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<document id="038774EE8F9EBF3D152EA7AECD845EB6" ID-CLB-Dataset="28065" ID-DOI="10.11646/zootaxa.4504.4.5" ID-GBIF-Dataset="343a6a67-e8bf-467c-9818-de9621bbe955" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606614" ID-ZooBank="08E1A79F-BAA4-4EAB-9636-FE44BF7EB7F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585132792" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2018" docId="03AB87D4E7626F6C12F9F9D6C8067FFF" docLanguage="en" docName="zootaxa.4504.4.5.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systaria princesa Jäger 2018, spec. nov." docType="treatment" docVersion="9" lastPageNumber="538" masterDocId="FF92FFACE76F6F62126EFFC4C9517912" masterDocTitle="On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests" masterLastPageNumber="544" masterPageNumber="524" pageNumber="537" updateTime="1698712839155" updateUser="ExternalLinkService">
<mods:mods id="AFE6AFC883DB2EC8C9F223128F7EAAD2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="51DD1BB149DF3D1D958E3FF14898779E">
<mods:title id="A0B3D97D7415DBFD22DF19945010BEFB">On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="FF58CCDBE3B29660761C010710071E58">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7283734CE8F79E58EE79E621AECF9C9B">Jäger, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource id="CBC02CF09BEABC95D9FE484947D62CB2">text</mods:typeOfResource>
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<mods:title id="8A9FC0C835440204A753CD852CD1DE82">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5EA06D7ED68A2429CCD0FBB311AD88C8">
<mods:date id="5C6B31F44F2D52915468A1CC168DC6B0">2018</mods:date>
<mods:detail id="815BED6A91D9BD292CEF6BA1FAEED580" type="pubDate">
<mods:number id="13599D3202E2EB812CAC294FC81A8900">2018-10-26</mods:number>
</mods:detail>
<mods:detail id="1B7DE63CE5A1BC6182320F7167F58D07" type="volume">
<mods:number id="B7EF2680A6051A219CC5EDF6D982A63D">4504</mods:number>
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<mods:number id="043E0D17E96B8EED586294D3143B9700">4</mods:number>
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<mods:classification id="E50077EE84276772C680DEE375EBB69E">journal article</mods:classification>
<mods:identifier id="EC110CD854F0601E464A05BB3FB96636" type="CLB-Dataset">28065</mods:identifier>
<mods:identifier id="CDAE490062B9C131ED6A638E9C2A08F5" type="DOI">10.11646/zootaxa.4504.4.5</mods:identifier>
<mods:identifier id="6A585A2605F1E602B67B760262B7464D" type="GBIF-Dataset">343a6a67-e8bf-467c-9818-de9621bbe955</mods:identifier>
<mods:identifier id="4D5B5ABB0D03CFBD29C4049833A1B6B5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="ACDEABF803A4EFB5E8503B148ED09F8A" type="Zenodo-Dep">2606614</mods:identifier>
<mods:identifier id="5C9F59E64A6EF2640993461C8830DF7B" type="ZooBank">08E1A79F-BAA4-4EAB-9636-FE44BF7EB7F3</mods:identifier>
</mods:mods>
<treatment id="03AB87D4E7626F6C12F9F9D6C8067FFF" ID-DOI="http://doi.org/10.5281/zenodo.5988018" ID-GBIF-Taxon="154495203" ID-Zenodo-Dep="5988018" LSID="urn:lsid:plazi:treatment:03AB87D4E7626F6C12F9F9D6C8067FFF" httpUri="http://treatment.plazi.org/id/03AB87D4E7626F6C12F9F9D6C8067FFF" lastPageId="14" lastPageNumber="538" pageId="13" pageNumber="537">
<subSubSection id="C3186549E7626F6F12F9F9D6C8B47F3F" box="[151,485,1554,1581]" pageId="13" pageNumber="537" type="nomenclature">
<paragraph id="8BBD36C2E7626F6F12F9F9D6C8B47F3F" blockId="13.[151,485,1554,1614]" box="[151,485,1554,1581]" pageId="13" pageNumber="537">
<heading id="D0F581AEE7626F6F12F9F9D6C8B47F3F" bold="true" box="[151,485,1554,1581]" fontSize="11" level="1" pageId="13" pageNumber="537" reason="1">
<emphasis id="B976EAD0E7626F6F12F9F9D6C8B47F3F" bold="true" box="[151,485,1554,1581]" pageId="13" pageNumber="537">
<taxonomicName id="4C024D41E7626F6F12F9F9D6C8377F3E" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[151,358,1554,1580]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="537" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="B976EAD0E7626F6F12F9F9D6C8377F3E" bold="true" box="[151,358,1554,1580]" italics="true" pageId="13" pageNumber="537">Systaria princesa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557ABE7626F6F1303F9D7C8B47F3F" box="[365,485,1555,1581]" pageId="13" pageNumber="537" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7626F6F12F9F9F1C8397F5C" box="[151,360,1589,1614]" pageId="13" pageNumber="537" type="description">
<paragraph id="8BBD36C2E7626F6F12F9F9F1C8397F5C" blockId="13.[151,485,1554,1614]" box="[151,360,1589,1614]" pageId="13" pageNumber="537">
<figureCitation id="13392A47E7626F6F12F9F9F1C8467F5C" box="[151,279,1589,1614]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="13" pageNumber="537">Figs 33–39</figureCitation>
,
<figureCitation id="13392A47E7626F6F134CF9F1C8397F5C" box="[290,360,1589,1614]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="13" pageNumber="537">45–46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7626F6C12F9F9B9CA5E79C6" lastPageId="14" lastPageNumber="538" pageId="13" pageNumber="537" type="materials_examined">
<paragraph id="8BBD36C2E7626F6F12F9F9B9CC757E7C" blockId="13.[151,1437,1661,2009]" pageId="13" pageNumber="537">
<emphasis id="B976EAD0E7626F6F12F9F9B9C81A7F84" bold="true" box="[151,331,1661,1686]" pageId="13" pageNumber="537">Type material:</emphasis>
<materialsCitation id="3B6A3C9FE7626F6F1338F9BACA6E7F84" ID-GBIF-Occurrence="2028138101" box="[342,831,1661,1686]" country="Philippines" location="Palawan" pageId="13" pageNumber="537" specimenCount="1" stateProvince="Palawan" typeStatus="holotype">
<collectingCountry id="F3157652E7626F6F1338F9BACB517F84" box="[342,512,1662,1686]" name="Philippines" pageId="13" pageNumber="537">PHILIPPINES</collectingCountry>
:
<emphasis id="B976EAD0E7626F6F1060F9B9CB2B7F84" box="[526,634,1661,1686]" italics="true" pageId="13" pageNumber="537">
<collectingRegion id="49C6F820E7626F6F1060F9B9CB247F84" box="[526,629,1661,1686]" country="Philippines" name="Palawan" pageId="13" pageNumber="537">Palawan</collectingRegion>
:
</emphasis>
<emphasis id="B976EAD0E7626F6F10E8F9B9CBA27F84" bold="true" box="[646,755,1661,1686]" pageId="13" pageNumber="537">
<typeStatus id="54B98860E7626F6F10E8F9B9CBA27F84" box="[646,755,1661,1686]" pageId="13" pageNumber="537" type="holotype">Holotype</typeStatus>
</emphasis>
: male
</materialsCitation>
,
<materialsCitation id="3B6A3C9FE7626F6F1120F9BAC81C7FCC" ID-GBIF-Occurrence="2028138112" collectingDate="2017-05-03" collectionCode="SMF" collectorName="Vanni, P. &amp; M. Ciaramella" country="Philippines" elevation="305" location="Tagusan cave" municipality="Underground River National Park" pageId="13" pageNumber="537" specimenCount="1" stateProvince="Puerto Princesa" typeStatus="holotype">
<collectingRegion id="49C6F820E7626F6F1120F9BACD537F84" box="[846,1026,1661,1686]" country="Philippines" name="Puerto Princesa" pageId="13" pageNumber="537">Puerto Princesa</collectingRegion>
<collectingMunicipality id="6BD9ACB8E7626F6F1663F9B9CCC97F84" box="[1037,1432,1661,1686]" pageId="13" pageNumber="537">Underground River National Park</collectingMunicipality>
,
<location id="8EDD6019E7626F6F12F9F966C87F7FAB" LSID="urn:lsid:plazi:treatment:03AB87D4E7626F6C12F9F9D6C8067FFF:8EDD6019E7626F6F12F9F966C87F7FAB" box="[151,302,1698,1722]" country="Philippines" municipality="Underground River National Park" name="Tagusan cave" pageId="13" pageNumber="537" stateProvince="Puerto Princesa">Tagusan cave</location>
[19], N 10°9'43.93'', E 118°53'30.05'',
<quantity id="4CFA9B27E7626F6F108FF965CA797FAB" box="[737,808,1697,1722]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="13" pageNumber="537" unit="m" value="305.0">
<elevation id="002FD1F1E7626F6F108FF965CA797FAB" box="[737,808,1697,1722]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="13" pageNumber="537" unit="m" value="305.0">305 m</elevation>
</quantity>
elevation, S.
<collectorName id="26F75314E7626F6F11AEF965CD727FAB" box="[960,1059,1697,1722]" pageId="13" pageNumber="537">Vanni, P.</collectorName>
Agnelli &amp;
<collectorName id="26F75314E7626F6F16CFF966CC127FA8" box="[1185,1347,1697,1722]" pageId="13" pageNumber="537">M. Ciaramella</collectorName>
leg. 2.–
<date id="FFBC1002E7626F6F12F9F901C9A97FCC" box="[151,248,1733,1758]" pageId="13" pageNumber="537" value="2017-05-03">
<collectingDate id="EFF8E9EAE7626F6F12F9F901C9A97FCC" box="[151,248,1733,1758]" pageId="13" pageNumber="537" value="2017-05-03">3.5.2017</collectingDate>
</date>
(
<collectionCode id="ED13AE07E7626F6F1369F902C8147FCC" box="[263,325,1734,1758]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="537">SMF</collectionCode>
)
</materialsCitation>
.
<emphasis id="B976EAD0E7626F6F1339F901CBE07FCC" bold="true" box="[343,689,1733,1758]" pageId="13" pageNumber="537">
<typeStatus id="54B98860E7626F6F1339F901C89E7FCC" box="[343,463,1733,1758]" pageId="13" pageNumber="537" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D04FD4BE7626F6F13B1F901CB7F7FCC" box="[479,558,1733,1758]" pageId="13" pageNumber="537" type="male">1 male</specimenCount>
,
<specimenCount id="9D04FD4BE7626F6F1054F901CBFB7FCC" box="[570,682,1733,1758]" pageId="13" pageNumber="537" type="female">4 females</specimenCount>
)
</emphasis>
.
<materialsCitation id="3B6A3C9FE7626F6F10D1F902CB8E7E10" ID-GBIF-Occurrence="2028138115" collectingDate="2017-05-03" collectionCode="SMF" collectorName="Vanni, P. &amp; M. Ciaramella" country="Philippines" elevation="305" location="Underground River National Park" municipality="Underground River National Park" pageId="13" pageNumber="537" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Puerto Princesa" typeStatus="paratype">
<collectingRegion id="49C6F820E7626F6F10D1F902CA3F7FCC" box="[703,878,1733,1758]" country="Philippines" name="Puerto Princesa" pageId="13" pageNumber="537">Puerto Princesa</collectingRegion>
<location id="8EDD6019E7626F6F111AF901CDA27FCC" LSID="urn:lsid:plazi:treatment:03AB87D4E7626F6C12F9F9D6C8067FFF:8EDD6019E7626F6F111AF901CDA27FCC" box="[884,1267,1733,1758]" country="Philippines" name="Underground River National Park" pageId="13" pageNumber="537" stateProvince="Puerto Princesa">Underground River National Park</location>
:
<specimenCount id="9D04FD4BE7626F6F1692F902CC317FCC" box="[1276,1376,1733,1758]" pageId="13" pageNumber="537" type="female">1 female</specimenCount>
, with same data as for holotype (
<collectionCode id="ED13AE07E7626F6F13A2F92ECB5B7E10" box="[460,522,1770,1794]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="537">SMF</collectionCode>
).
<specimenCount id="9D04FD4BE7626F6F1073F92ECB3C7E10" box="[541,621,1769,1794]" pageId="13" pageNumber="537" type="male">1 male</specimenCount>
,
<specimenCount id="9D04FD4BE7626F6F1014F92ECB8E7E10" box="[634,735,1769,1794]" pageId="13" pageNumber="537" type="female">1 female</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B6A3C9FE7626F6F1082F92ECC717E7C" ID-GBIF-Occurrence="2028138108" collectingDate="2017-04-26" collectingDateMax="2017-05-08" collectingDateMin="2017-04-26" collectionCode="PNM, MZUF" collectorName="Vanni, P. &amp; M. Ciaramella &amp; Cabayugan &amp; C. Paniccia" country="Philippines" county="Underground River cave" elevation="1" location="Lion cave" municipality="Tagusan cave" pageId="13" pageNumber="537" specimenCount="2" specimenCount-female="2" stateProvince="Puerto Princesa" typeStatus="paratype">
<collectingRegion id="49C6F820E7626F6F1082F92ECACC7E10" box="[748,925,1769,1794]" country="Philippines" name="Puerto Princesa" pageId="13" pageNumber="537">Puerto Princesa</collectingRegion>
<collectingCounty id="62DC4E4EE7626F6F11CBF92DCDEE7E13" box="[933,1215,1769,1794]" pageId="13" pageNumber="537">Underground River cave</collectingCounty>
, “gypsum and mud galleries”, entrance (outflow) at N 10°11’55.77’’, E 118°55’33.41’’,
<quantity id="4CFA9B27E7626F6F11D8F8CACAB47E37" box="[950,997,1806,1830]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="537" unit="m" value="1.0">
<elevation id="002FD1F1E7626F6F11D8F8CACAB47E37" box="[950,997,1806,1830]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="537" unit="m" value="1.0">1 m</elevation>
</quantity>
elevation, S.
<collectorName id="26F75314E7626F6F16E4F8C9CDA37E37" box="[1162,1266,1805,1830]" pageId="13" pageNumber="537">Vanni, P.</collectorName>
Agnelli &amp;
<collectorName id="26F75314E7626F6F1712F8CAC8437E58" pageId="13" pageNumber="537">M. Ciaramella</collectorName>
leg.
<date id="FFBC1002E7626F6F133EF8F5C8927E58" box="[336,451,1841,1866]" pageId="13" pageNumber="537" value="2017-04-26">
<collectingDate id="EFF8E9EAE7626F6F133EF8F5C8927E58" box="[336,451,1841,1866]" pageId="13" pageNumber="537" value="2017-04-26">26.4.2017</collectingDate>
</date>
, #18 (
<collectionCode id="ED13AE07E7626F6F1060F8F6CB007E58" box="[526,593,1842,1866]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="13" pageNumber="537">PNM</collectionCode>
).
<specimenCount id="9D04FD4BE7626F6F1009F8F6CB9F7E58" box="[615,718,1841,1866]" pageId="13" pageNumber="537" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BD9ACB8E7626F6F10B3F8F6CA2A7E5B" box="[733,891,1842,1866]" pageId="13" pageNumber="537">Tagusan cave</collectingMunicipality>
, S.
<collectorName id="26F75314E7626F6F11C4F8F5CD407E5B" box="[938,1041,1841,1866]" pageId="13" pageNumber="537">Vanni, P.</collectorName>
Agnelli &amp;
<collectorName id="26F75314E7626F6F16F4F8F6CC117E58" box="[1178,1344,1841,1866]" pageId="13" pageNumber="537">M. Ciaramella</collectorName>
leg. 2.-
<date id="FFBC1002E7626F6F12F9F891C9AA7E7C" box="[151,251,1877,1902]" pageId="13" pageNumber="537" value="2017-05-03">
<collectingDate id="EFF8E9EAE7626F6F12F9F891C9AA7E7C" box="[151,251,1877,1902]" pageId="13" pageNumber="537" value="2017-05-03">3.5.2017</collectingDate>
</date>
, #16 (
<collectionCode id="ED13AE07E7626F6F132EF892C8C57E7C" box="[320,404,1878,1902]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="13" pageNumber="537">MZUF</collectionCode>
).
<specimenCount id="9D04FD4BE7626F6F13C8F892CB5B7E7C" box="[422,522,1877,1902]" pageId="13" pageNumber="537" type="female">1 female</specimenCount>
,
<location id="8EDD6019E7626F6F1078F891CBD57E7F" LSID="urn:lsid:plazi:treatment:03AB87D4E7626F6C12F9F9D6C8067FFF:8EDD6019E7626F6F1078F891CBD57E7F" box="[534,644,1877,1902]" country="Philippines" county="Underground River cave" municipality="Tagusan cave" name="Lion cave" pageId="13" pageNumber="537" stateProvince="Puerto Princesa">Lion cave</location>
(
<collectorName id="26F75314E7626F6F10FDF891CA467E7C" box="[659,791,1877,1902]" pageId="13" pageNumber="537">Cabayugan</collectorName>
), S. Vanni &amp;
<collectorName id="26F75314E7626F6F11C3F892CD7C7E7C" box="[941,1069,1877,1902]" pageId="13" pageNumber="537">C. Paniccia</collectorName>
leg.
<date id="FFBC1002E7626F6F160AF891CD947E7C" box="[1124,1221,1877,1902]" pageId="13" pageNumber="537" value="2017-05-08">
<collectingDate id="EFF8E9EAE7626F6F160AF891CD947E7C" box="[1124,1221,1877,1902]" pageId="13" pageNumber="537" value="2017-05-08">8.5.2017</collectingDate>
</date>
(
<collectionCode id="ED13AE07E7626F6F16BBF892CC497E7C" box="[1237,1304,1878,1902]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="13" pageNumber="537">PNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBD36C2E7626F6F12A9F8BDCCCD7ECA" blockId="13.[151,1437,1661,2009]" pageId="13" pageNumber="537">
<emphasis id="B976EAD0E7626F6F12A9F8BDCD8E7E80" bold="true" box="[199,1247,1913,1938]" pageId="13" pageNumber="537">
Additional material examined (
<specimenCount id="9D04FD4BE7626F6F1039F8BDCB8F7E80" box="[599,734,1913,1938]" pageId="13" pageNumber="537" type="subadult">2 subadult</specimenCount>
males,
<specimenCount id="9D04FD4BE7626F6F1124F8BDCA817E80" box="[842,976,1913,1938]" pageId="13" pageNumber="537" type="subadult">1 subadult</specimenCount>
female,
<specimenCount id="9D04FD4BE7626F6F1629F8BDCD827E80" box="[1095,1235,1913,1938]" pageId="13" pageNumber="537" type="juvenile">3 juveniles</specimenCount>
).
</emphasis>
<materialsCitation id="3B6A3C9FE7626F6F1681F8BECCCD7ECA" ID-GBIF-Occurrence="2028138107" collectingDate="2017-05-03" collectionCode="SMF" collectorName="Vanni, P. &amp; M. Ciaramella" country="Philippines" elevation="1" location="Underground River cave" municipality="Underground River National Park" pageId="13" pageNumber="537" specimenCount="3" specimenCount-subadult="3" stateProvince="Palawan">
<collectingCountry id="F3157652E7626F6F1681F8BECCC87E80" box="[1263,1433,1914,1938]" name="Philippines" pageId="13" pageNumber="537">PHILIPPINES</collectingCountry>
:
<emphasis id="B976EAD0E7626F6F12F9F859C8557EA4" box="[151,260,1949,1974]" italics="true" pageId="13" pageNumber="537">
<collectingRegion id="49C6F820E7626F6F12F9F859C9AF7EA4" box="[151,254,1949,1974]" country="Philippines" name="Palawan" pageId="13" pageNumber="537">Palawan</collectingRegion>
:
</emphasis>
<emphasis id="B976EAD0E7626F6F137CF859C8A47EA4" bold="true" box="[274,501,1949,1974]" pageId="13" pageNumber="537">
<specimenCount id="9D04FD4BE7626F6F137CF859C8C67EA4" box="[274,407,1949,1974]" pageId="13" pageNumber="537" type="subadult">1 subadult</specimenCount>
female
</emphasis>
, with same data as for holotype (
<collectionCode id="ED13AE07E7626F6F11F1F859CA8C7EA7" box="[927,989,1949,1973]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="537">SMF</collectionCode>
).
<emphasis id="B976EAD0E7626F6F1199F859CD877EA4" bold="true" box="[1015,1238,1949,1974]" pageId="13" pageNumber="537">
<specimenCount id="9D04FD4BE7626F6F1199F859CD2D7EA4" box="[1015,1148,1949,1974]" pageId="13" pageNumber="537" type="subadult">2 subadult</specimenCount>
males,
</emphasis>
<collectingRegion id="49C6F820E7626F6F168BF85ACCCD7EA4" box="[1253,1436,1949,1974]" country="Philippines" name="Puerto Princesa" pageId="13" pageNumber="537">Puerto Princesa</collectingRegion>
<location id="8EDD6019E7626F6F12F9F804C8FC7ECA" LSID="urn:lsid:plazi:treatment:03AB87D4E7626F6C12F9F9D6C8067FFF:8EDD6019E7626F6F12F9F804C8FC7ECA" box="[151,429,1984,2009]" country="Philippines" name="Underground River cave" pageId="13" pageNumber="537" stateProvince="Palawan">Underground River cave</location>
, “gypsum and mud galleries”, entrance (outflow) at N 10°11'55.77'', E 118°55'33.41'',
<quantity id="4CFA9B27E7626F6F171CF804CCCD7ECA" box="[1394,1436,1984,2008]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="537" unit="m" value="1.0">
<elevation id="002FD1F1E7626F6F171CF804CCCD7ECA" box="[1394,1436,1984,2008]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="537" unit="m" value="1.0">1 m</elevation>
</quantity>
</materialsCitation>
</paragraph>
<paragraph id="8BBD36C2E7616F6C12F9FF53CA5E79C6" blockId="14.[151,1437,151,1773]" pageId="14" pageNumber="538">
elevation, S. Vanni, P. Agnelli &amp; M. Ciaramella leg.
<date id="FFBC1002E7616F6C108EFF53CA0279A2" box="[736,851,151,176]" pageId="14" pageNumber="538" value="2017-04-26">26.4.2017</date>
, #18 (PNM).
<emphasis id="B976EAD0E7616F6C1185FF53CD2179A2" bold="true" box="[1003,1136,151,176]" pageId="14" pageNumber="538">
<specimenCount id="9D04FD4BE7616F6C1185FF53CD3A79A2" box="[1003,1131,151,176]" pageId="14" pageNumber="538" type="juvenile">3 juveniles</specimenCount>
,
</emphasis>
Tagusan cave, S. Vanni, P. Agnelli &amp; M. Ciaramella leg. 2.-
<date id="FFBC1002E7616F6C106EFF7FCB3B79C6" box="[512,618,187,212]" pageId="14" pageNumber="538" value="2017-05-03">3.5.2017</date>
, #16 (MZUF).
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7616F6C12A9FF1ACD7479E5" box="[199,1061,222,248]" pageId="14" pageNumber="538" type="etymology">
<paragraph id="8BBD36C2E7616F6C12A9FF1ACD7479E5" blockId="14.[151,1437,151,1773]" box="[199,1061,222,248]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9FF1AC81F79E5" bold="true" box="[199,334,222,247]" pageId="14" pageNumber="538">Etymology.</emphasis>
The species is named after the
<typeStatus id="54B98860E7616F6C10C0FF24CB8F79EA" box="[686,734,224,248]" pageId="14" pageNumber="538">type</typeStatus>
locality; name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7616F6C12A9FEC6CA847B01" pageId="14" pageNumber="538" type="diagnosis">
<paragraph id="8BBD36C2E7616F6C12A9FEC6CA847B01" blockId="14.[151,1437,151,1773]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9FEC6C8107809" bold="true" box="[199,321,258,283]" pageId="14" pageNumber="538">Diagnosis.</emphasis>
Small to medium sized spiders with body length of males: 7.5–9.1, of females: 9.2–12.2. The species seems to be closely related to
<taxonomicName id="4C024D41E7616F6C1029FEE2CAD4782C" authority="(Rainbow 1901)" baseAuthorityName="Rainbow" baseAuthorityYear="1901" box="[583,901,293,318]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B976EAD0E7616F6C1029FEE2CB94782C" box="[583,709,293,318]" italics="true" pageId="14" pageNumber="538">S. insulana</emphasis>
(Rainbow 1901)
</taxonomicName>
from the
<collectingCountry id="F3157652E7616F6C1199FEE3CDC8782C" box="[1015,1177,293,318]" name="Vanuatu" pageId="14" pageNumber="538">New Hebrides</collectingCountry>
in having very similar copulatory organs (
<figureCitation id="13392A47E7616F6C131EFE8DCB167873" box="[368,583,329,354]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="14" pageNumber="538">Figs 33–35, 38–39</figureCitation>
; cf.
<bibRefCitation id="EF934B33E7616F6C1019FE8DCAC77870" author="Platnick, N. I. &amp; Bonaldo, A. B." box="[631,918,329,354]" pageId="14" pageNumber="538" pagination="13 - 16" refId="ref12358" refString="Platnick, N. I. &amp; Bonaldo, A. B. (1995) On the spider genus Hebrithele (Araneae, Miturgidae). Journal of Arachnology, 23, 13 - 16." type="journal article" year="1995">Platnick &amp; Bonaldo 1995</bibRefCitation>
and
<figureCitation id="13392A47E7616F6C11A3FE8DCD717873" box="[973,1056,329,354]" captionStart="FIGURES 40–44" captionStartId="17.[151,263,1890,1913]" captionTargetBox="[152,1436,203,1859]" captionTargetId="figure@17.[151,1436,203,1859]" captionTargetPageId="17" captionText="FIGURES 40–44. Systaria spp., copulatory organs. 40–43 Systaria elberti (Strand 1913) from Indonesia, Lombok Island (40– 41 holotype male, 42–43 female), 44 Systaria insulana (Rainbow 1809), male from New Hebrides. 40–41, 44 Left palp (40 ventral, 41, 44 retrolateral). 42 Epigyne, ventral. 43 Vulva, dorsal. For explanations of a, b see diagnosis (44 modified from Platnick &amp; Bonaldo 1995, not to scale). Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606656/files/figure.png" pageId="14" pageNumber="538">Fig. 44</figureCitation>
), but may be distinguished by the male RTA with wider base (=a) and shorter distal part (=b, best seen in retrolateral view,
<figureCitation id="13392A47E7616F6C16EFFEA8CC537897" box="[1153,1282,364,389]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="14" pageNumber="538">Figs 34, 36</figureCitation>
; with a/b=
<quantity id="4CFA9B27E7616F6C1714FEA8C9FC78BB" metricMagnitude="-2" metricUnit="m" metricValue="4.064" pageId="14" pageNumber="538" unit="in" value="1.6">1.6 in</quantity>
<taxonomicName id="4C024D41E7616F6C12DAFE55C86378BA" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[180,306,401,424]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="B976EAD0E7616F6C12DAFE55C86378BA" box="[180,306,401,424]" italics="true" pageId="14" pageNumber="538">S. princesa</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E7616F6C1357FE54C8FD78BB" bold="true" box="[313,428,400,425]" pageId="14" pageNumber="538">
<taxonomicNameLabel id="A24557ABE7616F6C1357FE54C8FD78BB" box="[313,428,400,425]" pageId="14" pageNumber="538" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
; a/b=
<quantity id="4CFA9B27E7616F6C139CFE54CB4978BB" box="[498,536,400,425]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="538" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C024D41E7616F6C104EFE55CBCD78BB" baseAuthorityName="Rainbow" baseAuthorityYear="1901" box="[544,668,400,425]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B976EAD0E7616F6C104EFE55CBCD78BB" box="[544,668,400,425]" italics="true" pageId="14" pageNumber="538">S. insulana</emphasis>
</taxonomicName>
; cf.
<figureCitation id="13392A47E7616F6C10BEFE54CA7078BA" box="[720,801,400,425]" captionStart="FIGURES 40–44" captionStartId="17.[151,263,1890,1913]" captionTargetBox="[152,1436,203,1859]" captionTargetId="figure@17.[151,1436,203,1859]" captionTargetPageId="17" captionText="FIGURES 40–44. Systaria spp., copulatory organs. 40–43 Systaria elberti (Strand 1913) from Indonesia, Lombok Island (40– 41 holotype male, 42–43 female), 44 Systaria insulana (Rainbow 1809), male from New Hebrides. 40–41, 44 Left palp (40 ventral, 41, 44 retrolateral). 42 Epigyne, ventral. 43 Vulva, dorsal. For explanations of a, b see diagnosis (44 modified from Platnick &amp; Bonaldo 1995, not to scale). Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606656/files/figure.png" pageId="14" pageNumber="538">Fig. 44</figureCitation>
) and the more distinctly arched atrial margin, especially medially protruding more posteriorly and more widely separated and slightly larger spermathecae. Moreover, both sexes of the new species have more lateral femoral spines (males: p3, r4; females p2, r3) of legs I and II especially in females (less in males [p2–3, r1–3] and none in females in
<taxonomicName id="4C024D41E7616F6C1126FE3FCA957B01" baseAuthorityName="Rainbow" baseAuthorityYear="1901" box="[840,964,506,531]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B976EAD0E7616F6C1126FE3FCA957B01" box="[840,964,506,531]" italics="true" pageId="14" pageNumber="538">S. insulana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7616F6C12A9FDD9CA4D7F0A" pageId="14" pageNumber="538" type="description">
<paragraph id="8BBD36C2E7616F6C12A9FDD9CA6C7AF2" blockId="14.[151,1437,151,1773]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9FDD9C8087B24" bold="true" box="[199,345,541,566]" pageId="14" pageNumber="538">Description.</emphasis>
<emphasis id="B976EAD0E7616F6C1308FDD9C8F17B24" box="[358,416,541,566]" italics="true" pageId="14" pageNumber="538">Male</emphasis>
(
<typeStatus id="54B98860E7616F6C13D8FDD9CB4E7B24" box="[438,543,541,566]" pageId="14" pageNumber="538" type="holotype">holotype</typeStatus>
): prosoma length 4.1, prosoma width 3.6, anterior width of prosoma 1.6, opisthosoma length 5.0, opisthosoma width 2.4. Eyes: AME 0.25, ALE 0.19, PME 0.16, PLE 0.20, AME–AME 0.05, AME–ALE 0.05, PME–PME 0.25, PME–PLE 0.22, AME–PME 0.13, ALE–PLE 0.02. Cheliceral furrow with 3 promarginal and 3 retromarginal teeth. Spination: Palp: femur p001, d012, r001; legs (IV lacking): femur I– II p0111, d111, r1111, III p1111, d111, r1111; patella I
<emphasis id="B976EAD0E7616F6C1097FD68CA567BD7" bold="true" box="[761,775,684,709]" pageId="14" pageNumber="538">–</emphasis>
III 000; tibia I–II v222, III p11, d10, r11, v222; metatarsus I
<emphasis id="B976EAD0E7616F6C12CEFD0BC9FF7BFA" bold="true" box="[160,174,719,744]" pageId="14" pageNumber="538">–</emphasis>
II v220, III p122, r112, v222. Leg formula: 123. Measurements of palp and legs: Palp 4.7 (1.8, 0.7, 0.8, -, 1.4), I 14.2 (3.9, 1.7, 3.6, 3.3, 1.7), II 12.9 (3.5, 1.7, 3.2, 3.0, 1.5), III 11.6 (3.3, 1.4, 2.7, 2.9, 1.3). Palp as in diagnosis (
<figureCitation id="13392A47E7616F6C12F1FCD2C8737A3D" box="[159,290,790,815]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="14" pageNumber="538">Figs 33–36</figureCitation>
). Cymbium distinctly longer than tibia. RTA arising distally from tibia with broad, roughly rectangular base in retrolateral view, its dorsal margin serrated, distal tip hooked. Embolus arising in 8-o’clock-position from tegulum, slightly bent, narrowing from broad base to narrow apical part. Tip situated close to membranous conductor in 12- to 12.30-o’clock-position. Broad part of spermophor visible in retrolateral and proximal part of tegulum. Colouration. Yellowish-brown without pattern. Fovea, eye region, chelicerae, labium gnathocoxae, sternum, cymbium and posterior half of opisthosoma darker.
</paragraph>
<paragraph id="8BBD36C2E7616F6C12A9FC2FCD397CBF" blockId="14.[151,1437,151,1773]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9FC2FC84D7D16" box="[199,284,1003,1028]" italics="true" pageId="14" pageNumber="538">Female</emphasis>
(
<typeStatus id="54B98860E7616F6C1347FC2FC8C17D11" box="[297,400,1003,1027]" pageId="14" pageNumber="538" type="paratype">paratype</typeStatus>
): prosoma length 5.0, prosoma width 3.5, anterior width of prosoma 2.2, opisthosoma length 6.6, opisthosoma width 3.8. Eyes: AME 0.22, ALE 0.20, PME 0.17, PLE 0.20, AME–AME 0.06, AME–ALE 0.09, PME–PME 0.28, PME–PLE 0.28, AME–PME 0.19, ALE–PLE 0.07. Cheliceral furrow with 3 promarginal and 3 retromarginal teeth. Spination: Palp: femur p001(0), d012, r001, patella p1, tibia p21, d11, metatarsus p21, r10, v022; legs: femur I–II p011, d111, r111, III p1111, d111, r1111, IV p1111, d111, r01(0)1; patella I–IV 000; tibia I v220, II v110, III–IV p11, d10, r11, v222; metatarsus I–II v200, III p122, r112, v222, IV p122, r122, v222. Leg formula: 4123. Measurements of palp and legs: Palp 5.2 (1.7, 0.8, 1.1, -, 1.6), I 14.2 (4.1, 1.8, 3.7, 3.0, 1.6), II 13.5 (3.9, 1.9, 3.4, 2.8, 1.5), III 11.5 (3.3, 1.4, 2.6, 2.8, 1.4), IV 16.1 (4.4, 1.9, 4.1, 4.1, 1.6). Copulatory organ as in diagnosis (
<figureCitation id="13392A47E7616F6C137DFAC2C8CB7C0D" box="[275,410,1286,1311]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="14" pageNumber="538">Figs 38–39</figureCitation>
). Epigynal field wider than long, transverse oval, with 2 pairs of slit sensilla and small separate anterior bands. Anterior margin of atrium bipartite, arched. Sclerotised spheres (~30) visible through cuticle in ventral view. Copulatory openings located laterally. Internal duct system with huge oval bursae, copulatory ducts recognisable postero-laterally behind bursae. Connecting duct narrow, spermathecae separated by almost one of their diameter, fertilisation ducts antero-laterad. Colouration. As in male.
</paragraph>
<paragraph id="8BBD36C2E7616F6C12A9FA7CCA4D7F0A" blockId="14.[151,1437,151,1773]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9FA7CC86D7CC3" bold="true" box="[199,316,1464,1489]" pageId="14" pageNumber="538">Variation.</emphasis>
Males (n=1) with prosoma length 3.2, opisthosoma length 4.3; females (n=4) with prosoma length 4.0–5.5, opisthosoma length 5.2–6.7. RTA tip may be more distinctly pointed in ventral view (
<figureCitation id="13392A47E7616F6C16ABFA1FCC467CE6" box="[1221,1303,1499,1524]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="14" pageNumber="538">Fig. 35</figureCitation>
) and dorsal margin of RTA less serrated in retrolateral view (
<figureCitation id="13392A47E7616F6C10D5FA3BCA5E7F0A" box="[699,783,1535,1560]" captionStart="FIGURES 33–39" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="14" pageNumber="538">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7616F6C12A9F9E6C8067FFF" pageId="14" pageNumber="538" type="distribution">
<paragraph id="8BBD36C2E7616F6C12A9F9E6CD1B7F29" blockId="14.[151,1437,151,1773]" box="[199,1098,1570,1595]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9F9E6C8317F29" bold="true" box="[199,352,1570,1595]" pageId="14" pageNumber="538">Distribution.</emphasis>
Known from several caves close to the
<typeStatus id="54B98860E7616F6C114EF9E7CA017F29" box="[800,848,1571,1595]" pageId="14" pageNumber="538">type</typeStatus>
locality (
<figureCitation id="13392A47E7616F6C11D4F9E6CD6F7F29" box="[954,1086,1570,1595]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="14" pageNumber="538">Figs 45–46</figureCitation>
).
</paragraph>
<paragraph id="8BBD36C2E7616F6C12A9F982C8067FFF" blockId="14.[151,1437,151,1773]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12A9F982C8D77F4D" bold="true" box="[199,390,1606,1631]" pageId="14" pageNumber="538">Natural history.</emphasis>
All specimens have been recorded from the dark parts of the cave system. Along with adult and subadult females as well as juveniles the following organisms have been found:
<taxonomicName id="4C024D41E7616F6C1625F9ADC84B7FB7" authority="Jager 2018" authorityName="Jager" authorityYear="2018" class="Arachnida" family="Sparassidae" genus="Heteropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="simoneallmannae">
<emphasis id="B976EAD0E7616F6C1625F9ADCCCD7F90" box="[1099,1436,1641,1666]" italics="true" pageId="14" pageNumber="538">Heteropoda simoneallmannae</emphasis>
<bibRefCitation id="EF934B33E7616F6C12F9F94AC84B7FB7" author="Zhong, Y. &amp; Jager, P. &amp; Chen, J. &amp; Liu, J." box="[151,282,1677,1702]" pageId="14" pageNumber="538" pagination="328 - 346" refId="ref12663" refString="Zhong, Y., Jager, P., Chen, J. &amp; Liu, J. (2018) Taxonomic review of the Sinopoda okinawana - group (Araneae: Sparassidae) in China. Zootaxa, 4388 (3), 328 - 346. https: // doi. org / 10.11646 / zootaxa. 4388.3.2" type="journal article" year="2018">Jäger 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C024D41E7616F6C1342F949C8E07FB4" box="[300,433,1677,1702]" class="Arachnida" family="Sparassidae" genus="Heteropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAD0E7616F6C1342F949C8E07FB4" box="[300,433,1677,1702]" italics="true" pageId="14" pageNumber="538">Heteropoda</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C024D41E7616F6C139DF949CB307FB4" box="[499,609,1677,1702]" class="Arachnida" family="Pholcidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
gen. sp. 1 &amp; 2,
<taxonomicName id="4C024D41E7616F6C1155F949CD7B7FB4" authority=", Psechrus" authorityName="Psechrus" box="[827,1066,1677,1702]" class="Arachnida" family="Pisauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">
Pisauridae,
<emphasis id="B976EAD0E7616F6C11AAF949CD7B7FB4" box="[964,1066,1677,1702]" italics="true" pageId="14" pageNumber="538">Psechrus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C024D41E7616F6C1602F949CD8C7FB4" box="[1132,1245,1677,1702]" class="Arachnida" family="Salticidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">Salticidae</taxonomicName>
,
<taxonomicName id="4C024D41E7616F6C1681F949CCC97FB4" box="[1263,1432,1677,1702]" class="Arachnida" family="Theraphosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">Theraphosidae</taxonomicName>
: (Selenocosmiinae),
<taxonomicName id="4C024D41E7616F6C1317F974CA5D7FDB" authority=", Opiliones (Laniatores)" box="[377,780,1712,1737]" class="Arachnida" family="Uloboridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">Uloboridae, Opiliones (Laniatores)</taxonomicName>
, Acari (Trombidiformes) and Isopoda (Agnelli, personal communication).
</paragraph>
</subSubSection>
</treatment>
</document>
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