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<mods:title id="AA4989C08C4BC81676EE4A3C845998F2">Revision of the higher taxonomy of Neotropical freshwater crabs of the family Pseudothelphusidae, based on multigene and morphological analyses</mods:title>
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<paragraph id="8BF8BB67FFE0FFBEFF77FC6BC5A72425" blockId="16.[163,742,985,1039]" lastBlockId="16.[145,761,1077,1130]" pageId="16" pageNumber="989">
SUBFAMILY
<taxonomicName id="4C47C0E4FFE0FFBEFEFEFC6BC6D423BD" ID-CoL="8H85S" authority="PRETZMANN, 1971" authorityName="Pretzmann" authorityYear="1971" box="[298,736,985,1010]" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" rank="subFamily" subFamily="Hypolobocerinae">
HYPOLOBOCERINAE
<bibRefCitation id="EFD6C696FFE0FFBEFDC4FC6BC6D423BD" author="Pretzmann G" box="[528,736,985,1010]" pageId="16" pageNumber="989" refId="ref21558" refString="Pretzmann G. 1971. Fortschritte in der Klassifizierung der Pseudothelphusidae. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften," type="book" year="1971">PRETZMANN, 1971</bibRefCitation>
</taxonomicName>
, NEW RANK
<emphasis id="B9336775FFE0FFBEFF45FB87C48A2405" box="[145,190,1077,1098]" italics="true" pageId="16" pageNumber="989">lsid</emphasis>
:
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="C35DE8ECFFE0FFBEFF45FB3AC1172094" pageId="16" pageNumber="989" type="diagnosis">
<paragraph id="8BF8BB67FFE0FFBEFF45FB3AC4C32710" blockId="16.[145,762,1160,1887]" pageId="16" pageNumber="989">
<emphasis id="B9336775FFE0FFBEFF45FB3AC4ED24F3" italics="true" pageId="16" pageNumber="989">
Diagnosis (modified from
<bibRefCitation id="EFD6C696FFE0FFBEFE13FB3AC6B224D2" author="Rodriguez G" box="[455,646,1160,1181]" pageId="16" pageNumber="989" pagination="224" refId="ref21823" refString="Rodriguez G. 1982. Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Collection Faune Tropicale 22. Paris: Editions ORSTOM, 224." type="journal article" year="1982">Rodriguez, 1982</bibRefCitation>
;
<bibRefCitation id="EFD6C696FFE0FFBEFD42FB3AC4FD24F3" author="Campos MR" pageId="16" pageNumber="989" refId="ref19132" refString="Campos MR. 2005. Freshwater crabs from Colombia: a taxonomic and distributional study. Coleccion Jorge Alvarez Lleras No. 24. Bogota: Academia Colombiana de Ciencias Exactas, Fisicas y Naturales." type="book" year="2005">Campos, 2005</bibRefCitation>
):
</emphasis>
Anterolateral margin of carapace often finely serrate to unarmed, lateral armature indistinct. Third maxilliped with exopod of variable length, not reaching distal border of ischium, ratio exopod/ischium length 0.20.8. Branchial efferent channel open, gap between jugal and lateral angles in
<taxonomicName id="4C47C0E4FFE0FFBEFE1FFAF2C60D251A" box="[459,569,1344,1365]" class="Malacostraca" family="Pseudothelphusidae" genus="Martiana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFE1FFAF2C60D251A" box="[459,569,1344,1365]" italics="true" pageId="16" pageNumber="989">Martiana</emphasis>
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partially closed. Gonopod morphology variable, with one lateral lobe of variable size and form (reduced in
<taxonomicName id="4C47C0E4FFE0FFBEFD8DFACCC6FE25DC" box="[601,714,1406,1427]" class="Malacostraca" family="Pseudothelphusidae" genus="Martiana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFD8DFACCC6FE25DC" box="[601,714,1406,1427]" italics="true" pageId="16" pageNumber="989">Martiana</emphasis>
</taxonomicName>
) on principal axis, in
<taxonomicName id="4C47C0E4FFE0FFBEFE8CFA2EC5C225FE" box="[344,502,1436,1457]" class="Malacostraca" family="Pseudothelphusidae" genus="Neostrengeria" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFE8CFA2EC5C225FE" box="[344,502,1436,1457]" italics="true" pageId="16" pageNumber="989">Neostrengeria</emphasis>
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with accessory lateral lobe. Gonopod somewhat slender, truncated and simple distally, caudal ridge strong, well defined, lateral lobe on principal axis evident, triangular, rounded or subquadrate, sometimes divided creating accessory lobe. In distal view, apical cavity transversal to main axis, rounded, oval or elongated caudocephalically: with two apical prominences (papillae), conical, acute, located on caudal margin of spermatic channel; aperture of spermatic channel in central position; mesial process small or of moderate size, subdistal, triangular, cephalically projected. Apex with apical elements, rarely ornamented with spines, spinules or granules;
<taxonomicName id="4C47C0E4FFE0FFBEFED5F898C5902770" box="[257,420,1834,1855]" class="Malacostraca" family="Pseudothelphusidae" genus="Lindacatalina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFED5F898C5902770" box="[257,420,1834,1855]" italics="true" pageId="16" pageNumber="989">Lindacatalina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C47C0E4FFE0FFBEFE0FF898C66D2770" authorityName="Pretzmann" authorityYear="1965" box="[475,601,1834,1855]" class="Malacostraca" family="Pseudothelphusidae" genus="Moritschus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFE0FF898C66D2770" box="[475,601,1834,1855]" italics="true" pageId="16" pageNumber="989">Moritschus</emphasis>
</taxonomicName>
could present spinules.
</paragraph>
<paragraph id="8BF8BB67FFE0FFBEFCFDFF74C1172094" blockId="16.[809,1315,197,219]" box="[809,1315,197,219]" pageId="16" pageNumber="989">
<emphasis id="B9336775FFE0FFBEFCFDFF74C7992094" box="[809,941,198,219]" italics="true" pageId="16" pageNumber="989">
<typeStatus id="54FC05C5FFE0FFBEFCFDFF74C76A2094" box="[809,862,198,219]" pageId="16" pageNumber="989">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="4C47C0E4FFE0FFBEFC6DFF77C12B2094" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" box="[953,1311,197,219]" class="Malacostraca" family="Pseudothelphusidae" genus="Hypolobocera" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFC6DFF77C0662095" box="[953,1106,197,218]" italics="true" pageId="16" pageNumber="989">Hypolobocera</emphasis>
Pretzmann, 1965
</taxonomicName>
.
</paragraph>
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<subSubSection id="C35DE8ECFFE0FFBEFCFDFEB6C123249E" pageId="16" pageNumber="989" type="discussion">
<paragraph id="8BF8BB67FFE0FFBEFCFDFEB6C7C721DB" blockId="16.[809,1425,260,404]" pageId="16" pageNumber="989">
<emphasis id="B9336775FFE0FFBEFCFDFEB6C0352156" box="[809,1025,260,281]" italics="true" pageId="16" pageNumber="989">Genera included:</emphasis>
<taxonomicName id="4C47C0E4FFE0FFBEFBC4FEB6C7CB2178" authority="Campos &amp; Magalhaes, 2014" authorityName="Campos &amp; Magalhaes" authorityYear="2014" class="Malacostraca" family="Pseudothelphusidae" genus="Colombiathelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFBC4FEB6C1372156" box="[1040,1283,260,281]" italics="true" pageId="16" pageNumber="989">Colombiathelphusa</emphasis>
<bibRefCitation id="EFD6C696FFE0FFBEFADBFEB6C7CB2178" author="Campos MR &amp; Magalhaes C" pageId="16" pageNumber="989" pagination="571 - 579" refId="ref19188" refString="Campos MR, Magalhaes C. 2014. Colombiathelphusa, a new genus of freshwater crab from Colombia, and the first location record of Eidocamptophallus chacei (Pretzmann, 1967) (Crustacea: Decapoda: Pseudothelphusidae). Zootaxa 3860: 571 - 579." type="journal article" year="2014">Campos &amp; Magalhães, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C47C0E4FFE0FFBEFBC7FE90C1BF2178" authority="Ortmann, 1897" authorityName="Ortmann" authorityYear="1897" box="[1043,1419,290,311]" class="Malacostraca" family="Pseudothelphusidae" genus="Hypolobocera" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFBC7FE90C0882178" box="[1043,1212,290,311]" italics="true" pageId="16" pageNumber="989">Hypolobocera</emphasis>
Ortmann, 1897
</taxonomicName>
,
<taxonomicName id="4C47C0E4FFE0FFBEFCFDFEF3C0AC2119" authority="Pretzmann, 1977" authorityName="Pretzmann" authorityYear="1977" box="[809,1176,321,342]" class="Malacostraca" family="Pseudothelphusidae" genus="Lindacatalina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFCFDFEF3C7F82119" box="[809,972,321,342]" italics="true" pageId="16" pageNumber="989">Lindacatalina</emphasis>
Pretzmann, 1977
</taxonomicName>
,
<taxonomicName id="4C47C0E4FFE0FFBEFB77FEF0C756213A" authority="Rodriguez, 1980" authorityName="Rodriguez" authorityYear="1980" class="Malacostraca" family="Pseudothelphusidae" genus="Martiana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFB77FEF0C1242118" box="[1187,1296,322,343]" italics="true" pageId="16" pageNumber="989">Martiana</emphasis>
Rodriguez, 1980
</taxonomicName>
,
<taxonomicName id="4C47C0E4FFE0FFBEFCB9FED2C08C2139" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" box="[877,1208,352,374]" class="Malacostraca" family="Pseudothelphusidae" genus="Moritschus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFCB9FED2C7D8213A" box="[877,1004,352,373]" italics="true" pageId="16" pageNumber="989">Moritschus</emphasis>
Pretzmann, 1965
</taxonomicName>
and
<taxonomicName id="4C47C0E4FFE0FFBEFB26FED2C7DB21DB" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" class="Malacostraca" family="Pseudothelphusidae" genus="Neostrengeria" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFB26FED2C1A5213A" box="[1266,1425,352,373]" italics="true" pageId="16" pageNumber="989">Neostrengeria</emphasis>
Pretzmann, 1965
</taxonomicName>
.
</paragraph>
<paragraph id="8BF8BB67FFE0FFBEFCFDFE0FC0DE232E" blockId="16.[809,1426,445,1233]" pageId="16" pageNumber="989">
<emphasis id="B9336775FFE0FFBEFCFDFE0FC7A3219D" box="[809,919,445,466]" italics="true" pageId="16" pageNumber="989">Remarks:</emphasis>
The former tribe
<taxonomicName id="4C47C0E4FFE0FFBEFBA6FE0FC111219C" baseAuthorityName="Campos &amp; Magalhaes" baseAuthorityYear="2014" box="[1138,1317,445,467]" pageId="16" pageNumber="989" rank="tribe" tribe="Hypolobocerini">Hypolobocerini</taxonomicName>
, now the subfamily
<taxonomicName id="4C47C0E4FFE0FFBEFC4AFE6EC06321BD" authorityName="Pretzmann" authorityYear="1971" box="[926,1111,476,498]" pageId="16" pageNumber="989" rank="subFamily" subFamily="Hypolobocerinae">Hypolobocerinae</taxonomicName>
was one of the most diverse and widely distributed groups of American freshwater crabs, ranging from southern
<collectingCountry id="F350FBF7FFE0FFBEFBAFFDABC0F82260" box="[1147,1228,537,559]" name="Mexico" pageId="16" pageNumber="989">Mexico</collectingCountry>
, through Central America, to northern SouthAmerica, including
<collectingCountry id="F350FBF7FFE0FFBEFAF6FD8AC1B92201" box="[1314,1421,568,590]" name="Colombia" pageId="16" pageNumber="989">Colombia</collectingCountry>
,
<collectingCountry id="F350FBF7FFE0FFBEFCFDFDE4C7B12223" box="[809,901,598,620]" name="Ecuador" pageId="16" pageNumber="989">Ecuador</collectingCountry>
,
<collectingCountry id="F350FBF7FFE0FFBEFC5BFDE5C7D82223" box="[911,1004,599,620]" name="Panama" pageId="16" pageNumber="989">Panama</collectingCountry>
,
<collectingCountry id="F350FBF7FFE0FFBEFC22FDE5C01F2223" box="[1014,1067,599,620]" name="Peru" pageId="16" pageNumber="989">Peru</collectingCountry>
and
<collectingCountry id="F350FBF7FFE0FFBEFBB5FDE4C0E12223" box="[1121,1237,598,620]" name="Venezuela" pageId="16" pageNumber="989">Venezuela</collectingCountry>
. Representatives of this group show a great variety of morphologies of the male gonopod, from the typical pattern of the genus
<taxonomicName id="4C47C0E4FFE0FFBEFCFDFD00C7F52288" authorityName="Ortmann" authorityYear="1897" box="[809,961,690,711]" class="Malacostraca" family="Pseudothelphusidae" genus="Hypolobocera" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFCFDFD00C7F52288" box="[809,961,690,711]" italics="true" pageId="16" pageNumber="989">Hypolobocera</emphasis>
</taxonomicName>
to the ornamented ones of
<taxonomicName id="4C47C0E4FFE0FFBEFB2CFD00C1BE2288" box="[1272,1418,690,711]" class="Malacostraca" family="Pseudothelphusidae" genus="Elsalvadoria" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFB2CFD00C1BE2288" box="[1272,1418,690,711]" italics="true" pageId="16" pageNumber="989">Elsalvadoria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C47C0E4FFE0FFBEFCFDFD63C04B22A9" authority=", Spirothelpusa" authorityName="Spirothelpusa" box="[809,1151,721,742]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFCFDFD63C7F922A9" box="[809,973,721,742]" italics="true" pageId="16" pageNumber="989">Ptychophallus</emphasis>
,
<emphasis id="B9336775FFE0FFBEFC08FD63C04B22A9" box="[988,1151,721,742]" italics="true" pageId="16" pageNumber="989">Spirothelpusa</emphasis>
</taxonomicName>
and others. In the new taxonomic scheme this taxon is restricted to the typical morphology of
<taxonomicName id="4C47C0E4FFE0FFBEFC1FFCBCC054236C" authorityName="Ortmann" authorityYear="1897" box="[971,1120,782,803]" class="Malacostraca" family="Pseudothelphusidae" genus="Hypolobocera" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFC1FFCBCC054236C" box="[971,1120,782,803]" italics="true" pageId="16" pageNumber="989">Hypolobocera</emphasis>
</taxonomicName>
and allied genera (
<tableCitation id="C6C58EDCFFE0FFBEFAE7FCBCC1B7236C" box="[1331,1411,782,804]" captionStart="Table 4" captionStartId="11.[143,207,552,574]" captionTargetPageId="11" captionText="Table 4. Proposal of the new classification of genera, families and subfamilies of the superfamily Pseudothelphusiodea. Abbreviations for countries of occurrence for each genus are: BE, Belize; BR, Brazil; CU, Cuba; COL, Colombia; CR, Costa Rica; DOM, Dominica; DR, Dominican Republic; EC, Ecuador; FGU, French Guiana; GUA, Guadeloupe; GT, Guatemala; GUY, Guyana; HA, Haiti; HON, Honduras; MX, Mexico; MAR, Martinique; NI, Nicaragua; PAN, Panama; PE, Peru; PR, Puerto Rico; STV, Saint Vicent; SAL, El Salvador; SC, Saint Croix, US Virgin Islands; SLU, Saint Lucia; SUR, Suriname; TR, Trinidad and Tobago; VE, Venezuela" httpUri="http://table.plazi.org/id/DF38EBEFFFFBFFA5FF5BFD9AC4CE22FC" pageId="16" pageNumber="989" tableUuid="DF38EBEFFFFBFFA5FF5BFD9AC4CE22FC">Table 4</tableCitation>
), and geographically occurring from south-western
<collectingCountry id="F350FBF7FFE0FFBEFCFDFCFEC7B1232E" box="[809,901,844,865]" name="Panama" pageId="16" pageNumber="989">Panama</collectingCountry>
to
<collectingCountry id="F350FBF7FFE0FFBEFC72FCFEC025232D" box="[934,1041,844,866]" name="Colombia" pageId="16" pageNumber="989">Colombia</collectingCountry>
,
<collectingCountry id="F350FBF7FFE0FFBEFBCFFCFEC04C232D" box="[1051,1144,844,866]" name="Ecuador" pageId="16" pageNumber="989">Ecuador</collectingCountry>
and
<collectingCountry id="F350FBF7FFE0FFBEFB7AFCFEC0D2232E" box="[1198,1254,844,865]" name="Peru" pageId="16" pageNumber="989">Peru</collectingCountry>
.
</paragraph>
<paragraph id="8BF8BB67FFE0FFBEFC95FCD8C123249E" blockId="16.[809,1426,445,1233]" pageId="16" pageNumber="989">
Although no DNA sequences were available for
<taxonomicName id="4C47C0E4FFE0FFBEFCFDFC38C7AD23D0" authorityName="Rodriguez" authorityYear="1980" box="[809,921,906,927]" class="Malacostraca" family="Pseudothelphusidae" genus="Martiana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFCFDFC38C7AD23D0" box="[809,921,906,927]" italics="true" pageId="16" pageNumber="989">Martiana</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD6C696FFE0FFBEFC7DFC3BC07923D0" author="Campos MR" box="[937,1101,905,927]" pageId="16" pageNumber="989" refId="ref19132" refString="Campos MR. 2005. Freshwater crabs from Colombia: a taxonomic and distributional study. Coleccion Jorge Alvarez Lleras No. 24. Bogota: Academia Colombiana de Ciencias Exactas, Fisicas y Naturales." type="book" year="2005">Campos, 2005</bibRefCitation>
: fig. 42) and
<taxonomicName id="4C47C0E4FFE0FFBEFB3DFC3BC1A523D1" authorityName="Pretzmann" authorityYear="1977" box="[1257,1425,905,926]" class="Malacostraca" family="Pseudothelphusidae" genus="Lindacatalina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFB3DFC3BC1A523D1" box="[1257,1425,905,926]" italics="true" pageId="16" pageNumber="989">Lindacatalina</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD6C696FFE0FFBEFCE5FC1AC7F923F2" author="Campos MR" box="[817,973,935,957]" pageId="16" pageNumber="989" refId="ref19132" refString="Campos MR. 2005. Freshwater crabs from Colombia: a taxonomic and distributional study. Coleccion Jorge Alvarez Lleras No. 24. Bogota: Academia Colombiana de Ciencias Exactas, Fisicas y Naturales." type="book" year="2005">Campos, 2005</bibRefCitation>
: figs 9194), these genera were placed in this subfamily due to morphological characters of their male first gonopod.
<taxonomicName id="4C47C0E4FFE0FFBEFBDFFC57C0DD23B5" authorityName="Campos &amp; Magalhaes" authorityYear="2014" box="[1035,1257,997,1018]" class="Malacostraca" family="Pseudothelphusidae" genus="Colombiathelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFBDFFC57C0DD23B5" box="[1035,1257,997,1018]" italics="true" pageId="16" pageNumber="989">Colombiathelphusa</emphasis>
</taxonomicName>
was originally placed in the
<taxonomicName id="4C47C0E4FFE0FFBEFC03FBB6C0A42455" baseAuthorityName="Campos &amp; Magalhaes" baseAuthorityYear="2014" box="[983,1168,1028,1050]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="994" phylum="Arthropoda" rank="tribe" tribe="Hypolobocerini">Hypolobocerini</taxonomicName>
recognizing that its gonopod morphology shows similarities with other groups (
<bibRefCitation id="EFD6C696FFE0FFBEFC52FBF3C0F32419" author="Campos MR &amp; Magalhaes C" box="[902,1223,1089,1111]" pageId="16" pageNumber="989" pagination="571 - 579" refId="ref19188" refString="Campos MR, Magalhaes C. 2014. Colombiathelphusa, a new genus of freshwater crab from Colombia, and the first location record of Eidocamptophallus chacei (Pretzmann, 1967) (Crustacea: Decapoda: Pseudothelphusidae). Zootaxa 3860: 571 - 579." type="journal article" year="2014">Campos &amp; Magalhães, 2014</bibRefCitation>
); future analyses should revisit the appropriateness of this placement. The affinity of the genus
<taxonomicName id="4C47C0E4FFE0FFBEFB99FBCCC09924DC" box="[1101,1197,1150,1171]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
<emphasis id="B9336775FFE0FFBEFB99FBCCC09924DC" box="[1101,1197,1150,1171]" italics="true" pageId="16" pageNumber="989">Chaceus</emphasis>
</taxonomicName>
needs to be further analysed to define if it belongs in the
<taxonomicName id="4C47C0E4FFE0FFBEFB07FB2FC1A524FC" authorityName="Pretzmann" authorityYear="1971" box="[1235,1425,1181,1203]" pageId="16" pageNumber="995" rank="subFamily" subFamily="Hypolobocerinae">Hypolobocerinae</taxonomicName>
or as proposed here in the
<taxonomicName id="4C47C0E4FFE0FFBEFB83FB09C127249E" authorityName="Rodriguez" authorityYear="1982" box="[1111,1299,1211,1233]" pageId="16" pageNumber="996" rank="subFamily" subFamily="Strengerianinae">Strengerianinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>