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<document id="D5E65BA3D72BED451581E30BBE178D98" ID-CLB-Dataset="7385" ID-DOI="10.11646/zootaxa.4951.3.5" ID-GBIF-Dataset="caee27e5-0786-4e95-84fb-ee5394fa79e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668365" ID-ZooBank="9BEB98F0-DEA7-4131-8E37-DFC4D5DFCB7B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617780432061" checkinUser="plazi" docAuthor="Bogorodsky, Sergey V., Zajonz, Uwe, Saeed, Fouad N. &amp; Weigmann, Simon" docDate="2021" docId="03B851366649FFA8FF6C14EBE47DEFD2" docLanguage="en" docName="zootaxa.4951.3.5.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pastinachus sephen" docType="treatment" docVersion="7" lastPageNumber="524" masterDocId="FF81294E6645FFA5FFFB1531E100ED7B" masterDocTitle="Notes on batoid fishes of the Socotra Archipelago (north-western Indian Ocean) with four new records" masterLastPageNumber="528" masterPageNumber="511" pageNumber="523" updateTime="1698936112385" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EF00ED508A3075BA0F1297333DA269B3">Notes on batoid fishes of the Socotra Archipelago (north-western Indian Ocean) with four new records</mods:title>
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<mods:namePart id="725B417015BD8389CA2D496B660AD3E7">Bogorodsky, Sergey V.</mods:namePart>
<mods:affiliation id="4BCE0BB25D016DA64499A08A27E85340">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK &amp; S ṫaṫẚon of ka ṫuralẚsṫsI OmskI ouss ẚaK ẚc 18 T 19 S @ yandexKru; h ṫṫps: LLorc ẚdKorgL 0000 - 0002 - 8679 - 9735</mods:affiliation>
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<mods:namePart id="BE7A3017E9F37D76B08E3A1589F06FF8">Zajonz, Uwe</mods:namePart>
<mods:affiliation id="2366BAEDF45DBF8786D1169E580AF8EA">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK</mods:affiliation>
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<mods:namePart id="ECE81A37A8A06558EBD1041F4B836397">Saeed, Fouad N.</mods:namePart>
<mods:affiliation id="FB51925D04971E646963E484672976B4">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK</mods:affiliation>
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<mods:namePart id="E068AB9F9D8737CD690038323194A2A2">Weigmann, Simon</mods:namePart>
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<mods:date id="C28A79961C22FA5C242E83CCDE129F29">2021</mods:date>
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<mods:number id="012B22EFFE0AC6FB2EEB89E1D9B63B59">4951</mods:number>
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<treatment id="03B851366649FFA8FF6C14EBE47DEFD2" ID-DOI="http://doi.org/10.5281/zenodo.4684879" ID-GBIF-Taxon="181003431" ID-Zenodo-Dep="4684879" LSID="urn:lsid:plazi:treatment:03B851366649FFA8FF6C14EBE47DEFD2" httpUri="http://treatment.plazi.org/id/03B851366649FFA8FF6C14EBE47DEFD2" lastPageId="13" lastPageNumber="524" pageId="12" pageNumber="523">
<subSubSection id="C30BB3AB6649FFA9FF6C14EBE0D6EF62" pageId="12" pageNumber="523" type="nomenclature">
<paragraph id="8BAEE0206649FFA9FF6C14EBE363EC8F" blockId="12.[151,611,474,573]" box="[151,611,474,501]" pageId="12" pageNumber="523">
<heading id="D0E6574C6649FFA9FF6C14EBE363EC8F" bold="true" box="[151,611,474,501]" fontSize="11" level="1" pageId="12" pageNumber="523" reason="1">
<taxonomicName id="4C119BA36649FFA9FF6C14EBE363EC8F" ID-CoL="4DZC8" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[151,611,474,501]" class="Elasmobranchii" family="Dasyatidae" genus="Pastinachus" kingdom="Animalia" order="Myliobatiformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="sephen">
<emphasis id="B9653C326649FFA9FF6C14EBE363EC8F" bold="true" box="[151,611,474,501]" pageId="12" pageNumber="523">
<emphasis id="B9653C326649FFA9FF6C14EBE088EC8E" bold="true" box="[151,392,474,501]" italics="true" pageId="12" pageNumber="523">Pastinachus sephen</emphasis>
(
<bibRefCitation id="EF809DD16649FFA9FE6C14EBE35AEC8F" author="Fabricius, J. C." box="[407,602,474,500]" pageId="12" pageNumber="523" refId="ref9562" refString="Fabricius, J. C. (1775) oaja sephen. fn: Niebuhr, C. (Ed.), aescr ẚptẚones an ẚmalẚum av ẚumI amph ẚbẚorumI p ẚscẚumI ẚnsectorumI verm ẚum; quae ẚn ẚtẚnere or ẚentalẚ observav ẚtKKK mos t mor tem auc torẚs ed ẚdẚt Cars ten k ẚebuhrK ex officina Molleri, Hauniae, pp. 17 + viii. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Fabricius, 1775</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAEE0206649FFA9FF6C14CEE0D6EF62" blockId="12.[151,611,474,573]" box="[151,470,511,537]" pageId="12" pageNumber="523">Fantail stingray, Cowtail ray</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB6649FFA9FF6C1712E075EFE5" pageId="12" pageNumber="523" type="description">
<paragraph id="8BAEE0206649FFA9FF6C1712E1E7EF46" blockId="12.[151,611,474,573]" box="[151,231,547,573]" pageId="12" pageNumber="523">
<figureCitation id="132AFCA56649FFA9FF6C1712E1E7EF46" box="[151,231,547,573]" captionStart="FIGURE 10" captionStartId="12.[151,250,1986,2010]" captionTargetBox="[151,1437,1126,1962]" captionTargetId="figure-317@12.[151,1437,1126,1962]" captionTargetPageId="12" captionText="FIGURE 10. masṫẚnachus sephen, underwater photograph, Ras Bidou, Socotra Island, 14 m depth. Photo by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.4668399" httpUri="https://zenodo.org/record/4668399/files/figure.png" pageId="12" pageNumber="523">Fig. 10</figureCitation>
</paragraph>
<paragraph id="8BAEE0206649FFA9FF6C1759E075EFE5" blockId="12.[151,1436,615,670]" pageId="12" pageNumber="523">
<emphasis id="B9653C326649FFA9FF6C1759E1C7EF04" box="[151,199,616,639]" italics="true" pageId="12" pageNumber="523">oaja</emphasis>
<taxonomicName id="4C119BA36649FFA9FF361756E076EF04" authority="Fabricius" authorityName="Fabricius" box="[205,374,615,639]" class="Elasmobranchii" family="Dasyatidae" genus="Pastinachus" kingdom="Animalia" order="Myliobatiformes" pageId="12" pageNumber="523" phylum="Chordata" rank="species" species="sephen">
<emphasis id="B9653C326649FFA9FF361756E013EF04" box="[205,275,615,639]" italics="true" pageId="12" pageNumber="523">sephen</emphasis>
<materialsCitation id="3B79EA7D6649FFA9FEE31756E076EF04" ID-GBIF-Occurrence="3071284313" box="[280,374,615,639]" country="Saudi Arabia" location="Jeddah" municipality="Al-Luhayya" pageId="12" pageNumber="523" specimenCount="2">Fabricius</materialsCitation>
</taxonomicName>
in Niebuhr (ex Forsskål), 1775: 17; no
<typeStatus id="54AA5E826649FFA9FCF91759E237EFFB" box="[770,823,616,640]" pageId="12" pageNumber="523">types</typeStatus>
known;
<typeStatus id="54AA5E826649FFA9FC741759E2BBEFFB" box="[911,955,616,640]" pageId="12" pageNumber="523">type</typeStatus>
locality:
<location id="8ECEB6FB6649FFA9FBE31756E561EF04" LSID="urn:lsid:plazi:treatment:03B851366649FFA8FF6C14EBE47DEFD2:8ECEB6FB6649FFA9FBE31756E561EF04" box="[1048,1121,615,639]" country="Saudi Arabia" municipality="Al-Luhayya" name="Jeddah" pageId="12" pageNumber="523">Jeddah</location>
,
<collectingCountry id="F306A0B06649FFA9FB911756E5EDEF04" box="[1130,1261,615,639]" name="Saudi Arabia" pageId="12" pageNumber="523">Saudi Arabia</collectingCountry>
and
<collectingMunicipality id="6BCA7A5A6649FFA9FAE01756E498EF04" box="[1307,1432,615,639]" pageId="12" pageNumber="523">Al-Luhayya</collectingMunicipality>
,
<collectingCountry id="F306A0B06649FFA9FF3C17B7E010EFE6" box="[199,272,646,669]" name="Yemen" pageId="12" pageNumber="523">Yemen</collectingCountry>
, Red Sea.
</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB6649FFA9FF6C17FDE37FE934" pageId="12" pageNumber="523" type="materials_examined">
<paragraph id="8BAEE0206649FFA9FF6C17FDE019EE71" blockId="12.[151,1437,716,1103]" pageId="12" pageNumber="523">
<emphasis id="B9653C326649FFA9FF6C17FDE085EF9D" bold="true" box="[151,389,716,742]" pageId="12" pageNumber="523">Material examined:</emphasis>
<materialsCitation id="3B79EA7D6649FFA9FE7617FCE015EE71" ID-GBIF-Occurrence="3071284309" collectingCountry="Yemen" collectingDate="2019-04-18" country="Socotra Island" county="Ras Bidou" elevation="14" location="No" municipality="Socotra Island" pageId="12" pageNumber="523" specimenCount="1">
<location id="8ECEB6FB6649FFA9FE7617FCE0AFEF9C" LSID="urn:lsid:plazi:treatment:03B851366649FFA8FF6C14EBE47DEFD2:8ECEB6FB6649FFA9FE7617FCE0AFEF9C" box="[397,431,717,743]" country="Socotra Island" county="Ras Bidou" municipality="Socotra Island" name="No" pageId="12" pageNumber="523">No</location>
specimen collected, underwater photograph,
<collectingCountry id="F306A0B06649FFA9FC4D17FDE55BEF9D" box="[950,1115,716,742]" name="Socotra Island" pageId="12" pageNumber="523">
<collectingMunicipality id="6BCA7A5A6649FFA9FC4D17FDE55BEF9D" box="[950,1115,716,742]" pageId="12" pageNumber="523">Socotra Island</collectingMunicipality>
</collectingCountry>
,
<collectingCounty id="62CF98AC6649FFA9FB9317FCE5E2EF9D" box="[1128,1250,716,743]" pageId="12" pageNumber="523">Ras Bidou</collectingCounty>
,
<elevation id="003C07136649FFA9FB1417FDE473EF9D" box="[1263,1395,716,743]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="523" unit="m" value="14.0">
<quantity id="4CE94DC56649FFA9FB1417FDE42AEF9C" box="[1263,1322,716,743]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="523" unit="m" value="14.0">14 m</quantity>
depth
</elevation>
,
<date id="FFAFC6E06649FFA9FA7B17FDE015EE71" pageId="12" pageNumber="523" value="2019-04-18">
<collectingDate id="EFEB3F086649FFA9FA7B17FDE015EE71" pageId="12" pageNumber="523" value="2019-04-18">18 April 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAEE0206649FFA9FF3C1625E2FCEE98" blockId="12.[151,1437,716,1103]" pageId="12" pageNumber="523">
<emphasis id="B9653C326649FFA9FF3C1625E0D7EE55" bold="true" box="[199,471,788,814]" pageId="12" pageNumber="523">Distinctive characters:</emphasis>
<emphasis id="B9653C326649FFA9FE1B1624E3C0EE55" box="[480,704,788,816]" italics="true" pageId="12" pageNumber="523">masṫẚnachus sephen</emphasis>
is characterised by having a broadly rhomboidal disc, its width 1.21.3 times its length; eyes small, length of orbit and spiracle
<quantity id="4CE94DC56649FFA9FC8C1609E2EAEE29" box="[887,1002,824,850]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="12" pageNumber="523" unit="in" value="1.7" valueMax="1.8" valueMin="1.6">1.61.8 in</quantity>
snout length; 2 or 3 large, pale, seedshaped tubercles mid-centrally on disc (merging with denticle band in adults); pelvic fins large, tips narrowly rounded; tail length usually less than twice disc width, with base broad and depressed; tail with a very deep cutaneous ventral fold, no dorsal fold, tip of tail usually free of fold and tapering continuously to terminal filament; usually one caudal sting well back on tail (Psomadakis
<emphasis id="B9653C326649FFA9FD5016F6E3DCEE99" box="[683,732,967,994]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2015; Last
<emphasis id="B9653C326649FFA9FC9316F6E299EE99" box="[872,921,967,994]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2016c).
</paragraph>
<paragraph id="8BAEE0206649FFA9FF3C16DCE37FE934" blockId="12.[151,1437,716,1103]" pageId="12" pageNumber="523">
<emphasis id="B9653C326649FFA9FF3C16DCE050E973" box="[199,336,1005,1032]" italics="true" pageId="12" pageNumber="523">Colouraṫẚon</emphasis>
: Dorsal disc uniform yellowish-brown, greyish or greyish brown, orbit and spiracle blackish, membranous ventral tail fold and posterior tail dark grey to black. Ventral surface white, often with narrow black margins (Psomadakis
<emphasis id="B9653C326649FFA9FED61102E05FE935" box="[301,351,1075,1102]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2015; Last
<emphasis id="B9653C326649FFA9FE111102E31BE935" box="[490,539,1075,1102]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2016c).
</paragraph>
</subSubSection>
<caption id="DF6EB0A86649FFA9FF6C12F3E49AEAA1" ID-DOI="http://doi.org/10.5281/zenodo.4668399" ID-Zenodo-Dep="4668399" box="[151,1434,1985,2010]" httpUri="https://zenodo.org/record/4668399/files/figure.png" pageId="12" pageNumber="523" startId="12.[151,250,1986,2010]" targetBox="[151,1437,1126,1962]" targetPageId="12">
<paragraph id="8BAEE0206649FFA9FF6C12F3E49AEAA1" blockId="12.[151,1434,1985,2010]" box="[151,1434,1985,2010]" pageId="12" pageNumber="523">
<emphasis id="B9653C326649FFA9FF6C12F3E020EAA1" bold="true" box="[151,288,1986,2010]" pageId="12" pageNumber="523">FIGURE 10.</emphasis>
<emphasis id="B9653C326649FFA9FEDD12F0E0EEEAA1" box="[294,494,1985,2010]" italics="true" pageId="12" pageNumber="523">masṫẚnachus sephen</emphasis>
, underwater photograph, Ras Bidou, Socotra Island, 14 m depth. Photo by S.V. Bogorodsky.
</paragraph>
</caption>
<subSubSection id="C30BB3AB6648FFA8FF3C15A7E387EC66" pageId="13" pageNumber="524" type="distribution">
<paragraph id="8BAEE0206648FFA8FF3C15A7E387EC66" blockId="13.[151,1437,150,681]" pageId="13" pageNumber="524">
<emphasis id="B9653C326648FFA8FF3C15A7E063EDCB" bold="true" box="[199,355,150,176]" pageId="13" pageNumber="524">Distribution:</emphasis>
The exact species distribution is unknown, positive records in the wider Arabian region include the Red Sea from its
<typeStatus id="54AA5E826648FFA8FE7C158AE0B7EDAE" box="[391,439,187,213]" pageId="13" pageNumber="524">type</typeStatus>
locality (
<bibRefCitation id="EF809DD16648FFA8FDD8158BE263EDAF" author="Golani, D. &amp; Bogorodsky, S. V." box="[547,867,186,212]" pageId="13" pageNumber="524" pagination="1 - 135" refId="ref10068" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The fishes of the Red Sea - reappraisal and updated checklist. Zoo taxa, 2463 (1), 1 - 135. https: // doi. org / 10.11646 / zootaxa. 2463.1.1" type="journal article" year="2010">Golani &amp; Bogorodsky 2010</bibRefCitation>
), Gulf of Tadjoura (Lips
<emphasis id="B9653C326648FFA8FB84158BE5B1EDAF" box="[1151,1201,186,213]" italics="true" pageId="13" pageNumber="524">eṫ al</emphasis>
. 2016),
<collectingCountry id="F306A0B06648FFA8FAEB158BE454EDAF" box="[1296,1364,186,212]" name="Oman" pageId="13" pageNumber="524">Oman</collectingCountry>
(Randall 1995), the Gulf (Almojil
<emphasis id="B9653C326648FFA8FE1815EFE314ED83" box="[483,532,222,249]" italics="true" pageId="13" pageNumber="524">eṫ al</emphasis>
. 2015) and
<collectingCountry id="F306A0B06648FFA8FD6215EFE3F8ED83" box="[665,760,222,248]" name="Pakistan" pageId="13" pageNumber="524">Pakistan</collectingCountry>
(Psomadakis
<emphasis id="B9653C326648FFA8FC6E15EFE2C7ED83" box="[917,967,222,249]" italics="true" pageId="13" pageNumber="524">eṫ al</emphasis>
. 2015). It has also been reported from the western coast of
<collectingCountry id="F306A0B06648FFA8FEAF1433E08EEC67" box="[340,398,258,284]" name="India" pageId="13" pageNumber="524">India</collectingCountry>
(Bineesh
<emphasis id="B9653C326648FFA8FE041433E330EC67" box="[511,560,258,285]" italics="true" pageId="13" pageNumber="524">eṫ al</emphasis>
. 2016).
</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB6648FFA8FF3C1417E47DEFD2" pageId="13" pageNumber="524" type="discussion">
<paragraph id="8BAEE0206648FFA8FF3C1417E47DEFD2" blockId="13.[151,1437,150,681]" pageId="13" pageNumber="524">
<emphasis id="B9653C326648FFA8FF3C1417E03DEC3B" bold="true" box="[199,317,294,320]" pageId="13" pageNumber="524">Remarks:</emphasis>
Photographed individuals match well all characters of
<emphasis id="B9653C326648FFA8FC4E1419E595EC3B" box="[949,1173,294,323]" italics="true" pageId="13" pageNumber="524">masṫẚnachus sephen</emphasis>
as given by Last
<emphasis id="B9653C326648FFA8FA991417E495EC3B" box="[1378,1429,294,321]" italics="true" pageId="13" pageNumber="524">eṫ al</emphasis>
. (2016c), particularly referring to the presence of prominent tubercles on dorsal middisc. The species occurs on sand and mud bottom of estuaries, bays and sheltered lagoons, at depths of
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(
<bibRefCitation id="EF809DD16648FFA8FBBB145FE5FDECF2" author="Weigmann, S." box="[1088,1277,366,393]" pageId="13" pageNumber="524" pagination="837 - 1037" refId="ref13064" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. gournal of c ẚsh B ẚology, 88 (3), 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874" type="journal article" year="2016">Weigmann 2016</bibRefCitation>
), often resting nearly buried in the sand or mud. The present confirmed record from the
<collectingCountry id="F306A0B06648FFA8FC3C14A3E51DECD7" box="[967,1053,402,428]" name="Yemen" pageId="13" pageNumber="524">Socotra</collectingCountry>
Archipelago is based on individuals observed on silty sand bottom close to large rocks with overhangs in Ras Bidou, western part of
<collectingCountry id="F306A0B06648FFA8FB0F1487E498ECAB" box="[1268,1432,438,464]" name="Socotra Island" pageId="13" pageNumber="524">Socotra Island</collectingCountry>
, at a depth of
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<quantity id="4CE94DC56648FFA8FED114EBE067EC8E" box="[298,359,474,501]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="524" unit="m" value="14.0">14 m</quantity>
.
</specimenCount>
<emphasis id="B9653C326648FFA8FE8914EDE350EC8F" box="[370,592,474,503]" italics="true" pageId="13" pageNumber="524">masṫẚnachus sephen</emphasis>
is included as near threatened in IUCN Red List Assessments and urgently requires conservation management (Kyne
<emphasis id="B9653C326648FFA8FD8E14CFE3A8EF63" box="[629,680,510,537]" italics="true" pageId="13" pageNumber="524">eṫ al</emphasis>
. 2017).
<emphasis id="B9653C326648FFA8FCFC1731E2C5EF62" box="[775,965,510,539]" italics="true" pageId="13" pageNumber="524">masṫẚnachus aṫer</emphasis>
(
<bibRefCitation id="EF809DD16648FFA8FC2F14CFE57FEF63" author="Macleay, W." box="[980,1151,510,536]" pageId="13" pageNumber="524" pagination="585 - 598" refId="ref12177" refString="Macleay, W. (1883) Contribution to a knowledge of the fishes of New Guinea. No. III. mroceed ẚngs of the i ẚnnean Soc ẚety of kew Sou th tales, 7 (4), 585 - 598. https: // doi. org / 10.5962 / bhl. part. 22766" type="journal article" year="1883">Macleay, 1883</bibRefCitation>
), reported from the Gulf,
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and
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eastward to
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, is a very similar species, which might possibly be found off
<collectingCountry id="F306A0B06648FFA8FF6C1777E1EDEF1B" box="[151,237,582,608]" name="Yemen" pageId="13" pageNumber="524">Socotra</collectingCountry>
Archipelago as well. It may be confused with
<emphasis id="B9653C326648FFA8FD0A1776E25AEF1B" box="[753,858,582,608]" italics="true" pageId="13" pageNumber="524">mK sephen</emphasis>
but differs by the length of eye and spiracle
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snout length (versus
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<emphasis id="B9653C326648FFA8FDE8175AE37CEFFF" box="[531,636,618,644]" italics="true" pageId="13" pageNumber="524">mK sephen</emphasis>
) and the presence of 34 tubercles mid-centrally on disc that are barely visible relative to the surrounding denticles (versus 23 tubercles prominent mid-centrally on disc in
<emphasis id="B9653C326648FFA8FAF817BEE46DEFD3" box="[1283,1389,654,680]" italics="true" pageId="13" pageNumber="524">mK sephen</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>