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<document id="7B734D03E76CC614E0CF760BFE75DBD3" ID-DOI="10.5281/zenodo.178011" ID-GBIF-Dataset="b02a028f-ba14-458e-be60-cd539f28d9d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="178011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459878369919" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2007" docId="03EE11580174FFBDFF68F92FFD8721DD" docLanguage="en" docName="zt01546p022.pdf" docOrigin="Zootaxa 1546" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brachinus piceus Chaudoir" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="FFD769200170FFB8FFFFFFDCFFB5251F" masterDocTitle="Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines" masterLastPageNumber="22" masterPageNumber="15" pageNumber="19" updateTime="1698227246572" updateUser="plazi">
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<mods:title id="13FDE2A8094AB9FDB14A28070754A532">Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines</mods:title>
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<mods:namePart id="C3FF248A5F3BBD11DAF769066ED4A2A0">Tian, Mingyi</mods:namePart>
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<mods:namePart id="190EA89DBFAED6EEF695C72C9DA09A0E">Deuve, Thierry</mods:namePart>
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<mods:typeOfResource id="64750F2FC3243DE75934430E24525176">text</mods:typeOfResource>
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<mods:title id="B1372F5AFFB4F23F056EEA9F07230129">Zootaxa</mods:title>
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<mods:date id="67AE6A8A41CCB02053C2D448D6FF8AFE">2007</mods:date>
<mods:detail id="6A9C9D94754853490F2D76F67D277447" type="volume">
<mods:number id="74FB8051B06757E2FA31BD2396A4D691">1546</mods:number>
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<mods:identifier id="B0FE9AC5073FDB01430E8CEFFDAA642F" type="DOI">10.5281/zenodo.178011</mods:identifier>
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<mods:identifier id="580A9ABC2706215F67D450C62CF5E73B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="345773167A5F90FD838B29A6359E809D" type="Zenodo-Dep">178011</mods:identifier>
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<treatment id="03EE11580174FFBDFF68F92FFD8721DD" ID-DOI="http://doi.org/10.5281/zenodo.6244094" ID-GBIF-Taxon="119370431" ID-Zenodo-Dep="6244094" LSID="urn:lsid:plazi:treatment:03EE11580174FFBDFF68F92FFD8721DD" httpUri="http://treatment.plazi.org/id/03EE11580174FFBDFF68F92FFD8721DD" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<subSubSection id="C35DF3C50174FFBCFF68F92FFEDB222F" pageId="4" pageNumber="19" type="nomenclature">
<paragraph id="8BF8A04E0174FFBCFF68F92FFE5C2212" blockId="4.[151,489,1779,1840]" box="[151,489,1779,1805]" pageId="4" pageNumber="19">
<heading id="D0B017220174FFBCFF68F92FFE5C2212" bold="true" box="[151,489,1779,1805]" fontSize="11" level="1" pageId="4" pageNumber="19" reason="1">
<taxonomicName id="4C47DBCD0174FFBCFF68F92FFE5C2212" ID-CoL="MTF6" authority="Chaudoir" authorityName="Chaudoir" box="[151,489,1779,1805]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B9337C5C0174FFBCFF68F92FFE5C2212" bold="true" box="[151,489,1779,1805]" pageId="4" pageNumber="19">
<emphasis id="B9337C5C0174FFBCFF68F92FFEDC2212" bold="true" box="[151,361,1779,1805]" italics="true" pageId="4" pageNumber="19">Brachinus piceus</emphasis>
Chaudoir
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF8A04E0174FFBCFF68F8CAFEDB222F" blockId="4.[151,489,1779,1840]" box="[151,366,1814,1840]" pageId="4" pageNumber="19">
(
<figureCitation id="137CBCCB0174FFBCFF60F8CAFF47222F" box="[159,242,1814,1840]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1773,1797]" captionTargetBox="[273,1303,204,1739]" captionTargetId="figure@2.[269,1317,197,1747]" captionTargetPageId="2" captionText="FIGURES 1 4. Habiti of Brachinus ssp. from the Philippines. 1. B. solidipalpis n. sp., paratype, male, 2. B. philippinensis n. sp., holotype, female, 3. B. piceus Chaudior, holotype, male, 4. B. luzonicus Chaudoir, female. Scale bar: 5.0 mm." httpUri="https://zenodo.org/record/178012/files/figure.png" pageId="4" pageNumber="19">Figs. 3</figureCitation>
,
<figureCitation id="137CBCCB0174FFBCFF00F8CAFEBA222F" box="[255,271,1814,1840]" captionStart="FIGURES 5 8" captionStartId="4.[151,269,1584,1608]" captionTargetBox="[195,1416,185,1554]" captionTargetId="figure@4.[151,1436,182,1560]" captionTargetPageId="4" captionText="FIGURES 5 8. Pronota and aedeagi of Brachinus ssp. from the Philippines. 5, 9 10. B. solidipalpis n. sp., holotype, 6. B. philippinensis n. sp., holotype, 7, 11 12. B. piceus Chaudior, holotype, 8, 13 14. B. luzonicus Chaudoir. Scale bar: 5.0 mm for 8, 10.0 mm for others." httpUri="https://zenodo.org/record/178013/files/figure.png" pageId="4" pageNumber="19">7</figureCitation>
, 1112)
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50174FFBCFF68F8BAFC692262" box="[151,988,1894,1917]" pageId="4" pageNumber="19" type="reference_group">
<paragraph id="8BF8A04E0174FFBCFF68F8BAFC692262" blockId="4.[151,988,1894,1917]" box="[151,988,1894,1917]" pageId="4" pageNumber="19">
<treatmentCitationGroup id="AB5787600174FFBCFF68F8BAFC692262" box="[151,988,1894,1917]" pageId="4" pageNumber="19">
<taxonomicName id="4C47DBCD0174FFBCFF68F8BAFD932262" ID-CoL="MTF6" authority="Chaudoir, 1876: 53" authorityName="Chaudoir" authorityPageNumber="53" authorityYear="1876" box="[151,550,1894,1917]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B9337C5C0174FFBCFF68F8BAFEE52262" box="[151,336,1894,1917]" italics="true" pageId="4" pageNumber="19">Brachinus piceus</emphasis>
<treatmentCitation id="0AE6865F0174FFBCFEA9F8BAFD932262" author="Chaudoir" box="[342,550,1894,1917]" page="53" pageId="4" pageNumber="19" year="1876">
<bibRefCitation id="EFD6DDBF0174FFBCFEA9F8BAFE4B2262" author="Chaudoir" box="[342,510,1894,1917]" pageId="4" pageNumber="22" refString="Chaudoir, M. de. (1876) Monographie des Brachynides. Annales de la Socit Entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
: 53
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AE6865F0174FFBCFDCEF8BAFCBE2262" author="Andrewes" box="[561,779,1894,1917]" page="51" pageId="4" pageNumber="19" year="1930">
<bibRefCitation id="EFD6DDBF0174FFBCFDCEF8BAFD512262" author="Andrewes" box="[561,740,1894,1917]" pageId="4" pageNumber="22" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects, 18, Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes, 1930</bibRefCitation>
: 51
</treatmentCitation>
;
<treatmentCitation id="0AE6865F0174FFBCFCE8F8BAFC6C2262" author="Csiki" box="[791,985,1894,1917]" page="1625" pageId="4" pageNumber="19" year="1933">
<bibRefCitation id="EFD6DDBF0174FFBCFCE8F8BAFC202262" author="Csiki" box="[791,917,1894,1917]" pageId="4" pageNumber="22" refString="Csiki, E. (1933) Carabidae III: Harpalinae VIII. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, part 126, Junk, Berlin, pp. 15991933." type="journal article" year="1933">Csiki, 1933</bibRefCitation>
: 1625
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50174FFBDFF68F872FEC526CD" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="diagnosis">
<paragraph id="8BF8A04E0174FFBDFF68F872FF6E25C5" blockId="4.[151,1436,1966,2032]" lastBlockId="5.[151,1437,152,1218]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<emphasis id="B9337C5C0174FFBCFF68F872FEAC22D7" bold="true" box="[151,281,1966,1992]" pageId="4" pageNumber="19">Diagnosis:</emphasis>
Medium sized, dark brown, legs yellow, mouthparts with palpi slender, mentum without pit or furrow, labial palpiger unisetose; head as wide as pronotum, lateral margins of pronotum deeply sinuate before hind angles, proepisternum invisible from above; elytra distinctly carinate, costae much narrower than depressions.
</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FF34FCF2241D" blockId="5.[151,1437,152,1218]" box="[199,839,232,258]" pageId="5" pageNumber="20">
Length: 10.0 mm; width: 4.0 mm. Habitus as in
<figureCitation id="137CBCCB0175FFBDFD04FF34FCF7241D" box="[763,834,232,258]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1773,1797]" captionTargetBox="[273,1303,204,1739]" captionTargetId="figure@2.[269,1317,197,1747]" captionTargetPageId="2" captionText="FIGURES 1 4. Habiti of Brachinus ssp. from the Philippines. 1. B. solidipalpis n. sp., paratype, male, 2. B. philippinensis n. sp., holotype, female, 3. B. piceus Chaudior, holotype, male, 4. B. luzonicus Chaudoir, female. Scale bar: 5.0 mm." httpUri="https://zenodo.org/record/178012/files/figure.png" pageId="5" pageNumber="20">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FECCFC0A2435" blockId="5.[151,1437,152,1218]" box="[199,959,272,298]" pageId="5" pageNumber="20">Whole body black, except yellowish on antennae, legs and palpi.</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FEE4FCC224ED" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">Head longer than wide (HL/HW=1.18); labrum straight, sexsetose, clypeus quadrisetose, the additional two setae on inner portion; mandibular scrobe plurisetose, frons and vertex convex, frontal impressions punctured and setose, compound eyes rather small; palpi subcylindrical, pubescent; ligula dilated, plurisetose, adnated to paraglossae, labial palpomere 2 plurisetose, mentum without tooth and pit, with four setae, submentum with three pairs of setae; labial palpiger with one seta.</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FDDCFC31275D" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">Antennae extending to middle of elytra, antennomere 1 with several long setae, pubescent from antennomere 2; antennomere 3 much longer than the 1 and 2 combined.</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FD8CFF7A27A5" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">
Pronotum quadrate (
<figureCitation id="137CBCCB0175FFBDFE49FD8CFE482775" box="[438,509,592,618]" captionStart="FIGURES 5 8" captionStartId="4.[151,269,1584,1608]" captionTargetBox="[195,1416,185,1554]" captionTargetId="figure@4.[151,1436,182,1560]" captionTargetPageId="4" captionText="FIGURES 5 8. Pronota and aedeagi of Brachinus ssp. from the Philippines. 5, 9 10. B. solidipalpis n. sp., holotype, 6. B. philippinensis n. sp., holotype, 7, 11 12. B. piceus Chaudior, holotype, 8, 13 14. B. luzonicus Chaudoir. Scale bar: 5.0 mm for 8, 10.0 mm for others." httpUri="https://zenodo.org/record/178013/files/figure.png" pageId="5" pageNumber="20">Fig. 7</figureCitation>
), as long as wide, disc moderately convex, lateral margin long and gently sinuate before hind angle, hind angle nearly rectangular, not sharp. Disc wrinkled, sparsely punctured and pubescent.
</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FD14FBB82645" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">Elytra without shoulders, elongate-ovate (EL/EW=1.63), sides not paralleled, widest at about apical onethird; depressions much wider than costae, punctures well marked, pubescent; pubescent distinct on base, edge and apex, but indistinct on median portion; costae distinctly carinate, very narrow and glabrous; apex of elytra truncated, with narrow membrane, fringe distinct, without long setae.</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FCB4FEC526CD" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">Fore tibia with subapical spur located externally; fore tarsomeres 13 of male slightly dilated, sponge setae diagnosed symmetric. Abdominal ventrites densely pubescent except proepisternum sparsely punctured and shortly setose.</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50175FFBDFF38FC03FE1B2155" pageId="5" pageNumber="20" type="description">
<paragraph id="8BF8A04E0175FFBDFF38FC03FD12213D" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">
<emphasis id="B9337C5C0175FFBDFF38FC03FE3126E6" bold="true" box="[199,388,991,1017]" pageId="5" pageNumber="20">Male genitalia:</emphasis>
Median lobe of aedeagus bent ventrally, apical lamella long, slightly constricted at subapex, apex broad; virga absent (Figs. 1112).
</paragraph>
<paragraph id="8BF8A04E0175FFBDFF38FBF3FE1B2155" blockId="5.[151,1437,152,1218]" box="[199,430,1071,1098]" pageId="5" pageNumber="20">
<emphasis id="B9337C5C0175FFBDFF38FBF3FE9E2156" bold="true" box="[199,299,1071,1097]" pageId="5" pageNumber="20">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50175FFBDFF38FB8BFEA32185" pageId="5" pageNumber="20" type="materials_examined">
<paragraph id="8BF8A04E0175FFBDFF38FB8BFEA32185" blockId="5.[151,1437,152,1218]" pageId="5" pageNumber="20">
<emphasis id="B9337C5C0175FFBDFF38FB8BFE2A216E" bold="true" box="[199,415,1111,1137]" pageId="5" pageNumber="20">Material studied:</emphasis>
Known only from the unique male
<typeStatus id="54FC1EEC0175FFBDFCBEFB84FC1F216D" box="[833,938,1112,1138]" pageId="5" pageNumber="20" type="holotype">holotype</typeStatus>
, labeled “
<taxonomicName id="4C47DBCD0175FFBDFBE0FB84FB48216D" authority="Chaudoir, I." authorityName="Chaudoir, I." box="[1055,1277,1112,1138]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B9337C5C0175FFBDFBE0FB84FBDF216E" box="[1055,1130,1112,1137]" italics="true" pageId="5" pageNumber="20">piceus</emphasis>
Chaudoir, I.
</taxonomicName>
<collectingCountry id="F350E0DE0175FFBDFAFBFB84FA3B216D" box="[1284,1422,1112,1138]" name="Philippines" pageId="5" pageNumber="20">Philippines</collectingCountry>
”, in
<collectionCode id="ED56388B0175FFBDFF4AFB5CFEA62185" LSID="urn:lsid:biocol.org:col:34988" box="[181,275,1152,1178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="20">MNHN</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50175FFBDFF38FB7BFD8721DD" box="[199,562,1191,1218]" pageId="5" pageNumber="20" type="distribution">
<paragraph id="8BF8A04E0175FFBDFF38FB7BFD8721DD" blockId="5.[151,1437,152,1218]" box="[199,562,1191,1218]" pageId="5" pageNumber="20">
<emphasis id="B9337C5C0175FFBDFF38FB7BFEDF21DE" bold="true" box="[199,362,1191,1217]" pageId="5" pageNumber="20">Distribution:</emphasis>
The
<collectingCountry id="F350E0DE0175FFBDFE59FB74FD9B21DD" box="[422,558,1192,1218]" name="Philippines" pageId="5" pageNumber="20">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>