35" captionText="FIGURE 23. Megisthanus zachariei sp. nov. (A) female anterior subcapitulum; (B) female palp, trochanter–genu; (C) major male anterior subcapitulum; (D) major male palp, trochanter–genu; (E) palp trochantal spur, aspect from right palp. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600904/files/figure.png" pageId="35" pageNumber="36">Fig. 23B</figureCitation>
|
||
) trochanter with ventro-distal spur, length 30, palp femur with distal spur, length 15. Chelicerae with entire fixed digit length 660, movable digit length 380. Morphology as per genus description. Fixed digit with 13 teeth; cheliceral seta length 30. Movable digit with 10 teeth; excrescences not apparent due to poor sclerotisation.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF3016FA03664149" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF3016FA07AA40B9" box="[199,252,153,176]" italics="true" pageId="36" pageNumber="37">Legs</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D59015BFEE716F4062940B9" box="[272,383,151,176]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="36" pageNumber="37">Fig. 22D</figureCitation>
|
||
). Setation as per genus description. Selected setal measurements: genu II setae ad1, pd1 subequal, length 90–110; seta pd1 on genu III long (length 525), much longer than ad1 (length 135), tarsus III ad3 moderately long, length 250, femur IV pd1 long, length 420, genu IV ad1 very long, length ca. 500, pd1 ca. 500, tibia IV pd1 long, length ca. 500 and tarsus IV ad4 long, length ca. 350, pd3 long, length ca. 300. Femur IV with rudimentary projections bearing setae pd2, ad2, larger projection bearing seta ad3.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF30172803B84211" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF3017280655416D" bold="true" box="[199,259,331,356]" italics="true" pageId="36" pageNumber="37">Male</emphasis>
|
||
(major) (n = 1). Lightly sclerotised specimen.
|
||
<emphasis id="B96B57548D59015BFCD3172E0428416D" box="[804,894,333,356]" italics="true" pageId="36" pageNumber="37">Dorsum</emphasis>
|
||
. Idiosomal length ca.
|
||
<quantity id="4CE726A38D59015BFB76172F03BC416D" box="[1153,1258,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="37" unit="mm" value="2.5">2.50 mm</quantity>
|
||
, width ca.
|
||
<quantity id="4CE726A38D59015BFA9C172F0793418E" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="36" pageNumber="37" unit="mm" value="1.7">1.70 mm</quantity>
|
||
, body length (including
|
||
<taxonomicName id="4C1FF0C58D59015BFE16170C053D4181" authorityName="Krefft" authorityYear="1871" box="[481,619,367,392]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="36" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
, to tip of gnathotectum) ca.
|
||
<quantity id="4CE726A38D59015BFC41170C0376418E" box="[950,1056,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" pageId="36" pageNumber="37" unit="mm" value="3.35">3.35 mm</quantity>
|
||
. Dorsal shield ovate, length
|
||
<quantity id="4CE726A38D59015BFA9C170C079341A5" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="36" pageNumber="37" unit="mm" value="2.45">2.45 mm</quantity>
|
||
, width
|
||
<quantity id="4CE726A38D59015BFEE817F706DC41A5" box="[287,394,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="36" pageNumber="37" unit="mm" value="1.45">1.45 mm</quantity>
|
||
; secondary sclerotisation absent. Dorsum hypertrichous. Most setae on dorsal shield tiny, smooth, length <5; some marginal setae longer (30–80), especially anteriorly and posteriorly. Setae in soft membrane close to dorsal shield sparsely barbed, short to moderate length (40–100), becoming more barbed and longer laterally; platelets and pores in soft cuticle not apparent, probably due to poor sclerotisation.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF301446039D431C" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF301446065A4235" box="[199,268,549,572]" italics="true" pageId="36" pageNumber="37">Venter</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D59015BFEE9144706DC4235" box="[286,394,548,573]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="36" pageNumber="37">Figs 21D</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D59015BFE601447069F4235" box="[407,457,548,572]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="36" pageNumber="37">22E</figureCitation>
|
||
). Setae st1 barbed, length 170. Sternoventral shield reticulate (poorly expressed due to weak sclerotisation), length from posterior margin of genital opening 940. Sternal setae st2 and st3 barbed, length 125–135; sternoventral shield (
|
||
<figureCitation id="132497C38D59015BFE05140F0503428D" box="[498,597,620,645]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="36" pageNumber="37">Fig. 22E</figureCitation>
|
||
) hypertrichous posterior to setae st3, with 25 setae around genital opening; posterior sternoventral shield with setae along margin; these setae smooth to sparsely barbed, shortest setae 80 (near genital opening) and longest setae 145 (posterior to coxa IV); posterior sternoventral shield (
|
||
<figureCitation id="132497C38D59015BFB0414D7020F42C5" box="[1267,1369,692,717]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="36" pageNumber="37">Fig. 21D</figureCitation>
|
||
) with two large discs of porose cuticle (diameter 90), each bearing an anterior (55) and posterior pair (35) of unmodified smooth setae, and a highly modified pair of setae centrally, shrivelled due to poor sclerotisation.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF30157C04664354" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
Genital opening (
|
||
<figureCitation id="132497C38D59015BFE7D157C06B84331" box="[394,494,799,824]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="36" pageNumber="37">Fig. 22E</figureCitation>
|
||
) length 135, width 115, surrounded by a ring of sclerotised cuticle; this cuticle with 12–15 pairs of loosely paired pores laterad genital opening.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF301504065E43C1" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
Ventrianal shield (
|
||
<figureCitation id="132497C38D59015BFE6C150405504389" box="[411,518,871,896]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="36" pageNumber="37">Fig. 21D</figureCitation>
|
||
) a curved trapezoid, length 355, anterior width ca. 525, maximum width 600, widest at mid-level of anus; shield reticulate, with 13 loosely paired setae. Peritreme and peritrematal shield similar to female.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF3015B704804474" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
<taxonomicName id="4C1FF0C58D59015BFF3015B7060043E5" authorityName="Krefft" authorityYear="1871" box="[199,342,980,1004]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="36" pageNumber="37" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B96B57548D59015BFF3015B7060043E5" box="[199,342,980,1004]" italics="true" pageId="36" pageNumber="37">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132497C38D59015BFE9115B706EA43E5" box="[358,444,980,1005]" captionStart="FIGURE 23" captionStartId="35.[151,250,1023,1045]" captionTargetBox="[176,1416,226,970]" captionTargetId="figure@35.[161,1426,193,1001]" captionTargetPageId="35" captionText="FIGURE 23. Megisthanus zachariei sp. nov. (A) female anterior subcapitulum; (B) female palp, trochanter–genu; (C) major male anterior subcapitulum; (D) major male palp, trochanter–genu; (E) palp trochantal spur, aspect from right palp. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600904/files/figure.png" pageId="36" pageNumber="37">Figs 23</figureCitation>
|
||
C–E). Gnathotectum similar to female, medial length 840; subcapitulum similar, except with submedial projections (inner length 40). Setae h1 smooth, blade-like, 200; setae h2 smooth, 180, setae h3 smooth, 180, palp coxal seta barbed, 180. Corniculi with small medial tooth-like process, corniculi length 195 (similar to female in length). Palp (
|
||
<figureCitation id="132497C38D59015BFDDD125C05D64451" box="[554,640,1087,1112]" captionStart="FIGURE 23" captionStartId="35.[151,250,1023,1045]" captionTargetBox="[176,1416,226,970]" captionTargetId="figure@35.[161,1426,193,1001]" captionTargetPageId="35" captionText="FIGURE 23. Megisthanus zachariei sp. nov. (A) female anterior subcapitulum; (B) female palp, trochanter–genu; (C) major male anterior subcapitulum; (D) major male palp, trochanter–genu; (E) palp trochantal spur, aspect from right palp. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600904/files/figure.png" pageId="36" pageNumber="37">Figs 23</figureCitation>
|
||
D–E) trochanter with two distoventral cusps on trochanter (length 25, 35) and one large cusp on femur (length 70). Chelicerae similar to female.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF3012EA04D744A9" blockId="36.[151,1437,151,1581]" box="[199,897,1159,1184]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF3012EA07AA44A9" box="[199,252,1161,1184]" italics="true" pageId="36" pageNumber="37">Legs</emphasis>
|
||
. Similar to female, distal spurs and spines similar in size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D59015BFF3012CF039144CD" box="[199,1223,1196,1221]" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="8BA08B468D59015BFF3012CF039144CD" blockId="36.[151,1437,151,1581]" box="[199,1223,1196,1221]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF3012CF061844CC" bold="true" box="[199,334,1196,1221]" pageId="36" pageNumber="37">Etymology.</emphasis>
|
||
It is with great pleasure that I name this species for my nephew Zacharie West.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D59015BFF3012AC07A84539" pageId="36" pageNumber="37" type="diagnosis">
|
||
<paragraph id="8BA08B468D59015BFF3012AC07A84539" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF3012AC069B44E1" bold="true" box="[199,461,1231,1256]" pageId="36" pageNumber="37">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D59015BFE2312B3058444E1" authorityName="Seeman" authorityYear="2019" box="[468,722,1232,1256]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="zachariei">
|
||
<emphasis id="B96B57548D59015BFE2312B3058444E1" box="[468,722,1232,1256]" italics="true" pageId="36" pageNumber="37">Megisthanus zachariei</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C1FF0C58D59015BFC5512B2036344E1" authorityName="Seeman" authorityYear="2019" box="[930,1077,1232,1256]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="southcotti">
|
||
<emphasis id="B96B57548D59015BFC5512B204EF44E1" box="[930,953,1233,1256]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFC3012B3036344E1" box="[967,1077,1232,1256]" italics="true" pageId="36" pageNumber="37">southcotti</emphasis>
|
||
</taxonomicName>
|
||
and their diagnostic differences are explained under the latter species. The species is also similar to
|
||
<taxonomicName id="4C1FF0C58D59015BFC61129603724505" authorityName="Berlese" authorityYear="1910" box="[918,1060,1268,1292]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="modestus">
|
||
<emphasis id="B96B57548D59015BFC61129604FB4505" box="[918,941,1269,1292]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFC4B129703724505" box="[956,1060,1268,1292]" italics="true" pageId="36" pageNumber="37">modestus</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs by lacking setae st4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D59015BFF30135F03A44625" pageId="36" pageNumber="37" type="discussion">
|
||
<paragraph id="8BA08B468D59015BFF30135F03A44625" blockId="36.[151,1437,151,1581]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96B57548D59015BFF30135F066D455C" bold="true" box="[199,315,1340,1365]" pageId="36" pageNumber="37">Remarks.</emphasis>
|
||
Both specimens of
|
||
<taxonomicName id="4C1FF0C58D59015BFDEF135E05F3455D" authorityName="Seeman" authorityYear="2019" box="[536,677,1340,1364]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="zachariei">
|
||
<emphasis id="B96B57548D59015BFDEF135E0579455D" box="[536,559,1341,1364]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFDCA135F05F3455D" box="[573,677,1340,1364]" italics="true" pageId="36" pageNumber="37">zachariei</emphasis>
|
||
</taxonomicName>
|
||
are lightly sclerotised, so several aspects are difficult to see, but the setation of the dorsal and ventral shields distinguish the two species of
|
||
<taxonomicName id="4C1FF0C58D59015BFBE6130303F64571" authorityName="Thorell" authorityYear="1882" box="[1041,1184,1376,1400]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D59015BFBE6130303F64571" box="[1041,1184,1376,1400]" italics="true" pageId="36" pageNumber="37">Megisthanus</emphasis>
|
||
</taxonomicName>
|
||
found on the genus
|
||
<taxonomicName id="4C1FF0C58D59015BFF6013E7067A4595" authorityName="Kaup" authorityYear="1868" box="[151,300,1412,1436]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D59015BFF6013E7067A4595" box="[151,300,1412,1436]" italics="true" pageId="36" pageNumber="37">Aulacocyclus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D59015BFEC913E606804595" authorityName="Seeman" authorityYear="2019" box="[318,470,1412,1436]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="southcotti">
|
||
<emphasis id="B96B57548D59015BFEC913E606034595" box="[318,341,1413,1436]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFE9F13E706804595" box="[360,470,1412,1436]" italics="true" pageId="36" pageNumber="37">southcotti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D59015BFDE113E605FE4595" authorityName="Seeman" authorityYear="2019" box="[534,680,1412,1436]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="zachariei">
|
||
<emphasis id="B96B57548D59015BFDE113E6057B4595" box="[534,557,1413,1436]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFDB713E705FE4595" box="[576,680,1412,1436]" italics="true" pageId="36" pageNumber="37">zachariei</emphasis>
|
||
</taxonomicName>
|
||
. The host beetle of
|
||
<taxonomicName id="4C1FF0C58D59015BFC6A13E603794595" authorityName="Seeman" authorityYear="2019" box="[925,1071,1412,1436]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="zachariei">
|
||
<emphasis id="B96B57548D59015BFC6A13E604E24595" box="[925,948,1413,1436]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFC3013E703794595" box="[967,1071,1412,1436]" italics="true" pageId="36" pageNumber="37">zachariei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D59015BFBB613E703FD4595" authorityName="Macleay" authorityYear="1871" box="[1089,1195,1412,1436]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kaupii">
|
||
<emphasis id="B96B57548D59015BFBB613E703044592" box="[1089,1106,1412,1435]" italics="true" pageId="36" pageNumber="37">A</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D59015BFB9213E703FD4595" box="[1125,1195,1412,1436]" italics="true" pageId="36" pageNumber="37">kaupii</emphasis>
|
||
</taxonomicName>
|
||
, is very common in southeast
|
||
<collectingRegion id="49DB45A48D59015BFEFF13C406DA45C9" box="[264,396,1447,1472]" country="Australia" name="Queensland" pageId="36" pageNumber="37">Queensland</collectingRegion>
|
||
where it tends to be found on the verges of rainforests and into drier habitats where other genera, such as
|
||
<taxonomicName id="4C1FF0C58D59015BFEBB13AF068245ED" authorityName="Kaup" authorityYear="1868" box="[332,468,1484,1508]" class="Insecta" family="Passallidae" genus="Pharochilus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D59015BFEBB13AF068245ED" box="[332,468,1484,1508]" italics="true" pageId="36" pageNumber="37">Pharochilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D59015BFDFA13AF05C045ED" authorityName="Kaup" authorityYear="1868" box="[525,662,1484,1508]" class="Insecta" family="Passallidae" genus="Mastachilus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D59015BFDFA13AF05C045ED" box="[525,662,1484,1508]" italics="true" pageId="36" pageNumber="37">Mastachilus</emphasis>
|
||
</taxonomicName>
|
||
, are rare. Despite this beetle species being collected frequently (n> 100), these two specimens are the only megisthanids collected from this host. Therefore it is a rare species on this host, at least in southeast
|
||
<collectingRegion id="49DB45A48D59015BFE421077056A4624" box="[437,572,1556,1581]" country="Australia" name="Queensland" pageId="36" pageNumber="37">Queensland</collectingRegion>
|
||
, where my collecting effort was concentrated (
|
||
<bibRefCitation id="EF8EF6B78D59015BFBBF107703B04625" author="Seeman, O. D." box="[1096,1254,1556,1580]" pageId="36" pageNumber="37" refId="ref22681" refString="Seeman, O. D. (2002) Mites and Passalid Beetles: Diversity, Taxonomy and Biogeography. PhD thesis. The University of Queensland: Brisbane, Australia." type="book" year="2002">Seeman 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |