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<document id="AFB4E73069B8C4116F15C9C1764D8501" ID-CLB-Dataset="22494" ID-DOI="10.11646/zootaxa.4765.1.1" ID-GBIF-Dataset="8af5a092-f062-48c7-8d7c-f96ae6282bfa" ID-ISSN="1175-5334" ID-Zenodo-Dep="3774174" ID-ZooBank="urn:lsid:zoobank.org:pub:C442D94C-0EB4-4509-B762-913707214819" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588094957734" checkinUser="carolina" docAuthor="Hodgson, Chris" docDate="2020" docId="03B2EA640A4A46372CFCFE04FB8AD2FD" docLanguage="en" docName="zootaxa.4765.1.1.pdf.imf" docOrigin="Zootaxa 4765 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cryptococcidae Kosztarab 1968" docType="treatment" docVersion="6" lastPageNumber="102" masterDocId="FF8B921C0A2F46522C6BFF8EFFCBD747" masterDocTitle="A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males" masterLastPageNumber="264" masterPageNumber="1" pageNumber="102" updateTime="1698837970293" updateUser="ExternalLinkService">
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<mods:title id="4F9A71F97BB819CC55B6C22EE44CD8CF">A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males</mods:title>
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<treatment id="03B2EA640A4A46372CFCFE04FB8AD2FD" ID-DOI="http://doi.org/10.5281/zenodo.3796770" ID-GBIF-Taxon="163857819" ID-Zenodo-Dep="3796770" LSID="urn:lsid:plazi:treatment:03B2EA640A4A46372CFCFE04FB8AD2FD" httpUri="http://treatment.plazi.org/id/03B2EA640A4A46372CFCFE04FB8AD2FD" lastPageNumber="102" pageId="101" pageNumber="102">
<subSubSection id="C30108F90A4A46372CFCFE04FDB6D6E3" box="[151,637,394,421]" pageId="101" pageNumber="102" type="nomenclature">
<paragraph id="8BA45B720A4A46372CFCFE04FDB6D6E3" blockId="101.[151,637,394,421]" box="[151,637,394,421]" pageId="101" pageNumber="102">
<taxonomicName id="4C1B20F10A4A46372CFCFE04FDB6D6E3" authority="Kosztarab 1968" authorityName="Kosztarab" authorityYear="1968" box="[151,637,394,421]" class="Insecta" family="Cryptococcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="family">
<emphasis id="B96F87600A4A46372CFCFE04FDB6D6E3" bold="true" box="[151,637,394,421]" pageId="101" pageNumber="102">
CRYPTOCOCCIDAE
<bibRefCitation id="EF8A26830A4A46372DD3FE05FDB6D6E3" author="Kosztarab" box="[440,637,394,421]" firstAuthor="Kosztarab" pageId="101" pageNumber="102" pagination="12" refId="ref205898" refString="Kosztarab, M. P. (1968) Cryptococcidae, a new family of the Coccoidea (Homoptera). Virginia Journal of Science, 19, 12." type="journal article" year="1968">Kosztarab 1968</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
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<subSubSection id="C30108F90A4A46372CFCFE44FF31D547" pageId="101" pageNumber="102" type="reference_group">
<paragraph id="8BA45B720A4A46372CFCFE44FF31D547" blockId="101.[151,1436,457,512]" pageId="101" pageNumber="102">
<treatmentCitationGroup id="AB0B7C5C0A4A46372CFCFE44FF31D547" pageId="101" pageNumber="102">
<treatmentCitation id="0ABA7D630A4A46372CFCFE44FE76D6A6" box="[151,445,457,482]" pageId="101" pageNumber="102">
<taxonomicName id="4C1B20F10A4A46372CFCFE44FE76D6A6" ID-CoL="9G33N" authority="Douglas 1890" authorityName="Douglas" authorityYear="1890" box="[151,445,457,482]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A46372CFCFE44FEE9D6A6" box="[151,290,458,481]" italics="true" pageId="101" pageNumber="102">Cryptococcus</emphasis>
<bibRefCitation id="EF8A26830A4A46372D40FE47FE76D6A6" author="Douglas" box="[299,445,457,482]" firstAuthor="Douglas" pageId="101" pageNumber="102" pagination="153 - 155" refId="ref200157" refString="Douglas, J. W. (1890) Notes on some British and exotic Coccidae (No. 16). Entomologist's Monthly Magazine, 26, 153 - 155." type="journal article" year="1890">Douglas 1890</bibRefCitation>
</taxonomicName>
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, 155.
<typeStatus id="54A0E5D00A4A46372D95FE44FDFBD6A5" box="[510,560,458,482]" pageId="101" pageNumber="102">Type</typeStatus>
species:
<taxonomicName id="4C1B20F10A4A46372EF9FE44FC69D6A5" authority="Baerensprung" authorityName="Baerensprung" authorityYear="1849" box="[658,930,457,482]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="fagi">
<emphasis id="B96F87600A4A46372EF9FE44FCC7D6A6" box="[658,780,457,481]" italics="true" pageId="101" pageNumber="102">Coccus fagi</emphasis>
Baerensprung
</taxonomicName>
(=
<taxonomicName id="4C1B20F10A4A46372FA2FE44FAEED6A5" ID-CoL="ZWNL" authority="Lindinger" authorityName="Lindinger" authorityYear="1936" box="[969,1317,457,482]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="fagisuga">
<emphasis id="B96F87600A4A46372FA2FE44FB7FD6A6" box="[969,1204,457,481]" italics="true" pageId="101" pageNumber="102">Cryptococcus fagisuga</emphasis>
Lindinger
</taxonomicName>
), by monotypy.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30108F90A4A46372CFCFDA6FAE9D4AB" pageId="101" pageNumber="102" type="discussion">
<paragraph id="8BA45B720A4A46372CFCFDA6FAE9D4AB" blockId="101.[151,1437,552,1466]" pageId="101" pageNumber="102">
<emphasis id="B96F87600A4A46372CFCFDA6FEFDD504" bold="true" box="[151,310,552,579]" pageId="101" pageNumber="102">Introduction.</emphasis>
This small family, known as the bark-crevice scales, has now been shown to be non-monophyletic (
<bibRefCitation id="EF8A26830A4A46372CF4FDC2FE75D521" author="Gwiazdowski" box="[159,446,588,614]" etAl="et al." firstAuthor="Gwiazdowski" pageId="101" pageNumber="102" pagination="9 - 18" refId="ref202748" refString="Gwiazdowski, R. A., van Driesche, R. G., Desnoyers, A., Lyon, S., Wu, S. A., Kamata, N. &amp; Normark, B. B. (2006) Possible geographic origin of beech scale, Cryptococcus fagisuga (Hemiptera: Eriococcidae), an invasive pest in North America. Biological Control: Theory and Applications in Pest Management, 39 (1), 9 - 18. https: // doi. org / 10.1016 / j. biocontrol. 2006.04.009" type="journal article" year="2006">
Gwiazdowski
<emphasis id="B96F87600A4A46372D29FDC3FEB0D521" box="[322,379,588,614]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="EF8A26830A4A46372DA2FDC2FDB7D521" author="Nan" box="[457,636,588,614]" etAl="et al." firstAuthor="Nan" pageId="101" pageNumber="102" pagination="170" refId="ref209879" refString="Nan, N., Dean, G. J. &amp; Wu, S. A. (2013) A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang &amp; Hao) comb. n. Zootaxa, 3722 (2), 170. https: // doi. org / 10.11646 / zootaxa. 3722.2.3" type="journal article" year="2013">
Nan
<emphasis id="B96F87600A4A46372D94FDC3FDFAD521" box="[511,561,588,614]" italics="true" pageId="101" pageNumber="102">et al</emphasis>
. 2013
</bibRefCitation>
).
<bibRefCitation id="EF8A26830A4A46372EE5FDC2FC76D520" author="Gwiazdowski" box="[654,957,588,615]" etAl="et al." firstAuthor="Gwiazdowski" pageId="101" pageNumber="102" pagination="9 - 18" refId="ref202748" refString="Gwiazdowski, R. A., van Driesche, R. G., Desnoyers, A., Lyon, S., Wu, S. A., Kamata, N. &amp; Normark, B. B. (2006) Possible geographic origin of beech scale, Cryptococcus fagisuga (Hemiptera: Eriococcidae), an invasive pest in North America. Biological Control: Theory and Applications in Pest Management, 39 (1), 9 - 18. https: // doi. org / 10.1016 / j. biocontrol. 2006.04.009" type="journal article" year="2006">
Gwiazdowski
<emphasis id="B96F87600A4A46372F5BFDC3FCAAD521" box="[816,865,588,614]" italics="true" pageId="101" pageNumber="102">et al</emphasis>
. (2006)
</bibRefCitation>
, in a molecular study using 18S ribosomal DNA sequences of the
<typeStatus id="54A0E5D00A4A46372DF7FDFEFE07D5CD" box="[412,460,624,650]" pageId="101" pageNumber="102">type</typeStatus>
species,
<taxonomicName id="4C1B20F10A4A46372E58FDFEFCE1D5CE" authority="Lindinger" authorityName="Lindinger" authorityYear="1936" box="[563,810,623,650]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="fagisuga">
<emphasis id="B96F87600A4A46372E58FDFEFD7FD5CE" box="[563,692,623,650]" italics="true" pageId="101" pageNumber="102">C. fagisuga</emphasis>
Lindinger
</taxonomicName>
plus
<taxonomicName id="4C1B20F10A4A46372F03FDFEFBFDD5CE" authority="Brittin" authorityName="Brittin" authorityYear="1915" box="[872,1078,623,650]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="nudatus">
<emphasis id="B96F87600A4A46372F03FDFEFC28D5CE" box="[872,995,623,650]" italics="true" pageId="101" pageNumber="102">C. nudatus</emphasis>
Brittin
</taxonomicName>
,
<taxonomicName id="4C1B20F10A4A4637282AFDFEFA57D5CE" authority="Kosztarab &amp; Hale" authorityName="Kosztarab &amp; Hale" box="[1089,1436,623,650]" class="Insecta" family="Eriococcidae" genus="Lobimargo" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B96F87600A4A4637282AFDFEFB01D5CE" box="[1089,1226,623,650]" italics="true" pageId="101" pageNumber="102">C. williamsi</emphasis>
Kosztarab &amp; Hale
</taxonomicName>
and
<taxonomicName id="4C1B20F10A4A46372CADFD1DFE60D5EA" authority="Tang &amp; Hao" authorityName="Tang &amp; Hao (Wu" authorityYear="2000" box="[198,427,659,685]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="ulmi">
<emphasis id="B96F87600A4A46372CADFD1DFED2D5EA" box="[198,281,659,685]" italics="true" pageId="101" pageNumber="102">C. ulmi</emphasis>
Tang &amp; Hao
</taxonomicName>
, showed that the first three species fell within the Gondwanan clade of the “
<taxonomicName id="4C1B20F10A4A46372890FD1DFA58D5EA" box="[1275,1427,659,685]" class="Insecta" family="Eriococcidae" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="family">Eriococcidae</taxonomicName>
” of
<bibRefCitation id="EF8A26830A4A46372CDEFD38FE0CD597" author="Cook and Gullan" box="[181,455,694,720]" firstAuthor="Cook" pageId="101" pageNumber="102" pagination="441 - 452" refId="ref199531" refString="Cook, L. G. &amp; Gullan, P. J. (2004) The gall-inducing habit has evolved multiple times among the eriococcid scale insects (Sternorrhyncha: Coccoidea: Eriococcidae). Biological Journal of the Linnean Society, 83 (4), 441 - 452. https: // doi. org / 10.1111 / j. 1095 - 8312.2004.00396. x" type="journal article" year="2004">Cook and Gullan (2004)</bibRefCitation>
, but that
<taxonomicName id="4C1B20F10A4A46372E47FD39FDB5D597" authorityName="Tang &amp; Hao (Wu" authorityYear="2000" box="[556,638,694,721]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="ulmi">
<emphasis id="B96F87600A4A46372E47FD39FDB5D597" box="[556,638,694,721]" italics="true" pageId="101" pageNumber="102">C. ulmi</emphasis>
</taxonomicName>
did not but was closer to
<bibRefCitation id="EF8A26830A4A46372FF0FD38FB76D597" author="Cook and Gullan's" box="[923,1213,694,720]" firstAuthor="Cook" pageId="101" pageNumber="102" pagination="441 - 452" refId="ref199531" refString="Cook, L. G. &amp; Gullan, P. J. (2004) The gall-inducing habit has evolved multiple times among the eriococcid scale insects (Sternorrhyncha: Coccoidea: Eriococcidae). Biological Journal of the Linnean Society, 83 (4), 441 - 452. https: // doi. org / 10.1111 / j. 1095 - 8312.2004.00396. x" type="journal article" year="2004">Cook and Gullan’s (2004)</bibRefCitation>
“BSE” clade. Since then,
<bibRefCitation id="EF8A26830A4A46372CBFFD54FE5BD5B3" author="Nan" box="[212,400,730,756]" etAl="et al." firstAuthor="Nan" pageId="101" pageNumber="102" pagination="170" refId="ref209879" refString="Nan, N., Dean, G. J. &amp; Wu, S. A. (2013) A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang &amp; Hao) comb. n. Zootaxa, 3722 (2), 170. https: // doi. org / 10.11646 / zootaxa. 3722.2.3" type="journal article" year="2013">
Nan
<emphasis id="B96F87600A4A46372D62FD55FEF2D5B3" box="[265,313,730,756]" italics="true" pageId="101" pageNumber="102">et al</emphasis>
. (2013)
</bibRefCitation>
have confirmed these findings and transferred
<taxonomicName id="4C1B20F10A4A46372FF2FD54FC21D5B3" authorityName="Tang &amp; Hao (Wu" authorityYear="2000" box="[921,1002,730,756]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="ulmi">
<emphasis id="B96F87600A4A46372FF2FD54FC21D5B3" box="[921,1002,730,756]" italics="true" pageId="101" pageNumber="102">C. ulmi</emphasis>
</taxonomicName>
to a new genus (
<taxonomicName id="4C1B20F10A4A463728C8FD55FAACD5B3" box="[1187,1383,731,756]" class="Insecta" family="Eriococcidae" genus="Macroporicoccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A463728C8FD55FAACD5B3" box="[1187,1383,731,756]" italics="true" pageId="101" pageNumber="102">Macroporicoccus</emphasis>
</taxonomicName>
Nan &amp; Wu). As all known cryptococcid males are apterous, members of this family were not included in
<bibRefCitation id="EF8A26830A4A4637296CFD73FE88D47C" author="Hodgson and Hardy's" firstAuthor="Hodgson" pageId="101" pageNumber="102" pagination="794 - 804" refId="ref203953" refString="Hodgson, C. J. &amp; Hardy, N. B. (2013) The phylogeny of the superfamily Coccoidea (Hemiptera: Sternorrhyncha) based on the morphology of extant and extinct macropterous males. Systematic Entomology, 38, 794 - 804. https: // doi. org / 10.1111 / syen. 12030" type="journal article" year="2013">Hodgson and Hardy’s (2013)</bibRefCitation>
study based on adult male morphology. The family
<taxonomicName id="4C1B20F10A4A46372FFFFCAFFB8FD47C" authorityName="Kosztarab" authorityYear="1968" box="[916,1092,801,827]" class="Insecta" family="Cryptococcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="family">Cryptococcidae</taxonomicName>
therefore does not really exist but this group is dealt with here (despite apparently belonging to separate eriococcid clades) because of the apterous males. Thus, this group includes just 9 species in 3 genera, namely
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<taxonomicName id="4C1B20F10A4A46372FDAFCE6FB7AD4C5" authority="Douglas" authorityName="Douglas" authorityYear="1890" box="[945,1201,872,898]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A46372FDAFCE6FB80D4C5" box="[945,1099,872,898]" italics="true" pageId="101" pageNumber="102">Cryptococcus</emphasis>
Douglas
</taxonomicName>
,
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<taxonomicName id="4C1B20F10A4A46372887FCE6FF26D4E2" authority="Nitsche" authorityName="Nitsche" authorityYear="1895" class="Insecta" family="Eriococcidae" genus="Pseudochermes" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A46372887FCE6FA57D4C5" box="[1260,1436,872,898]" italics="true" pageId="101" pageNumber="102">Pseudochermes</emphasis>
Nitsche
</taxonomicName>
and
<quantity id="4CE3F6970A4A46372D49FC05FE86D4E2" box="[290,333,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="101" pageNumber="102" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C1B20F10A4A46372D38FC02FC90D4E2" authority="(Garcia Morales et al. 2019)" baseAuthorityName="Garcia Morales" baseAuthorityYear="2019" box="[339,859,907,933]" class="Insecta" family="Eriococcidae" genus="Macroporicoccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A46372D38FC02FDDCD4E2" box="[339,535,908,933]" italics="true" pageId="101" pageNumber="102">Macroporicoccus</emphasis>
(
<bibRefCitation id="EF8A26830A4A46372E4DFC05FC9FD4E2" author="Garcia Morales" box="[550,852,907,933]" etAl="et al." firstAuthor="Garcia Morales" pageId="101" pageNumber="102" refId="ref201635" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. &amp; Hardy, N. B. (2019) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: https: // data. nal. usda. gov / dataset / scalenet-scaleinsects-coccoidea-database (accessed 12 August 2019)" type="url" year="2019">
García Morales
<emphasis id="B96F87600A4A46372EB0FC02FCD8D4E2" box="[731,787,907,933]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
. These species have a mainly Holarctic distribution but with one species (
<taxonomicName id="4C1B20F10A4A46372DE7FC21FDCDD48E" authorityName="Brittin" authorityYear="1915" box="[396,518,943,969]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="nudatus">
<emphasis id="B96F87600A4A46372DE7FC21FDCDD48E" box="[396,518,943,969]" italics="true" pageId="101" pageNumber="102">C. nudatus</emphasis>
</taxonomicName>
) described from
<collectingCountry id="F30C1BE20A4A46372EADFC21FCABD48F" box="[710,864,942,969]" name="New Zealand" pageId="101" pageNumber="102">New Zealand</collectingCountry>
. The adult males of only
<taxonomicName id="4C1B20F10A4A46372817FC21FF12D4AB" authority="Tang &amp; Hao (Wu 2000)" authorityName="Tang &amp; Hao (Wu" authorityYear="2000" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="ulmi">
<emphasis id="B96F87600A4A46372817FC21FB1FD48E" box="[1148,1236,943,969]" italics="true" pageId="101" pageNumber="102">M. ulmi</emphasis>
Tang &amp; Hao (
<bibRefCitation id="EF8A26830A4A4637291EFC21FF1AD4AB" author="Wu, S. A." pageId="101" pageNumber="102" pagination="251 - 256" refId="ref213278" refString="Wu, S. A. (2000) Descriptions of male and immature stages of Cryptococcus ulmi Tang &amp; Hao (Homoptera: Coccoidea: Eriococcidae) with brief notes on its biology. Entomotaxonomia, 22 (4), 251 - 256." type="journal article" year="2000">Wu 2000</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1B20F10A4A46372D64FC5CFD2ED4AA" authority="(Kalt.) (Afifi 1968)" authorityName="Afifi" authorityYear="1968" baseAuthorityName="Kalt." box="[271,741,978,1005]" class="Insecta" family="Cryptococcidae" genus="Peudochermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="fraxini">
<emphasis id="B96F87600A4A46372D64FC5CFDCDD4AB" box="[271,518,978,1004]" italics="true" pageId="101" pageNumber="102">Peudochermes fraxini</emphasis>
(Kalt.) (
<bibRefCitation id="EF8A26830A4A46372E0EFC5CFD16D4AA" author="Afifi" box="[613,733,978,1005]" firstAuthor="Afifi" pageId="101" pageNumber="102" pagination="3 - 210" refId="ref197178" refString="Afifi, S. A. (1968) Morphology and taxonomy of the adult males of the families Pseudococcidae and Eriococcidae (Homoptera: Coccoidea). British Museum (Natural History), Second Report on Economic Zoology, 13, 3 - 210." type="journal article" year="1968">Afifi 1968</bibRefCitation>
)
</taxonomicName>
have been described and appear to be very similar.
</paragraph>
</subSubSection>
<subSubSection id="C30108F90A4A46372CACFC7BFB50D2D1" pageId="101" pageNumber="102" type="diagnosis">
<paragraph id="8BA45B720A4A46372CACFC7BFB50D2D1" blockId="101.[151,1437,552,1466]" pageId="101" pageNumber="102">
<emphasis id="B96F87600A4A46372CACFC7BFAABD357" bold="true" box="[199,1376,1013,1040]" pageId="101" pageNumber="102">
Family diagnosis based on the adult male morphology of
<taxonomicName id="4C1B20F10A4A46372F04FC78FBAFD357" authority="Tang &amp; Hao" authorityName="Tang &amp; Hao (Wu" authorityYear="2000" box="[879,1124,1013,1040]" class="Insecta" family="Eriococcidae" genus="Cryptococcus" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="ulmi">
<emphasis id="B96F87600A4A46372F04FC78FC00D357" bold="true" box="[879,971,1013,1040]" italics="true" pageId="101" pageNumber="102">M. ulmi</emphasis>
Tang &amp; Hao
</taxonomicName>
and
<taxonomicName id="4C1B20F10A4A463728F4FC78FAABD357" authority="(Kalt.)" authorityName="Afifi" authorityYear="1968" baseAuthorityName="Kalt." box="[1183,1376,1013,1040]" class="Insecta" family="Cryptococcidae" genus="Peudochermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="fraxini">
<emphasis id="B96F87600A4A463728F4FC78FAC1D357" bold="true" box="[1183,1290,1013,1040]" italics="true" pageId="101" pageNumber="102">P. fraxini</emphasis>
(Kalt.)
</taxonomicName>
</emphasis>
(
<figureCitation id="132047F70A4A46372904FC7BFF7DD374" captionStart="FIGURE 41" captionStartId="102.[151,250,1705,1730]" captionTargetBox="[161,1430,188,1680]" captionTargetId="figure@102.[151,1436,181,1680]" captionTargetPageId="102" captionText="FIGURE 41. Pseudochermes fraxini (Kalt.). Apterous adult male.(Cryptococcidae; Gondwanan group). (AfterAfifi 1968). Where C=apical antennal segments, F=metathoracic leg and H=side view of posterior abdominal segments and penial sheath. For abbreviations see p. 8." figureDoi="http://doi.org/10.5281/zenodo.3774256" httpUri="https://zenodo.org/record/3774256/files/figure.png" pageId="101" pageNumber="102">Fig. 41</figureCitation>
).
<emphasis id="B96F87600A4A46372CA7FB97FEDBD374" bold="true" box="[204,272,1049,1075]" pageId="101" pageNumber="102">Body.</emphasis>
<emphasis id="B96F87600A4A46372D72FB97FEB6D374" box="[281,381,1049,1075]" italics="true" pageId="101" pageNumber="102">Apterous</emphasis>
, without a neck; body roundly pointed anteriorly, abdomen narrowing to a point posteriorly.
<emphasis id="B96F87600A4A46372CFCFBB3FEF5D310" box="[151,318,1085,1111]" italics="true" pageId="101" pageNumber="102">Setae very few</emphasis>
, all hs (
<taxonomicName id="4C1B20F10A4A46372DCBFBB2FD9BD311" authorityName="Nitsche" authorityYear="1895" box="[416,592,1084,1110]" class="Insecta" family="Eriococcidae" genus="Pseudochermes" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A46372DCBFBB2FD9BD311" box="[416,592,1084,1110]" italics="true" pageId="101" pageNumber="102">Pseudochermes</emphasis>
</taxonomicName>
) or some peg-like (
<taxonomicName id="4C1B20F10A4A46372F5DFBB3FC31D311" box="[822,1018,1085,1110]" class="Insecta" family="Eriococcidae" genus="Macroporicoccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4A46372F5DFBB3FC31D311" box="[822,1018,1085,1110]" italics="true" pageId="101" pageNumber="102">Macroporicoccus</emphasis>
</taxonomicName>
); pores absent.
<emphasis id="B96F87600A4A463728DFFBB3FB3FD310" bold="true" box="[1204,1268,1085,1111]" pageId="101" pageNumber="102">Head</emphasis>
<emphasis id="B96F87600A4A46372896FBB2FE7ED33D" italics="true" pageId="101" pageNumber="102">with a pair of ocelli but no simple eyes</emphasis>
;
<emphasis id="B96F87600A4A46372DADFBEFFD35D33D" box="[454,766,1120,1146]" italics="true" pageId="101" pageNumber="102">antennae 7 or 8 segmented</emphasis>
, segments varied in shape; some segments with fs and hs setae; apex with a few short capitate setae and antennal bristles barely differentiated; with vestigial mouthparts; all cranial ridges absent.
<emphasis id="B96F87600A4A46372DFAFB29FE21D386" bold="true" box="[401,490,1191,1217]" pageId="101" pageNumber="102">Thorax</emphasis>
.
<emphasis id="B96F87600A4A46372D91FB29FC4AD386" box="[506,897,1191,1217]" italics="true" pageId="101" pageNumber="102">Pronotal ridge and sclerite absent</emphasis>
; prosternum with transverse ridge only; with a sparse band of hs.
<emphasis id="B96F87600A4A46372D03FB44FCF4D3A3" box="[360,831,1226,1252]" italics="true" pageId="101" pageNumber="102">Mesothorax and metathorax membranous</emphasis>
, with a few hs. Spiracles normal, without pores. Legs moderately developed, five segmented; arrangement of trochanter sensilla and presence of tarsal campaniform pore unknown; setae few; tibia with 2 spurs;
<emphasis id="B96F87600A4A46372E3BFA9FFC08D26C" box="[592,963,1297,1323]" italics="true" pageId="101" pageNumber="102">tarsal and claw digitules capitate</emphasis>
; claw without a denticle.
<emphasis id="B96F87600A4A4637288BFA9EFA9ED26C" bold="true" box="[1248,1365,1296,1323]" pageId="101" pageNumber="102">Abdomen</emphasis>
membranous, gradually narrowing to a short penial sheath; with segmental bands of hs (
<emphasis id="B96F87600A4A46372824FAB8FAD5D208" box="[1103,1310,1334,1359]" italics="true" pageId="101" pageNumber="102">Macroporicoccuss</emphasis>
with some peg-like setae in pleural areas). Caudal extensions, ostioles and glandular pouches absent. Penial sheath about as long as broad, abdominal segment IX fused to style, with a few short setae. Aedeagus short.
</paragraph>
</subSubSection>
<subSubSection id="C30108F90A4A46372CACFA2EFB8AD2FD" box="[199,1089,1439,1466]" pageId="101" pageNumber="102" type="discussion">
<paragraph id="8BA45B720A4A46372CACFA2EFB8AD2FD" blockId="101.[151,1437,552,1466]" box="[199,1089,1439,1466]" pageId="101" pageNumber="102">
<bibRefCitation id="EF8A26830A4A46372CACFA2EFEA9D2FD" author="Afifi's" box="[199,354,1439,1466]" firstAuthor="Afifi's" pageId="101" pageNumber="102" pagination="3 - 210" refId="ref197178" refString="Afifi, S. A. (1968) Morphology and taxonomy of the adult males of the families Pseudococcidae and Eriococcidae (Homoptera: Coccoidea). British Museum (Natural History), Second Report on Economic Zoology, 13, 3 - 210." type="journal article" year="1968">Afifi’s (1968)</bibRefCitation>
figure of the adult male of
<taxonomicName id="4C1B20F10A4A46372EFEFA2EFD37D2FD" authorityName="Afifi" authorityYear="1968" baseAuthorityName="Kalt." box="[661,764,1440,1466]" class="Insecta" family="Cryptococcidae" genus="Peudochermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="fraxini">
<emphasis id="B96F87600A4A46372EFEFA2EFD37D2FD" box="[661,764,1440,1466]" italics="true" pageId="101" pageNumber="102">P. fraxini</emphasis>
</taxonomicName>
is reproduced here (
<figureCitation id="132047F70A4A46372F8BFA2EFBFFD2FD" box="[992,1076,1440,1466]" captionStart="FIGURE 41" captionStartId="102.[151,250,1705,1730]" captionTargetBox="[161,1430,188,1680]" captionTargetId="figure@102.[151,1436,181,1680]" captionTargetPageId="102" captionText="FIGURE 41. Pseudochermes fraxini (Kalt.). Apterous adult male.(Cryptococcidae; Gondwanan group). (AfterAfifi 1968). Where C=apical antennal segments, F=metathoracic leg and H=side view of posterior abdominal segments and penial sheath. For abbreviations see p. 8." figureDoi="http://doi.org/10.5281/zenodo.3774256" httpUri="https://zenodo.org/record/3774256/files/figure.png" pageId="101" pageNumber="102">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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