365 lines
60 KiB
XML
365 lines
60 KiB
XML
<document id="5F3B3A80DF49078D40F650954C4B8465" ID-CLB-Dataset="9118" ID-DOI="10.11646/zootaxa.4852.2.7" ID-GBIF-Dataset="d3b8188f-cbf5-4420-b0a7-416610730865" ID-ISSN="1175-5326" ID-Zenodo-Dep="4409834" ID-ZooBank="082655D6-BC36-4B11-9468-15E5FE6DBCFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609594253387" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03F62C2BCC1BFFB657E5FBCAFBF3FD93" docLanguage="en" docName="zootaxa.4852.2.7.pdf" docOrigin="Zootaxa 4852 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marcepania halva Jałoszyński 2020, sp. n." docType="treatment" docVersion="5" lastPageNumber="226" masterDocId="FFCF5453CC1AFFB55772FFF5FFBDFF8A" masterDocTitle="Three new species of Marcepania Jałoszyński (Coleoptera: Staphylinidae Scydmaeninae)" masterLastPageNumber="230" masterPageNumber="223" pageNumber="224" updateTime="1698850971170" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C7C50F34A996DDEDB594DA6FE2A33F84">Three new species of Marcepania Jałoszyński (Coleoptera: Staphylinidae Scydmaeninae)</mods:title>
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<mods:name id="E4A66C4BB4A9736897ED5472AA96330E" type="personal">
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<mods:namePart id="5D2ADEE96888AD858CEB2707111E88A4">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="1A730A16E94943B47CA872DD890B0D1A">Zootaxa</mods:title>
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<mods:part id="9271CE0D411E425D03DA236800829E0A">
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<mods:date id="A080ED09961552FE4CFEA72EBF8A8D40">2020</mods:date>
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<mods:number id="6F837E29749674BBCE6F1A5B493C1866">2020-09-15</mods:number>
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<mods:number id="22C367F374C8D5B0C4354C1AB9A02033">4852</mods:number>
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<mods:identifier id="7ECA25C957EF167AF4D23E1D15364C56" type="DOI">10.11646/zootaxa.4852.2.7</mods:identifier>
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<treatment id="03F62C2BCC1BFFB657E5FBCAFBF3FD93" ID-DOI="http://doi.org/10.5281/zenodo.4409836" ID-GBIF-Taxon="176416152" ID-Zenodo-Dep="4409836" LSID="urn:lsid:plazi:treatment:03F62C2BCC1BFFB657E5FBCAFBF3FD93" httpUri="http://treatment.plazi.org/id/03F62C2BCC1BFFB657E5FBCAFBF3FD93" lastPageId="3" lastPageNumber="226" pageId="1" pageNumber="224">
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<subSubSection id="C345CEB6CC1BFFB457E5FBCAFE03FBD3" box="[151,446,1086,1113]" pageId="1" pageNumber="224" type="nomenclature">
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<paragraph id="8BE09D3DCC1BFFB457E5FBCAFE03FBD3" blockId="1.[151,446,1086,1149]" box="[151,446,1086,1113]" pageId="1" pageNumber="224">
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<heading id="D0A82A51CC1BFFB457E5FBCAFE03FBD3" bold="true" box="[151,446,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="224" reason="1">
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<emphasis id="B92B412FCC1BFFB457E5FBCAFE03FBD3" bold="true" box="[151,446,1086,1113]" pageId="1" pageNumber="224">
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<taxonomicName id="4C5FE6BECC1BFFB457E5FBCAFEC9FBD3" ID-CoL="8Y67N" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,372,1086,1113]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="halva" status="sp. nov.">
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<emphasis id="B92B412FCC1BFFB457E5FBCAFEC9FBD3" bold="true" box="[151,372,1086,1113]" italics="true" pageId="1" pageNumber="224">Marcepania halva</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A218FC54CC1BFFB45609FBCAFE03FBD3" box="[379,446,1087,1113]" pageId="1" pageNumber="224" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C345CEB6CC1BFFB457E5FB96FE9AFBF7" box="[151,295,1123,1149]" pageId="1" pageNumber="224" type="description">
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<paragraph id="8BE09D3DCC1BFFB457E5FB96FE9AFBF7" blockId="1.[151,446,1086,1149]" box="[151,295,1123,1149]" pageId="1" pageNumber="224">
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(
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<figureCitation id="136481B8CC1BFFB457EDFB96FF5BFBF7" box="[159,230,1123,1149]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[161,1426,280,1273]" captionTargetId="figure-43@2.[151,1436,270,1283]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of Marcepania halva sp. n. (1), M. krowka sp. n. (2), and M. daifuku sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4409838" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="1" pageNumber="224">Figs 1</figureCitation>
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,
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<figureCitation id="136481B8CC1BFFB45786FB96FEA2FBF7" box="[244,287,1123,1149]" captionStart-0="FIGURES 4–7" captionStart-1="FIGURES 8–13" captionStartId-0="3.[151,264,1835,1859]" captionStartId-1="4.[151,264,1860,1884]" captionTargetBox-0="[170,1401,600,1782]" captionTargetBox-1="[159,1424,435,1829]" captionTargetId-0="figure-198@3.[170,1416,561,1782]" captionTargetId-1="figure-84@4.[151,1436,417,1836]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." captionText-1="FIGURES 8–13. Aedeagus in ventral view. Marcepania halva sp. n. (8–9), M. krowka sp. n. (10–11; one paramere broken off), and M. daifuku sp. n. (12–13; 12 includes also a fragment of abdominal segment, omitted in 13)." figureDoi-0="http://doi.org/10.5281/zenodo.4409840" figureDoi-1="http://doi.org/10.5281/zenodo.4409842" httpUri-0="https://zenodo.org/record/4409840/files/figure.png" httpUri-1="https://zenodo.org/record/4409842/files/figure.png" pageId="1" pageNumber="224">4–9</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C345CEB6CC1BFFB457E5FB5EFB78FABB" pageId="1" pageNumber="224" type="materials_examined">
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<paragraph id="8BE09D3DCC1BFFB457E5FB5EFB78FABB" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="224">
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<emphasis id="B92B412FCC1BFFB457E5FB5EFEF8FB4E" bold="true" box="[151,325,1194,1221]" pageId="1" pageNumber="224">Type material.</emphasis>
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<materialsCitation id="3B379760CC1BFFB45639FB5EFC8CFA87" ID-GBIF-Occurrence="3008330304" collectingDate="1989-12-02" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Indonesia" elevation="1500" location="Brastagi" pageId="1" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
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||
<emphasis id="B92B412FCC1BFFB45639FB5EFC13FB4E" bold="true" box="[331,942,1194,1221]" pageId="1" pageNumber="224">
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||
<typeStatus id="54E4239FCC1BFFB45639FB5EFE01FB4F" box="[331,444,1195,1221]" pageId="1" pageNumber="224" type="holotype">Holotype</typeStatus>
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:
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||
<collectingCountry id="F348DDADCC1BFFB456BAFB5FFDD8FB4E" box="[456,613,1194,1220]" name="Indonesia" pageId="1" pageNumber="224">INDONESIA</collectingCountry>
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||
(
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||
<collectingRegion id="499B53DFCC1BFFB45506FB5FFC1FFB4E" box="[628,930,1194,1220]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="224">North Sumatra Province</collectingRegion>
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):
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</emphasis>
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<specimenCount id="9D5956B4CC1BFFB454C7FB5EFC71FB4F" box="[949,972,1195,1221]" pageId="1" pageNumber="224" type="male">♂</specimenCount>
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, two labels: “
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<collectingRegion id="499B53DFCC1BFFB45310FB5EFB52FB4F" box="[1122,1263,1195,1221]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="224">SUMATRA</collectingRegion>
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: N Sum. /
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||
<quantity id="4CA730D8CC1BFFB45216FB5EFA21FB4F" box="[1380,1436,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="224" unit="km" value="7.0">7 km</quantity>
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N
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<location id="8E80CBE6CC1BFFB457C0FB3AFEB2FB63" LSID="urn:lsid:plazi:treatment:03F62C2BCC1BFFB657E5FBCAFBF3FD93:8E80CBE6CC1BFFB457C0FB3AFEB2FB63" box="[178,271,1231,1257]" country="Indonesia" name="Brastagi" pageId="1" pageNumber="224" stateProvince="Sumatera Utara">Brastagi</location>
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/
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<quantity id="4CA730D8CC1BFFB45656FB3BFEC6FB63" box="[292,379,1230,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="224" unit="m" value="1500.0">
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<elevation id="00727A0ECC1BFFB45656FB3BFEC6FB63" box="[292,379,1230,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="224" unit="m" value="1500.0">1500 m</elevation>
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</quantity>
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||
,
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||
<date id="FFE1BBFDCC1BFFB456F5FB3AFDBFFB63" box="[391,514,1231,1257]" pageId="1" pageNumber="224" value="1989-12-02">
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<collectingDate id="EFA54215CC1BFFB456F5FB3AFDBFFB63" box="[391,514,1231,1257]" pageId="1" pageNumber="224" value="1989-12-02">2.XII.1989</collectingDate>
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||
</date>
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||
/ Agosti, Löbl / Burckhardt # 28a” [white, printed], “
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||
<taxonomicName id="4C5FE6BECC1BFFB45326FB3AFB40FB63" box="[1108,1277,1231,1257]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92B412FCC1BFFB45326FB3AFB40FB63" box="[1108,1277,1231,1257]" italics="true" pageId="1" pageNumber="224">MARCEPANIA</emphasis>
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||
</taxonomicName>
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||
/
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||
<taxonomicName id="4C5FE6BECC1BFFB45260FB3AFAEDFB63" authorityName="Jałoszyński" authorityYear="2020" box="[1298,1360,1231,1257]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="halva">
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<emphasis id="B92B412FCC1BFFB45260FB3AFAEDFB63" box="[1298,1360,1231,1257]" italics="true" pageId="1" pageNumber="224">halva</emphasis>
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</taxonomicName>
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m. /
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<collectorName id="26AAF8EBCC1BFFB452FBFB3AFEA6FA87" pageId="1" pageNumber="224">P. Jałoszyński</collectorName>
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, 2020 /
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<typeStatus id="54E4239FCC1BFFB45600FB06FDA1FA87" box="[370,540,1267,1293]" pageId="1" pageNumber="224" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="ED4E05F8CC1BFFB455BEFB06FC95FA87" box="[716,808,1267,1293]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="224" type="Museum">MHNG</collectionCode>
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)
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</materialsCitation>
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||
.
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<materialsCitation id="3B379760CC1BFFB45448FB06FB7CFABB" ID-GBIF-Occurrence="3008330301" collectingDate="1989-12-02" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Indonesia" elevation="1500" location="Brastagi" pageId="1" pageNumber="224" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paratype">
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<emphasis id="B92B412FCC1BFFB45448FB06FC0FFA87" bold="true" box="[826,946,1267,1293]" pageId="1" pageNumber="224">
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||
<typeStatus id="54E4239FCC1BFFB45448FB06FC0FFA87" box="[826,946,1267,1293]" pageId="1" pageNumber="224" type="paratype">Paratypes</typeStatus>
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</emphasis>
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||
: 1 disarticulated female mounted in Canada balsam and 3 dry-mounted specimen of unknown sex, same data as for holotype (cPJ,
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<collectionCode id="ED4E05F8CC1BFFB4532DFAE2FB07FABB" box="[1119,1210,1303,1329]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="224" type="Museum">MHNG</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345CEB6CC1BFFB457B5FACFFBBAFA4B" pageId="1" pageNumber="224" type="diagnosis">
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<paragraph id="8BE09D3DCC1BFFB457B5FACFFBBAFA4B" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="224">
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<emphasis id="B92B412FCC1BFFB457B5FACFFEFFFADE" bold="true" box="[199,322,1338,1364]" pageId="1" pageNumber="224">Diagnosis.</emphasis>
|
||
Body relatively stout, EI approximately 1.3, and large (BL slightly above
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<quantity id="4CA730D8CC1BFFB453E2FACEFB6BFADF" box="[1168,1238,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="224" unit="mm" value="1.0">1 mm</quantity>
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||
); head, pronotum and elytra with distinct, dense punctures; aedeagus elongate and subtrapezoidal with oblique distal margin; endophallus with a pair of well demarcated elongate structures, only slightly darker than surrounding areas; parameres much longer than median lobe, each with two conspicuously long apical setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C345CEB6CC1BFFB757B5FA3FFAABFA4C" lastPageId="2" lastPageNumber="225" pageId="1" pageNumber="224" type="description">
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<paragraph id="8BE09D3DCC1BFFB457B5FA3FFE26F983" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="224">
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||
<emphasis id="B92B412FCC1BFFB457B5FA3FFEE4FA6E" bold="true" box="[199,345,1482,1508]" pageId="1" pageNumber="224">Description.</emphasis>
|
||
Body of male (
|
||
<figureCitation id="136481B8CC1BFFB45570FA3EFDFFFA6F" box="[514,578,1483,1509]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[161,1426,280,1273]" captionTargetId="figure-43@2.[151,1436,270,1283]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of Marcepania halva sp. n. (1), M. krowka sp. n. (2), and M. daifuku sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4409838" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="1" pageNumber="224">Fig. 1</figureCitation>
|
||
) elongate and slender, moderately convex, light brown, covered with yellowish setae; BL
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||
<quantity id="4CA730D8CC1BFFB4565DFA1AFE2BF983" box="[303,406,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="1" pageNumber="224" unit="mm" value="1.08">1.08 mm</quantity>
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||
.
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||
</paragraph>
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<paragraph id="8BE09D3DCC1BFFB457B5F9E6FD40F96B" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="224">
|
||
Head (
|
||
<figureCitation id="136481B8CC1BFFB45662F9E6FEE8F9A7" box="[272,341,1555,1581]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[161,1426,280,1273]" captionTargetId="figure-43@2.[151,1436,270,1283]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of Marcepania halva sp. n. (1), M. krowka sp. n. (2), and M. daifuku sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4409838" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="1" pageNumber="224">Figs 1</figureCitation>
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||
,
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<figureCitation id="136481B8CC1BFFB45610F9E6FECCF9A7" box="[354,369,1555,1581]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="1" pageNumber="224">4</figureCitation>
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) broadest at moderately large, weakly convex, oval and coarsely faceted eyes, HL
|
||
<quantity id="4CA730D8CC1BFFB4538DF9E6FAD9F9A7" box="[1279,1380,1555,1581]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="224" unit="mm" value="0.1">0.10 mm</quantity>
|
||
, HW
|
||
<quantity id="4CA730D8CC1BFFB457E5F9C3FF43F9DB" box="[151,254,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="224" unit="mm" value="0.25">0.25 mm</quantity>
|
||
; vertex and posterior region of frons confluent and weakly convex; anterior portion of frons flattened; supraantennal tubercles barely marked; frons and vertex covered with dense and distinct but shallow punctures; setae sparse and short, suberect.Antennae (
|
||
<figureCitation id="136481B8CC1BFFB4554AF98AFDC3F913" box="[568,638,1663,1689]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[161,1426,280,1273]" captionTargetId="figure-43@2.[151,1436,270,1283]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of Marcepania halva sp. n. (1), M. krowka sp. n. (2), and M. daifuku sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4409838" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="1" pageNumber="224">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="136481B8CC1BFFB455F9F98AFD27F913" box="[651,666,1663,1689]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="1" pageNumber="224">4</figureCitation>
|
||
) short, AnL
|
||
<quantity id="4CA730D8CC1BFFB45456F98AFC37F913" box="[804,906,1663,1689]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="224" unit="mm" value="0.43">0.43 mm</quantity>
|
||
; antennomeres 1–2 strongly elongate; 4 slightly transverse, 5–7 each elongate; 8 and 9 each almost as long as broad, 10 indistinctly transverse, 11 much shorter than 9–10, about 1.8 × as long as broad, with truncate apex.
|
||
</paragraph>
|
||
<paragraph id="8BE09D3DCC1BFFB457B5F91EFE2BF833" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="224">
|
||
Prothorax (
|
||
<figureCitation id="136481B8CC1BFFB45631F91EFE34F88F" box="[323,393,1771,1797]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[161,1426,280,1273]" captionTargetId="figure-43@2.[151,1436,270,1283]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of Marcepania halva sp. n. (1), M. krowka sp. n. (2), and M. daifuku sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4409838" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="1" pageNumber="224">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="136481B8CC1BFFB456E4F91FFE19F88F" box="[406,420,1770,1797]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="1" pageNumber="224">5</figureCitation>
|
||
) nearly semi-oval, broadest slightly in front of middle; PL
|
||
<quantity id="4CA730D8CC1BFFB45342F91FFB28F88F" box="[1072,1173,1770,1797]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="224" unit="mm" value="0.35">0.35 mm</quantity>
|
||
, PW
|
||
<quantity id="4CA730D8CC1BFFB453BDF91EFA88F88F" box="[1231,1333,1771,1797]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="224" unit="mm" value="0.43">0.43 mm</quantity>
|
||
.Anterior margin rounded; lateral margins strongly and nearly evenly rounded in anterior half, nearly straight and weakly convergent posterad in posterior half; hind corners weakly obtuse-angled; posterior margin shallowly bisinuate; lateral ante-basal impressions small and shallow but with relatively sharp margins. Punctures on disc distinct, relatively sharply marked and dense, those near middle separated by spaces slightly shorter than puncture diameters; setae sparse, short, suberect.
|
||
</paragraph>
|
||
<paragraph id="8BE09D3DCC1BFFB757B5F836FDA7FF5F" blockId="1.[151,1437,1194,2013]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="225" pageId="1" pageNumber="224">
|
||
Elytra (
|
||
<figureCitation id="136481B8CC1BFFB4566BF836FEE2F857" box="[281,351,1987,2013]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1309,1333]" captionTargetBox="[161,1426,280,1273]" captionTargetId="figure-43@2.[151,1436,270,1283]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus of Marcepania halva sp. n. (1), M. krowka sp. n. (2), and M. daifuku sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.4409838" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="1" pageNumber="224">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="136481B8CC1BFFB4561EF836FEC6F857" box="[364,379,1987,2013]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="1" pageNumber="224">7</figureCitation>
|
||
) together oval, broadest near anterior third; EL
|
||
<quantity id="4CA730D8CC1BFFB454FAF836FC50F857" box="[904,1005,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="224" unit="mm" value="0.63">0.63 mm</quantity>
|
||
, EW
|
||
<quantity id="4CA730D8CC1BFFB4535AF836FB30F857" box="[1064,1165,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="224" unit="mm" value="0.48">0.48 mm</quantity>
|
||
, EI 1.32; apices broadly, separately rounded; punctures similar to those on pronotum but denser, partly arranged into irregular longitudinal rows; setae short, sparse, suberect.
|
||
</paragraph>
|
||
<paragraph id="8BE09D3DCC18FFB757B5FF2AFD95FF73" blockId="2.[151,1436,151,249]" box="[199,552,223,249]" pageId="2" pageNumber="225">Legs short, robust; unmodified.</paragraph>
|
||
<caption id="DF20CDB5CC18FFB757E5FAE8FB40FABF" ID-DOI="http://doi.org/10.5281/zenodo.4409838" ID-Zenodo-Dep="4409838" box="[151,1277,1309,1333]" httpUri="https://zenodo.org/record/4409838/files/figure.png" pageId="2" pageNumber="225" startId="2.[151,264,1309,1333]" targetBox="[161,1426,280,1273]" targetPageId="2">
|
||
<paragraph id="8BE09D3DCC18FFB757E5FAE8FB40FABF" blockId="2.[151,1277,1309,1333]" box="[151,1277,1309,1333]" pageId="2" pageNumber="225">
|
||
<emphasis id="B92B412FCC18FFB757E5FAE8FE87FABF" bold="true" box="[151,314,1309,1333]" pageId="2" pageNumber="225">FIGURES 1–3.</emphasis>
|
||
Dorsal habitus of
|
||
<taxonomicName id="4C5FE6BECC18FFB75684FAEBFD12FABF" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[502,687,1309,1333]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="halva" status="sp. nov.">
|
||
<emphasis id="B92B412FCC18FFB75684FAEBFD12FABF" box="[502,687,1309,1333]" italics="true" pageId="2" pageNumber="225">Marcepania halva</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92B412FCC18FFB755C7FAE8FD53FABF" bold="true" box="[693,750,1309,1333]" pageId="2" pageNumber="225">
|
||
<taxonomicNameLabel id="A218FC54CC18FFB755C7FAE8FD53FABF" box="[693,750,1309,1333]" pageId="2" pageNumber="225" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(1),
|
||
<taxonomicName id="4C5FE6BECC18FFB7546DFAEBFC36FABF" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[799,907,1309,1333]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="krowka" status="sp. nov.">
|
||
<emphasis id="B92B412FCC18FFB7546DFAEBFC36FABF" box="[799,907,1309,1333]" italics="true" pageId="2" pageNumber="225">M. krowka</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92B412FCC18FFB754E3FAE8FC77FABF" bold="true" box="[913,970,1309,1333]" pageId="2" pageNumber="225">
|
||
<taxonomicNameLabel id="A218FC54CC18FFB754E3FAE8FC77FABF" box="[913,970,1309,1333]" pageId="2" pageNumber="225" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(2), and
|
||
<taxonomicName id="4C5FE6BECC18FFB75357FAEBFB2EFABF" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[1061,1171,1309,1333]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="daifuku" status="sp. nov.">
|
||
<emphasis id="B92B412FCC18FFB75357FAEBFB2EFABF" box="[1061,1171,1309,1333]" italics="true" pageId="2" pageNumber="225">M. daifuku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92B412FCC18FFB753EBFAE8FB6FFABF" bold="true" box="[1177,1234,1309,1333]" pageId="2" pageNumber="225">
|
||
<taxonomicNameLabel id="A218FC54CC18FFB753EBFAE8FB6FFABF" box="[1177,1234,1309,1333]" pageId="2" pageNumber="225" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(3).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE09D3DCC18FFB757B5FA96FAABFA4C" blockId="2.[151,1437,1379,2018]" pageId="2" pageNumber="225">
|
||
Aedeagus (
|
||
<figureCitation id="136481B8CC18FFB75631FA96FE1BFAF4" box="[323,422,1379,1406]" captionStart="FIGURES 8–13" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[159,1424,435,1829]" captionTargetId="figure-84@4.[151,1436,417,1836]" captionTargetPageId="4" captionText="FIGURES 8–13. Aedeagus in ventral view. Marcepania halva sp. n. (8–9), M. krowka sp. n. (10–11; one paramere broken off), and M. daifuku sp. n. (12–13; 12 includes also a fragment of abdominal segment, omitted in 13)." figureDoi="http://doi.org/10.5281/zenodo.4409842" httpUri="https://zenodo.org/record/4409842/files/figure.png" pageId="2" pageNumber="225">Figs 8–9</figureCitation>
|
||
) moderately stout; AeL
|
||
<quantity id="4CA730D8CC18FFB755C0FA96FCA5FAF4" box="[690,792,1379,1406]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="225" unit="mm" value="0.15">0.15 mm</quantity>
|
||
; median lobe in ventral view elongate subtrapezoidal with oblique distal margin; endophallic structures weakly sclerotized, with pair of elongate fusiform components only slightly darker than surrounding areas; parameres conspicuously long, each with two long apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345CEB6CC18FFB757B5FA3AFD1CFA60" box="[199,673,1487,1514]" pageId="2" pageNumber="225" type="distribution">
|
||
<paragraph id="8BE09D3DCC18FFB757B5FA3AFD1CFA60" blockId="2.[151,1437,1379,2018]" box="[199,673,1487,1514]" pageId="2" pageNumber="225">
|
||
<emphasis id="B92B412FCC18FFB757B5FA3AFEDCFA63" bold="true" box="[199,353,1487,1513]" pageId="2" pageNumber="225">Distribution.</emphasis>
|
||
Northern
|
||
<collectingRegion id="499B53DFCC18FFB756A6FA25FD8CFA60" box="[468,561,1488,1514]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="225">Sumatra</collectingRegion>
|
||
(
|
||
<figureCitation id="136481B8CC18FFB75532FA3AFD29FA60" box="[576,660,1487,1514]" captionStart="FIGURE 14" captionStartId="6.[151,250,1505,1529]" captionTargetBox="[151,1436,741,1481]" captionTargetId="figure-226@6.[151,1436,741,1481]" captionTargetPageId="6" captionText="FIGURE 14. Distribution of Marcepania species." figureDoi="http://doi.org/10.5281/zenodo.4409846" httpUri="https://zenodo.org/record/4409846/files/figure.png" pageId="2" pageNumber="225">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345CEB6CC18FFB757B5FA01FD14F9B8" pageId="2" pageNumber="225" type="etymology">
|
||
<paragraph id="8BE09D3DCC18FFB757B5FA01FD14F9B8" blockId="2.[151,1437,1379,2018]" pageId="2" pageNumber="225">
|
||
<emphasis id="B92B412FCC18FFB757B5FA01FEF3F984" bold="true" box="[199,334,1524,1550]" pageId="2" pageNumber="225">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C5FE6BECC18FFB756BFFA06FDB6F987" authorityName="Jałoszyński" authorityYear="2020" box="[461,523,1523,1549]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="halva">
|
||
<emphasis id="B92B412FCC18FFB756BFFA06FDB6F987" box="[461,523,1523,1549]" italics="true" pageId="2" pageNumber="225">halva</emphasis>
|
||
</taxonomicName>
|
||
(a noun in apposition) refers to the Middle East sweet confection (as fitting well the genus name that is derived from marzipan).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345CEB6CC18FFB657B5F9C9FBF3FD93" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225" type="discussion">
|
||
<paragraph id="8BE09D3DCC18FFB657B5F9C9FC90FEEF" blockId="2.[151,1437,1379,2018]" lastBlockId="3.[151,1437,151,537]" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225">
|
||
<emphasis id="B92B412FCC18FFB757B5F9C9FE86F9DC" bold="true" box="[199,315,1596,1622]" pageId="2" pageNumber="225">Remarks.</emphasis>
|
||
This species has the largest and most convex adults; this is the only known
|
||
<taxonomicName id="4C5FE6BECC18FFB7530CF9C9FABBF9DF" box="[1150,1286,1596,1621]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B412FCC18FFB7530CF9C9FABBF9DF" box="[1150,1286,1596,1621]" italics="true" pageId="2" pageNumber="225">Marcepania</emphasis>
|
||
</taxonomicName>
|
||
whose adults exceed
|
||
<quantity id="4CA730D8CC18FFB7579EF995FE93F9F0" box="[236,302,1632,1658]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="225" unit="mm" value="1.0">1 mm</quantity>
|
||
in length. The unusually convex body prompted detailed morphological examination to confirm that this is indeed a
|
||
<taxonomicName id="4C5FE6BECC18FFB75634F971FE73F917" box="[326,462,1668,1693]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B412FCC18FFB75634F971FE73F917" box="[326,462,1668,1693]" italics="true" pageId="2" pageNumber="225">Marcepania</emphasis>
|
||
</taxonomicName>
|
||
and not an unknown genus. Structures of a disarticulated female (
|
||
<figureCitation id="136481B8CC18FFB753C1F976FAABF914" box="[1203,1302,1667,1694]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Figs 4–7</figureCitation>
|
||
) do not differ in any detail from those known in previously examined species (
|
||
<bibRefCitation id="EFCEE0CCCC18FFB754E1F952FBE7F948" author="Jaloszynski, P." box="[915,1114,1703,1730]" pageId="2" pageNumber="225" pagination="159 - 189" refId="ref3559" refString="Jaloszynski, P. (2013) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. https: // doi. org / 10.1111 / syen. 12044" type="journal article" year="2013">Jałoszyński 2013</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C5FE6BECC18FFB7531FF95DFB6CF94B" authorityName="Jałoszyński" authorityYear="2020" box="[1133,1233,1703,1729]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="225" phylum="Arthropoda" rank="species" species="halva">
|
||
<emphasis id="B92B412FCC18FFB7531FF95DFB6CF94B" box="[1133,1233,1703,1729]" italics="true" pageId="2" pageNumber="225">M. halva</emphasis>
|
||
</taxonomicName>
|
||
has asymmetrical mandibles, the right one with pre-apical tooth (
|
||
<figureCitation id="136481B8CC18FFB755C7F93EFD41F96C" box="[693,764,1739,1766]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75478F939FC93F96F" box="[778,814,1740,1765]" italics="true" pageId="2" pageNumber="225">pat</emphasis>
|
||
, lacking in the left one), button-shaped maxillary palpomere 4 (
|
||
<figureCitation id="136481B8CC18FFB75662F91AFEE8F880" box="[272,341,1775,1802]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75613F905FE23F880" box="[353,414,1776,1802]" italics="true" pageId="2" pageNumber="225">mxp4</emphasis>
|
||
), gradually thickened antennae, open procoxal cavities (
|
||
<figureCitation id="136481B8CC18FFB75364F91AFBE6F880" box="[1046,1115,1775,1802]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB7531AF905FB26F883" box="[1128,1179,1776,1801]" italics="true" pageId="2" pageNumber="225">pcxc</emphasis>
|
||
), complete notosternal sutures (
|
||
<figureCitation id="136481B8CC18FFB75787F8E6FE85F8A4" box="[245,312,1811,1838]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75631F8E1FEDAF8A7" box="[323,359,1812,1837]" italics="true" pageId="2" pageNumber="225">nss</emphasis>
|
||
; poorly visible in a photograph, but confirmed as complete under microscope), narrow inner (i.e., adcoxal) portions of hypomera (
|
||
<figureCitation id="136481B8CC18FFB75570F8C2FDFAF8D8" box="[514,583,1847,1874]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75526F8C2FDCBF8DB" box="[596,630,1847,1873]" italics="true" pageId="2" pageNumber="225">ihy</emphasis>
|
||
), a massive mesoventral intercoxal process (
|
||
<figureCitation id="136481B8CC18FFB75302F8C2FB08F8DB" box="[1136,1205,1847,1873]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB753B1F8CDFB40F8DB" box="[1219,1277,1848,1873]" italics="true" pageId="2" pageNumber="225">msvp</emphasis>
|
||
) clearly separating mesocoxae, its posterior margin bearing a long and narrow median longitudinal carina (
|
||
<figureCitation id="136481B8CC18FFB753C8F8AEFB42F8FF" box="[1210,1279,1883,1909]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB7527EF8AEFA89F8FF" box="[1292,1332,1883,1909]" italics="true" pageId="2" pageNumber="225">mlc</emphasis>
|
||
) running posteriorly, lateral metaventral carinae (
|
||
<figureCitation id="136481B8CC18FFB75523F88AFD29F813" box="[593,660,1919,1945]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB755D2F88AFD68F813" box="[672,725,1919,1945]" italics="true" pageId="2" pageNumber="225">lmvc</emphasis>
|
||
) and lateral metaventral foveae (
|
||
<figureCitation id="136481B8CC18FFB75333F88AFB38F813" box="[1089,1157,1919,1945]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 6</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB753E2F88AFB7DF813" box="[1168,1216,1919,1945]" italics="true" pageId="2" pageNumber="225">lmvf</emphasis>
|
||
), broadly separated metacoxae, a transverse mesoscutum (
|
||
<figureCitation id="136481B8CC18FFB75533F856FD39F834" box="[577,644,1955,1982]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB755E2F851FD0BF834" box="[656,694,1956,1982]" italics="true" pageId="2" pageNumber="225">sc2</emphasis>
|
||
) and mesoscutellum (
|
||
<figureCitation id="136481B8CC18FFB754D8F856FC53F834" box="[938,1006,1955,1982]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75488F856FB9AF837" box="[1018,1063,1955,1981]" italics="true" pageId="2" pageNumber="225">scl2</emphasis>
|
||
) with a distinct scutoscutellar suture (
|
||
<figureCitation id="136481B8CC18FFB757A0F832FEABF868" box="[210,278,1991,2018]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75650F83DFEFEF86B" box="[290,323,1992,2017]" italics="true" pageId="2" pageNumber="225">sss</emphasis>
|
||
), and well-marked humeral denticles (
|
||
<figureCitation id="136481B8CC18FFB75580F832FC88F868" box="[754,821,1991,2018]" captionStart="FIGURES 4–7" captionStartId="3.[151,264,1835,1859]" captionTargetBox="[170,1401,600,1782]" captionTargetId="figure-198@3.[170,1416,561,1782]" captionTargetPageId="3" captionText="FIGURES 4–7. Morphological structures of Marcepania halva sp. n. with structures important for genus identification indicated. Head in ventral view (4); prothorax in ventral view (5); pterothorax in ventral view (6); elytral base with meso- and metanotum in dorsal view (7). Abbreviations: hd, humeral denticle; ihy, inner portion of hypomeron; lmvc, lateral metaventral carina; lmvf, lateral metaventral fovea; mlc, median longitudinal carina; msvp, mesoventral intercoxal process; mxp4, maxillary palpomere 4; nss, notosternal suture; pat, pre-apical tooth; pcxc, procoxal cavity; sc2, mesoscutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4409840" httpUri="https://zenodo.org/record/4409840/files/figure.png" pageId="2" pageNumber="225">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="B92B412FCC18FFB75430F832FCE3F86B" box="[834,862,1991,2017]" italics="true" pageId="2" pageNumber="225">hd</emphasis>
|
||
). Together with a weakly sclerotized, plate-shaped, simple aedeagus with slender parameres, this set of characters agrees with that given in the diagnosis of
|
||
<emphasis id="B92B412FCC19FFB6525DFF62FF06FF5E" italics="true" pageId="3" pageNumber="226">Marcepa- nia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCEE0CCCC19FFB657BBFF4EFE2DFF5F" author="Jaloszynski, P." box="[201,400,187,213]" pageId="3" pageNumber="226" pagination="159 - 189" refId="ref3559" refString="Jaloszynski, P. (2013) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. https: // doi. org / 10.1111 / syen. 12044" type="journal article" year="2013">Jałoszyński 2013</bibRefCitation>
|
||
). An additional character was found in
|
||
<taxonomicName id="4C5FE6BECC19FFB65434FF4EFC17FF5F" authorityName="Jałoszyński" authorityYear="2020" box="[838,938,187,213]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="halva">
|
||
<emphasis id="B92B412FCC19FFB65434FF4EFC17FF5F" box="[838,938,187,213]" italics="true" pageId="3" pageNumber="226">M. halva</emphasis>
|
||
</taxonomicName>
|
||
(and, less pronounced, in the two remaining newly described species), i.e., setiferous punctures on elytra partly arranged in longitudinal rows. This feature is poorly visible in small specimens, and it may occur in all
|
||
<taxonomicName id="4C5FE6BECC19FFB65467FEF6FC20FE96" box="[789,925,259,284]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B412FCC19FFB65467FEF6FC20FE96" box="[789,925,259,284]" italics="true" pageId="3" pageNumber="226">Marcepania</emphasis>
|
||
</taxonomicName>
|
||
species, but only in the large-bodied
|
||
<taxonomicName id="4C5FE6BECC19FFB65248FEF6FA21FE97" authorityName="Jałoszyński" authorityYear="2020" box="[1338,1436,259,285]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="halva">
|
||
<emphasis id="B92B412FCC19FFB65248FEF6FA21FE97" box="[1338,1436,259,285]" italics="true" pageId="3" pageNumber="226">M. halva</emphasis>
|
||
</taxonomicName>
|
||
it is distinct enough to be easily noticed under a stereomicroscope. The rows of setae and setiferous punctures are better visible in transparent mounts of disarticulated elytra.
|
||
</paragraph>
|
||
<paragraph id="8BE09D3DCC19FFB657B5FE9AFBF3FD93" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="226">
|
||
The aedeagus of
|
||
<taxonomicName id="4C5FE6BECC19FFB656FDFE9AFE4BFE03" authorityName="Jałoszyński" authorityYear="2020" box="[399,502,367,393]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="halva">
|
||
<emphasis id="B92B412FCC19FFB656FDFE9AFE4BFE03" box="[399,502,367,393]" italics="true" pageId="3" pageNumber="226">M. halva</emphasis>
|
||
</taxonomicName>
|
||
is most similar to that of
|
||
<taxonomicName id="4C5FE6BECC19FFB65459FE9AFB6EFE03" authority="Jaloszynski, 2013" authorityName="Jaloszynski" authorityYear="2013" box="[811,1235,367,393]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="semengohensis">
|
||
<emphasis id="B92B412FCC19FFB65459FE9AFC46FE03" box="[811,1019,367,393]" italics="true" pageId="3" pageNumber="226">M. semengohensis</emphasis>
|
||
<bibRefCitation id="EFCEE0CCCC19FFB65377FE9AFB6EFE03" author="Jaloszynski, P." box="[1029,1235,367,393]" pageId="3" pageNumber="226" pagination="159 - 189" refId="ref3559" refString="Jaloszynski, P. (2013) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. https: // doi. org / 10.1111 / syen. 12044" type="journal article" year="2013">Jałoszyński, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5FE6BECC19FFB65261FE9AFED8FE27" authority="Jaloszynski, 2019" authorityName="Jaloszynski" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="princesa">
|
||
<emphasis id="B92B412FCC19FFB65261FE9AFA21FE02" box="[1299,1436,367,392]" italics="true" pageId="3" pageNumber="226">M. princesa</emphasis>
|
||
<bibRefCitation id="EFCEE0CCCC19FFB657E5FE66FED8FE27" author="Jaloszynski, P." box="[151,357,403,429]" pageId="3" pageNumber="226" pagination="141 - 144" refId="ref3613" refString="Jaloszynski, P. (2019) The first Philippine species of Marcepania (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4668 (1), 141 - 144. https: // doi. org / 10.11646 / zootaxa. 4668.1.10" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
. In addition to proportionally much longer parameres and different endophallic structures,
|
||
<taxonomicName id="4C5FE6BECC19FFB6520CFE66FF68FE5B" authorityName="Jałoszyński" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="halva">
|
||
<emphasis id="B92B412FCC19FFB6520CFE66FF68FE5B" italics="true" pageId="3" pageNumber="226">M. halva</emphasis>
|
||
</taxonomicName>
|
||
differs from these two species in external characters: the male of the newly described species is distinctly stouter, with EI 1.32, versus
|
||
<quantity id="4CA730D8CC19FFB65697FE2EFDC9FE7F" box="[485,628,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.6449" metricValueMax="3.7338" metricValueMin="3.5559999999999996" pageId="3" pageNumber="226" unit="in" value="1.435" valueMax="1.47" valueMin="1.4">1.40–1.47 in</quantity>
|
||
<taxonomicName id="4C5FE6BECC19FFB6550CFE2EFCF0FE7F" authorityName="Jaloszynski" authorityYear="2013" box="[638,845,475,501]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="semengohensis">
|
||
<emphasis id="B92B412FCC19FFB6550CFE2EFCF0FE7F" box="[638,845,475,501]" italics="true" pageId="3" pageNumber="226">M. semengohensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="4CA730D8CC19FFB654FBFE2FFC64FE7F" box="[905,985,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.8608" pageId="3" pageNumber="226" unit="in" value="1.52">1.52 in</quantity>
|
||
<taxonomicName id="4C5FE6BECC19FFB65491FE2EFBD6FE7E" authorityName="Jaloszynski" authorityYear="2019" box="[995,1131,475,500]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="princesa">
|
||
<emphasis id="B92B412FCC19FFB65491FE2EFBD6FE7E" box="[995,1131,475,500]" italics="true" pageId="3" pageNumber="226">M. princesa</emphasis>
|
||
</taxonomicName>
|
||
. Together with
|
||
<taxonomicName id="4C5FE6BECC19FFB65251FE2EFA21FE7F" authorityName="Jałoszyński" authorityYear="2020" box="[1315,1436,475,501]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="species" species="krowka">
|
||
<emphasis id="B92B412FCC19FFB65251FE2EFA21FE7F" box="[1315,1436,475,501]" italics="true" pageId="3" pageNumber="226">M. krowka</emphasis>
|
||
</taxonomicName>
|
||
described below, this is the first record of the occurrence of
|
||
<taxonomicName id="4C5FE6BECC19FFB65445FE0AFC02FD92" box="[823,959,511,536]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="226" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B412FCC19FFB65445FE0AFC02FD92" box="[823,959,511,536]" italics="true" pageId="3" pageNumber="226">Marcepania</emphasis>
|
||
</taxonomicName>
|
||
on
|
||
<collectingRegion id="499B53DFCC19FFB6549AFE0AFBF4FD93" box="[1000,1097,511,537]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="226">Sumatra</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |