470 lines
60 KiB
XML
470 lines
60 KiB
XML
<document id="63666ECEF9FF84E8F7DE15F129570C39" ID-DOI="10.5281/zenodo.173907" ID-GBIF-Dataset="29f50131-188d-4263-a803-f6fab5f75be9" ID-ISSN="11755326" ID-Zenodo-Dep="173907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459786406446" checkinUser="plazi" docAuthor="Aplin, Kenneth P., Fitch, Alison J. & King, Dennis J." docDate="2006" docId="03A587F0FF83FF94FEF2FE15A7FCFE64" docLanguage="en" docName="zt01313p038.pdf" docOrigin="Zootaxa 1313" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Varanus bushi Aplin, Fitch & King, 2006, sp. nov." docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="FF9CFF88FF94FF88FFFAFFE9A631FF9A" masterDocTitle="A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species" masterLastPageNumber="38" masterPageNumber="1" pageNumber="24" updateTime="1698218667939" updateUser="plazi">
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<mods:titleInfo id="EF82ACB63B0313B44035E8162DBD3CD7">
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<mods:title id="71B37ECEF98C854652AE94B3876EA176">A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species</mods:title>
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<mods:name id="00CC9BCCCD71E1D35EB9B5A82C55B4DA" type="personal">
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<mods:roleTerm id="DD9ED09077D0CAF80B3065A78397D446">Author</mods:roleTerm>
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<mods:namePart id="A4A798BA26DB846F0699934B6B7526B1">Aplin, Kenneth P.</mods:namePart>
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<mods:roleTerm id="D78660EE62D34C46F2C166533561EEEA">Author</mods:roleTerm>
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<mods:namePart id="C0829B2C8FC06EA54EDFBD5D3120D934">Fitch, Alison J.</mods:namePart>
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<mods:name id="9BB2C69C115F2F0A62E72C9DFAD68E2E" type="personal">
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<mods:roleTerm id="6E9867466B368D8AA6DBFAE58CA671EC">Author</mods:roleTerm>
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<mods:namePart id="DF5A281BE06C39403D405CE21EA73CC3">King, Dennis J.</mods:namePart>
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<mods:typeOfResource id="8A02CDF1C864FB1D691FA0CEFAD2BF21">text</mods:typeOfResource>
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<mods:title id="9C38B68D9D27240DB8F9788184C88C78">Zootaxa</mods:title>
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<mods:part id="03940996A1EFD8FF4C02533C9FCF22AE">
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<mods:date id="3630B6F00EAE2DE7CEC121BB5E20C194">2006</mods:date>
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<mods:detail id="56E2ED1EE52C159A0F08492956CE2D64" type="volume">
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<mods:number id="F9B11E20BB72AC37F50246428089B9C4">1313</mods:number>
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<mods:classification id="9D8053A4183598238B47DF3E6A061474">journal article</mods:classification>
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<mods:identifier id="5E6A9CDB7D67839386E75462ABF97ECF" type="DOI">10.5281/zenodo.173907</mods:identifier>
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<mods:identifier id="FDD67DA12491B029DF3CE7553CF72D17" type="GBIF-Dataset">29f50131-188d-4263-a803-f6fab5f75be9</mods:identifier>
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<mods:identifier id="B43938AE2E9CA218BE4F911650E7C866" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="E11FD6A84AE89C55BCE1B456703C1BB1" type="Zenodo-Dep">173907</mods:identifier>
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<treatment id="03A587F0FF83FF94FEF2FE15A7FCFE64" ID-DOI="http://doi.org/10.5281/zenodo.6262551" ID-GBIF-Taxon="119349185" ID-Zenodo-Dep="6262551" LSID="urn:lsid:plazi:treatment:03A587F0FF83FF94FEF2FE15A7FCFE64" httpUri="http://treatment.plazi.org/id/03A587F0FF83FF94FEF2FE15A7FCFE64" lastPageId="28" lastPageNumber="29" pageId="23" pageNumber="24">
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<subSubSection id="C316656DFF83FF9FFEF2FE15A781FDA3" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BB336E6FF83FF9FFEF2FE15A42DFD8F" blockId="23.[264,540,507,569]" box="[264,540,507,534]" pageId="23" pageNumber="24">
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<heading id="D0FB818AFF83FF9FFEF2FE15A42DFD8F" bold="true" box="[264,540,507,534]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B978EAF4FF83FF9FFEF2FE15A42DFD8F" bold="true" box="[264,540,507,534]" pageId="23" pageNumber="24">
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<taxonomicName id="4C0C4D65FF83FF9FFEF2FE15A786FD8C" box="[264,439,508,534]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
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<emphasis id="B978EAF4FF83FF9FFEF2FE15A786FD8C" bold="true" box="[264,439,508,534]" italics="true" pageId="23" pageNumber="24">Varanus bushi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24B578FFF83FF9FFE44FE12A42DFD8F" box="[446,540,507,533]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB336E6FF83FF9FFEF2FDF6A781FDA3" blockId="23.[264,540,507,569]" box="[264,432,543,569]" pageId="23" pageNumber="24">
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<figureCitation id="13372A63FF83FF9FFEF2FDF6A747FDA3" box="[264,374,543,569]" captionStart="FIGURE 1" captionStartId="2.[264,368,1268,1292]" captionTargetBox="[267,1319,526,1244]" captionTargetId="figure@2.[267,1319,526,1244]" captionTargetPageId="2" captionText="FIGURE 1. An adult Varanus bushi sp. nov. from near Roebourne, photographed in life by Robert BrowneCooper." httpUri="https://zenodo.org/record/173908/files/figure.png" pageId="23" pageNumber="24">Figures 1</figureCitation>
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,
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<figureCitation id="13372A63FF83FF9FFE7EFDF6A781FDA3" box="[388,432,543,569]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="8.[264,368,1242,1266]" captionStartId-1="13.[264,368,995,1019]" captionStartId-2="13.[264,367,1691,1715]" captionStartId-3="14.[264,368,790,814]" captionTargetBox-0="[280,1304,751,1234]" captionTargetBox-1="[294,1294,442,964]" captionTargetBox-2="[312,1294,1163,1660]" captionTargetBox-3="[277,1309,319,771]" captionTargetId-0="figure@8.[275,1313,748,1236]" captionTargetId-1="figure@13.[267,1319,442,979]" captionTargetId-2="figure@13.[267,1319,1138,1673]" captionTargetId-3="figure@14.[265,1321,308,784]" captionTargetPageId-0="8" captionTargetPageId-1="13" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 3. Details of scalation on dorsal midbody of representative specimens of three Varanus species. (a) WAM R 135340 V. b u s h i sp. nov. Paratype Ψ; (b) WAM R 75792 V. gilleni Ψ; c) WAM R 135601 V. caudolineatus ♂." captionText-1="FIGURE 4. Dorsal views of representative specimens of three Varanus species. (a) WAM R 125521 V. b u s h i sp. nov. Holotype ♂; (b) WAM R 135340 V. bushi sp. nov. Paratype Ψ; (c) WAM R 75792 V. gilleni Ψ; (d) WAM R 135601 V. caudolineatus ♂; (e) WAM R 138950 V. caudolineatus ♂." captionText-2="FIGURE 5. Ventral views of representative specimens of three Varanus species. Same individuals as Figure 3." captionText-3="FIGURE 6. Dorsal views of head and neck of representative specimens of three Varanus species. Same individuals as Figure 3." httpUri-0="https://zenodo.org/record/173910/files/figure.png" httpUri-1="https://zenodo.org/record/173911/files/figure.png" httpUri-2="https://zenodo.org/record/173912/files/figure.png" httpUri-3="https://zenodo.org/record/173913/files/figure.png" pageId="23" pageNumber="24">3–6</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C316656DFF83FF9FFEF2FD87A2A8FCE4" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8BB336E6FF83FF9FFEF2FD87A7C8FC9C" blockId="23.[264,1324,622,1894]" pageId="23" pageNumber="24">
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<typeStatus id="54B78844FF83FF9FFEF2FD87A74BFD12" box="[264,378,622,648]" pageId="23" pageNumber="24" type="holotype">
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<emphasis id="B978EAF4FF83FF9FFEF2FD87A74BFD12" bold="true" box="[264,378,622,648]" pageId="23" pageNumber="24">Holotype</emphasis>
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</typeStatus>
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: Western Australian Museum R108999, adult male from Marandoo, Western
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<collectingCountry id="F31B7676FF83FF9FFEF2FD73A742FD2E" box="[264,371,666,692]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
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in 22
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<geoCoordinate id="EE385021FF83FF9FFE40FD7FA43AFD2E" box="[442,523,662,692]" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="-0.6166667">0 37’ S</geoCoordinate>
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118
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<geoCoordinate id="EE385021FF83FF9FFDC5FD7FA4A4FD2E" box="[575,661,662,692]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="0.13333334">0 08’ E</geoCoordinate>
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. Collected on
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<date id="FFB21026FF83FF9FFCB9FD73A5D2FD2E" box="[835,995,666,692]" pageId="23" pageNumber="24" value="1991-06-20">20 June 1991</date>
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by Greg Harold. Specimen fixed in 10% formalin then stored in 70% ethanol. Liver sample stored in 80 0 C ultrafreeze at
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<collectionCode id="ED1DAE23FF83FF9FFE53FD05A7C4FC9C" LSID="urn:lsid:biocol.org:col:34981" box="[425,501,748,774]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="24">WAM</collectionCode>
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.
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</paragraph>
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<paragraph id="8BB336E6FF83FF9FFEC2FCFDA2A8FCE4" blockId="23.[264,1324,622,1894]" pageId="23" pageNumber="24">
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<typeStatus id="54B78844FF83FF9FFEC2FCFDA784FCB4" box="[312,437,788,814]" pageId="23" pageNumber="24" type="paratype">
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<emphasis id="B978EAF4FF83FF9FFEC2FCFDA784FCB4" bold="true" box="[312,437,788,814]" pageId="23" pageNumber="24">Paratypes</emphasis>
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</typeStatus>
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: Three additional specimens from Marandoo:
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<collectionCode id="ED1DAE23FF83FF9FFC29FCFDA22AFCB4" LSID="urn:lsid:biocol.org:col:34981" box="[979,1051,788,814]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="24">WAM</collectionCode>
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R54230, an adult male collected on
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<date id="FFB21026FF83FF9FFE66FCD5A455FCCC" box="[412,612,828,854]" pageId="23" pageNumber="24" value="1976-10-25">25 October 1976</date>
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;
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<collectionCode id="ED1DAE23FF83FF9FFD95FCD5A486FCCC" LSID="urn:lsid:biocol.org:col:34981" box="[623,695,828,854]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="24">WAM</collectionCode>
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R56834, an adult female collected in
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<date id="FFB21026FF83FF9FFB8AFCD5A2C2FCCC" box="[1136,1267,828,854]" pageId="23" pageNumber="24" value="1977-04">April 1977</date>
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; and
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<collectionCode id="ED1DAE23FF83FF9FFEF2FC8DA77EFCE4" LSID="urn:lsid:biocol.org:col:34981" box="[264,335,868,894]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="24">WAM</collectionCode>
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R62171 a juvenile of undetermined sex collected on
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<date id="FFB21026FF83FF9FFC3BFC8DA2A5FCE4" box="[961,1172,868,894]" pageId="23" pageNumber="24" value="1979-02-28">28 February 1979</date>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C316656DFF83FF9FFEC2FC65A4CBFA64" pageId="23" pageNumber="24" type="diagnosis">
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<paragraph id="8BB336E6FF83FF9FFEC2FC65A4CBFA64" blockId="23.[264,1324,622,1894]" pageId="23" pageNumber="24">
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<emphasis id="B978EAF4FF83FF9FFEC2FC65A781FC3C" bold="true" box="[312,432,908,934]" pageId="23" pageNumber="24">Diagnosis</emphasis>
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: A smallbodied member of the subgenus
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<taxonomicName id="4C0C4D65FF83FF9FFC5DFC64A235FC3C" box="[935,1028,909,934]" class="Squamata" family="Varanidae" genus="Odatria" higherTaxonomySource="CoL" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
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<emphasis id="B978EAF4FF83FF9FFC5DFC64A235FC3C" box="[935,1028,909,934]" italics="true" pageId="23" pageNumber="24">Odatria</emphasis>
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</taxonomicName>
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distinguished from most others by the combination of a longitudinally striped and only moderately spinose tail, unkeeled head and body scales, nonoverlapping ventral primary scales, and an absence of longitudinal streaks on throat. Distinguished from
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<emphasis id="B978EAF4FF83FF9FFC9EFBECA5FBFB84" box="[868,970,1029,1054]" italics="true" pageId="23" pageNumber="24">V. g i l l e n i</emphasis>
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by its slightly lesser average size, more elongate dorsal scales, more densely spotted venter and more irregularly spotted dorsum, less prominent linear patterning on the head and neck, and its more numerous presacral vertebrae, pedal subdigital lamellae and ventral scales. Males are further distinguished from
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<taxonomicName id="4C0C4D65FF83FF9FFDA7FB4CA4FFFB24" box="[605,718,1189,1214]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="gilleni">
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<emphasis id="B978EAF4FF83FF9FFDA7FB4CA4FFFB24" box="[605,718,1189,1214]" italics="true" pageId="23" pageNumber="24">V. gilleni</emphasis>
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</taxonomicName>
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by having hemipenes with an undivided inner hemibaculum. Distinguished from
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<taxonomicName id="4C0C4D65FF83FF9FFD4BFB24A546FB7C" box="[689,887,1229,1254]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caudolineatus">
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<emphasis id="B978EAF4FF83FF9FFD4BFB24A546FB7C" box="[689,887,1229,1254]" italics="true" pageId="23" pageNumber="24">V. caudolineatus</emphasis>
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</taxonomicName>
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by its slightly greater average and maximum size, proportionally shorter fore and hindlimbs, more elongate snout, higher average midbody and ventral scale counts, higher average subdigital lamellar counts on pes, more finely scaled and less rugose proximal portion of the tail, more numerous presacral vertebrae, less conspicuously spotted head, more orderly alignment of dorsal pattern into transverse rows, and presence of transverse bands on the basal onethird of the tail. Males are further distinguished from
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<taxonomicName id="4C0C4D65FF83FF9FFD03FA54A58AFA4C" box="[761,955,1469,1494]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caudolineatus">
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<emphasis id="B978EAF4FF83FF9FFD03FA54A58AFA4C" box="[761,955,1469,1494]" italics="true" pageId="23" pageNumber="24">V. caudolineatus</emphasis>
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</taxonomicName>
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by having a shorter hemipenis with more numerous papillose distal frills.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C316656DFF83FF9FFEC2F9E5A5AEF9EC" pageId="23" pageNumber="24" type="etymology">
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<paragraph id="8BB336E6FF83FF9FFEC2F9E5A5AEF9EC" blockId="23.[264,1324,622,1894]" pageId="23" pageNumber="24">
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<emphasis id="B978EAF4FF83FF9FFEC2F9E5A78EF9BC" bold="true" box="[312,447,1548,1574]" pageId="23" pageNumber="24">Etymology</emphasis>
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: We take pleasure in naming this species after naturalist and educator Brian Bush who has contributed enormously to our knowledge of the herpetofauna of Western
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<collectingCountry id="F31B7676FF83FF9FFE94F9B5A7EBF9EC" box="[366,474,1628,1654]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
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and of the Pilbara region in particular.
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</paragraph>
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</subSubSection>
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<subSubSection id="C316656DFF83FF90FEC2F96DA7A1FEAC" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="distribution">
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<paragraph id="8BB336E6FF83FF90FEC2F96DA7A1FEAC" blockId="23.[264,1324,622,1894]" lastBlockId="24.[264,1325,284,1110]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<emphasis id="B978EAF4FF83FF9FFEC2F96DA4A0F904" bold="true" box="[312,657,1668,1694]" pageId="23" pageNumber="24">Distribution and sympatry</emphasis>
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: Endemic to the Pilbara region of Western
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<collectingCountry id="F31B7676FF83FF9FFB42F96DA316F904" box="[1208,1319,1668,1694]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
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, northwest to the vicinity of Cooya Poonya and Tambrey, southwest to Mt Brockman and Mt Tom Price in the Hamersley Range, southeast to Mt Whaleback and northeast to Marillana and Hope Downs (
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<figureCitation id="13372A63FF83FF9FFD7EF915A4E7F88C" box="[644,726,1788,1814]" captionStart="FIGURE 9" captionStartId="24.[264,368,1779,1803]" captionTargetBox="[465,1125,1158,1761]" captionTargetId="figure@24.[465,1125,1158,1766]" captionTargetPageId="24" captionText="FIGURE 9. Distribution of the three Varanus species based on examined materials and type localities. The approximate wider distribution of V. g i l l e n i is indicated by the dashed line. Symbols: Stars — V. caudolineatus; Triangles — V. gilleni; Circles — V. b u s h i sp. nov.; Circles with cross — s V. caudolineatus and V. b u s h i sp. nov. in local sympatry; Circles with dot – type localities." httpUri="https://zenodo.org/record/173916/files/figure.png" pageId="23" pageNumber="24">Fig. 9</figureCitation>
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||
). Two instances of regional sympatry with
|
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<taxonomicName id="4C0C4D65FF83FF9FFAEFF914A79CF8A4" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caudolineatus">
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<emphasis id="B978EAF4FF83FF9FFAEFF914A79CF8A4" italics="true" pageId="23" pageNumber="24">V. caudolineatus</emphasis>
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</taxonomicName>
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are recorded—at West Angelas and at Hope Downs, both on the southern margin of the Pilbara Uplands. Details of the relevant specimens are provided in the Discussion.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C316656DFF8CFF92FEC2FEAAA435FC8C" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BB336E6FF8CFF90FEC2FEAAA757FE34" blockId="24.[264,1325,284,1110]" pageId="24" pageNumber="25">
|
||
<emphasis id="B978EAF4FF8CFF90FEC2FEAAA45CFEC7" bold="true" box="[312,621,323,349]" pageId="24" pageNumber="25">
|
||
Description of
|
||
<typeStatus id="54B78844FF8CFF90FDF8FEAAA45CFEC7" box="[514,621,323,349]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: Adult male measuring SVL
|
||
<quantity id="4CF49B03FF8CFF90FC02FEADA254FEC4" box="[1016,1125,324,350]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="24" pageNumber="25" unit="mm" value="145.0">145 mm</quantity>
|
||
, Tail
|
||
<quantity id="4CF49B03FF8CFF90FB42FEADA317FEC4" box="[1208,1318,324,350]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="25" unit="mm" value="190.0">190 mm</quantity>
|
||
, Forelimb
|
||
<quantity id="4CF49B03FF8CFF90FE84FE85A7E7FE1C" box="[382,470,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="24" pageNumber="25" unit="mm" value="32.0">32 mm</quantity>
|
||
,
|
||
<collectingCountry id="F31B7676FF8CFF90FE1FFE85A411FE1C" box="[485,544,364,390]" name="India" pageId="24" pageNumber="25">Hind</collectingCountry>
|
||
limb
|
||
<quantity id="4CF49B03FF8CFF90FD91FE85A4F5FE1C" box="[619,708,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="24" pageNumber="25" unit="mm" value="43.0">43 mm</quantity>
|
||
, Head and neck
|
||
<quantity id="4CF49B03FF8CFF90FC68FE85A5DAFE1C" box="[914,1003,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="mm" value="50.0">50 mm</quantity>
|
||
. Both hemipenes are fully everted.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8CFF90FEC2FE55A45DFE64" blockId="24.[264,1325,284,1110]" pageId="24" pageNumber="25">Vertebral column includes 30 presacral vertebrae. Phalangeal formula of manus 1.2.3.4.2; and of pes 3.4.3.2.1.</paragraph>
|
||
<paragraph id="8BB336E6FF8CFF90FEC2FDE5A297FDEC" blockId="24.[264,1325,284,1110]" pageId="24" pageNumber="25">
|
||
Head moderately depressed, depth at pineal organ
|
||
<quantity id="4CF49B03FF8CFF90FC6BFDE5A5C1FDBC" box="[913,1008,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="24" pageNumber="25" unit="mm" value="8.7">8.7 mm</quantity>
|
||
. Snout relatively elongate; lacking canthus rostralis. Nostril positioned
|
||
<quantity id="4CF49B03FF8CFF90FCD6FDDDA5BDFDD4" box="[812,908,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="24" pageNumber="25" unit="mm" value="6.8">6.8 mm</quantity>
|
||
from tip of snout;
|
||
<quantity id="4CF49B03FF8CFF90FB7CFDDDA2D6FDD4" box="[1158,1255,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="24" pageNumber="25" unit="mm" value="3.4">3.4 mm</quantity>
|
||
from anterior corner of eye. Ear aperture is vertically narrow and obliquely oriented.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8CFF90FEC2FD6DA48CFBCC" blockId="24.[264,1325,284,1110]" pageId="24" pageNumber="25">
|
||
Dorsal head scales unornamented; most are irregular polygons, subrounded in shape, and majority have a single scale organ. Supraocular scales are smaller than those on the rostrum, frontal and parietal regions. Granules are absent from head except in temporal region and on throat forward into genal groove. Dorsal primary scales on neck are raised but lack keels; all have scale organs. The dorsal primary scales on neck are rounded anteriorly but become more ovate posteriorly, merging smoothly with body scalation. Ventral primary scales on neck are small and almost round anteriorly, but become larger and progressively more elongate to rear; all ventral primary scales on neck are ringed laterally and posteriorly by small granules. Approximately one third of ventral neck scales bear scale organs, usually one per scale but not infrequently two or three. Gular fold is distinct, located
|
||
<quantity id="4CF49B03FF8CFF90FE34FBFDA425FBB4" box="[462,532,1044,1070]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="mm" value="5.0">5 mm</quantity>
|
||
forward of anterior base of forelimb, and consisting of 4 rows of small scales, all lacking scale organs.
|
||
</paragraph>
|
||
<caption id="DF73666EFF8CFF90FEF2F91AA2EDF8F1" httpUri="https://zenodo.org/record/173916/files/figure.png" pageId="24" pageNumber="25" targetBox="[465,1125,1158,1761]" targetPageId="24">
|
||
<paragraph id="8BB336E6FF8CFF90FEF2F91AA2EDF8F1" blockId="24.[264,1324,1779,1899]" pageId="24" pageNumber="25">
|
||
<emphasis id="B978EAF4FF8CFF90FEF2F91AA7BEF891" bold="true" box="[264,399,1779,1803]" pageId="24" pageNumber="25">FIGURE 9.</emphasis>
|
||
Distribution of the three
|
||
<taxonomicName id="4C0C4D65FF8CFF90FD41F91DA523F891" box="[699,786,1780,1803]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
|
||
<emphasis id="B978EAF4FF8CFF90FD41F91DA523F891" box="[699,786,1780,1803]" italics="true" pageId="24" pageNumber="25">Varanus</emphasis>
|
||
</taxonomicName>
|
||
species based on examined materials and type localities. The approximate wider distribution of
|
||
<emphasis id="B978EAF4FF8CFF90FCE9F8FDA55CF8B1" box="[787,877,1812,1835]" italics="true" pageId="24" pageNumber="25">V. g i l l e n i</emphasis>
|
||
is indicated by the dashed line. Symbols: Stars —
|
||
<taxonomicName id="4C0C4D65FF8CFF90FE9FF8DDA425F8D1" box="[357,532,1844,1867]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8CFF90FE9FF8DDA425F8D1" box="[357,532,1844,1867]" italics="true" pageId="24" pageNumber="25">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
; Triangles —
|
||
<taxonomicName id="4C0C4D65FF8CFF90FD54F8DDA53DF8D1" box="[686,780,1844,1867]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="gilleni">
|
||
<emphasis id="B978EAF4FF8CFF90FD54F8DDA53DF8D1" box="[686,780,1844,1867]" italics="true" pageId="24" pageNumber="25">V. gilleni</emphasis>
|
||
</taxonomicName>
|
||
; Circles —
|
||
<emphasis id="B978EAF4FF8CFF90FC77F8DDA5EFF8D1" box="[909,990,1844,1867]" italics="true" pageId="24" pageNumber="25">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8CFF90FC1DF8DAA20CF8D1" bold="true" box="[999,1085,1843,1867]" pageId="24" pageNumber="25">sp. nov.</emphasis>
|
||
; Circles with cross — s
|
||
<taxonomicName id="4C0C4D65FF8CFF90FEE3F8BDA7F8F8F1" box="[281,457,1876,1899]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8CFF90FEE3F8BDA7F8F8F1" box="[281,457,1876,1899]" italics="true" pageId="24" pageNumber="25">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B978EAF4FF8CFF90FE07F8BDA47CF8F1" box="[509,589,1876,1899]" italics="true" pageId="24" pageNumber="25">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8CFF90FDADF8BAA49CF8F1" bold="true" box="[599,685,1875,1899]" pageId="24" pageNumber="25">sp. nov.</emphasis>
|
||
in local sympatry; Circles with dot – type localities.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB336E6FF8DFF91FEC2FEF5A788FD5C" blockId="25.[264,1325,284,1870]" pageId="25" pageNumber="26">
|
||
Body scalation is relatively unspecialised. Dorsal primary scales are ovate, approximately twice as long as wide and lack keels; the majority have a single scale organ. Transverse scale rows are wellorganised on the dorsum but tend to subdivide on flanks. Ventral primary scales are approximately 50% longer than and twice as wide as dorsal primary scales. All are ringed laterally and posteriorly by small granules. Single scale organs are present on the majority of scales towards the flanks but only about 15% of scales located along the midventral area. Inguinal fold indistinct but located
|
||
<quantity id="4CF49B03FF8DFF91FB36FDE5A31AFDBC" box="[1228,1323,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
|
||
forward of the anterior base of hindlimb. Position is marked by a change in scale size and shape, those behind the fold and back to vent being smaller, rounded and not arranged in transverse rows, and by occurrence of incomplete transverse series that terminate before reaching flank.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8DFF91FEC2FD3DA2CAFB9C" blockId="25.[264,1325,284,1870]" pageId="25" pageNumber="26">Scales on the inner surface of the fore and hindlimbs are small and rounded while those on the outer surfaces are larger, more elongate and weakly keeled. Plantar surfaces of manus and pes with primary scales on small mounds of fine granules; all primary scales bear scale organs and some are pigmented. Subdigital surfaces with transverse lamellae made up of two or three rounded primary scales surrounded by granules. The apical lamella on each digit is single and more intensely pigmented. Longest digit (IV) of manus has 20 lamellae; longest digit (IV) of pes has 23. Claws on manus and pes are moderately large and darkly pigmented; all are laterally compressed and bear sharp, recurved tips.</paragraph>
|
||
<paragraph id="8BB336E6FF8DFF91FEC2FBFDA489FA0C" blockId="25.[264,1325,284,1870]" pageId="25" pageNumber="26">Tail is slightly wider than deep at base, becoming more rounded in crosssection towards tip; it lacks a dorsal keel or other obvious specialisation. Dorsal primary scales near tail base are elongate but otherwise unspecialised. Moving distally, the primary scales first become more elongate, then develop a distinct midline keel, and finally develop smaller lateral keels. Ventral primary scales also change from elongate and lacking keels near the tail base, to even longer and with a strong midline keel distally. The overall effect is that the tail feels relatively smooth near the base but becomes more distinctly rasplike distally. Circumferential scale counts decrease from 52 near tail base to 32 at one third of total length, then to 22 at twothirds of total length. Longitudinal scale counts at the same positions are 8, 7 and 6 rows per cm.</paragraph>
|
||
<paragraph id="8BB336E6FF8DFF91FEC2FA4DA784F9AC" blockId="25.[264,1325,284,1870]" pageId="25" pageNumber="26">The postcloacal scale cluster is relatively poorly developed and consists of two rows of modified but unpigmented scales. The outer row consists of six nodular scales of which the lateral four scales are distinctly spinose. The inner row consists of four smaller, bluntly pointed scales.</paragraph>
|
||
<paragraph id="8BB336E6FF8DFF91FEC2F9ADA7C7F934" blockId="25.[264,1325,284,1870]" pageId="25" pageNumber="26">
|
||
The hemipenis measures
|
||
<quantity id="4CF49B03FF8DFF91FD98F9ADA48DF9C4" box="[610,700,1604,1630]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="25" pageNumber="26" unit="mm" value="6.3">6.3 mm</quantity>
|
||
without the projecting hemibacula and 6.8 including these structures. The nude basal portion measures
|
||
<quantity id="4CF49B03FF8DFF91FCA5F985A58CF91C" box="[863,957,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="25" pageNumber="26" unit="mm" value="3.3">3.3 mm</quantity>
|
||
. Further details were provided in an earlier section.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8DFF92FEC2F955A435FC8C" blockId="25.[264,1325,284,1870]" lastBlockId="26.[264,1324,284,1870]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||
Ground colour of dorsum in preservative is a uniform palebrown from rostrum to base of tail, becoming greyer on the flanks. Patterning on the head includes an irregular mottling of dark brown on the dorsum and sides, tending towards longitudinal streaking on the occiput, and a subdued temporal stripe on each side, running from the posterior corner of the eye to above the ear. Irregular dark brown mottling is also present on the dorsum of the body, each ‘spot’ usually consisting of one or a few dark primary scales. The pattern is irregular on the anterior body but tends towards transverse alignment on the lower half of the back and above the hind limbs. The last three bars of this series are quite distinct and alternate with rows of small dark spots. The under surface of the throat, neck and body bears distinct pale grey spotting, each ‘spot’ usually comprised of two or three pigmented primary scales. These are most abundant on the throat and neck, and along the sides of the body, and less so in the midventral region. The insides of the limbs are similarly patterned. The proximal
|
||
<quantity id="4CF49B03FF8EFF92FDCBFDB5A4B6FDEC" box="[561,647,604,630]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="27" unit="mm" value="35.0">35 mm</quantity>
|
||
of the tail bears a series of eight more or less complete transverse bars. The next
|
||
<quantity id="4CF49B03FF8EFF92FDB8FD6DA4A9FD04" box="[578,664,644,670]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="26" pageNumber="27" unit="mm" value="38.0">38 mm</quantity>
|
||
bears a series of broken bars and irregular spots. The remainder of the tail supports a linear pattern consisting of five, more or less continuous dark stripes, each of which is one scale wide. The ventral surface of the tail is unpatterned throughout its length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316656DFF8EFF93FEC2FCCDA2DDFE1C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="materials_examined">
|
||
<paragraph id="8BB336E6FF8EFF92FEC2FCCDA5A7FC2C" blockId="26.[264,1324,284,1870]" pageId="26" pageNumber="27">
|
||
<emphasis id="B978EAF4FF8EFF92FEC2FCCDA534FCA4" bold="true" box="[312,773,804,830]" pageId="26" pageNumber="27">Variation among referred specimens</emphasis>
|
||
: Mensural and meristic data for the complete sample of
|
||
<emphasis id="B978EAF4FF8EFF92FE7EFCA5A7ECFCFF" box="[388,477,844,869]" italics="true" pageId="26" pageNumber="27">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8EFF92FE10FCA5A479FCFC" bold="true" box="[490,584,844,870]" pageId="26" pageNumber="27">sp. nov.</emphasis>
|
||
is presented in
|
||
<tableCitation id="C68E035DFF8EFF92FCFEFCA5A56DFCFC" box="[772,860,844,870]" captionStart="TABLE 1" captionStartId="10.[264,352,283,307]" captionTargetBox="[269,1320,564,1819]" captionTargetPageId="10" captionText="TABLE 1. Mensural and meristic data for each Varanus caudolineatus, V. gilleni and V. bushi sp. nov. Data are presented separately for each sex, and for the total sample including unsexed individuals. Analysis of Variance (ANOVA) results indicate extent of sexual dimorphism for each variable; NS indicates nonsignificant contrasts with p> 0.05 (i. e. not significantly dimorphic). Abbreviations for variables are explained in Methods." httpUri="http://table.plazi.org/id/DF73666EFF9EFF82FEF2FEF2A579FE27" pageId="26" pageNumber="27" tableUuid="DF73666EFF9EFF82FEF2FEF2A579FE27">Table 1</tableCitation>
|
||
; locality details for referred specimens are given in Appendix II. The
|
||
<typeStatus id="54B78844FF8EFF92FD8EFC9DA4EAFC14" box="[628,731,884,910]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
is the largest male specimen of
|
||
<emphasis id="B978EAF4FF8EFF92FBA7FC9DA286FC17" box="[1117,1207,884,909]" italics="true" pageId="26" pageNumber="27">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8EFF92FB39FC9DA313FC14" bold="true" box="[1219,1314,884,910]" pageId="26" pageNumber="27">sp. nov.</emphasis>
|
||
;
|
||
<collectionCode id="ED1DAE23FF8EFF92FEF2FC75A70EFC2C" box="[264,319,924,950]" pageId="26" pageNumber="27">SVL</collectionCode>
|
||
of the largest female (
|
||
<collectionCode id="ED1DAE23FF8EFF92FDBFFC75A4BEFC2C" LSID="urn:lsid:biocol.org:col:34981" box="[581,655,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="26" pageNumber="27">WAM</collectionCode>
|
||
R135340) is
|
||
<quantity id="4CF49B03FF8EFF92FCD7FC75A5A2FC2C" box="[813,915,924,950]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="26" pageNumber="27" unit="mm" value="140.0">140 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8EFF92FEC2FC2DA430FA0C" blockId="26.[264,1324,284,1870]" pageId="26" pageNumber="27">
|
||
Dorsal patterning in most individuals is similar to that of the
|
||
<typeStatus id="54B78844FF8EFF92FBBAFC2DA29BFC44" box="[1088,1194,964,990]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
, with fine spotting on the anterior body, conspicuous banding only on the rear of the body and on the proximal segment of the tail, and longitudinal striping of the distal tail. In some individuals (e.g.
|
||
<collectionCode id="ED1DAE23FF8EFF92FE37FBD5A424FBCC" LSID="urn:lsid:biocol.org:col:34981" box="[461,533,1084,1110]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="26" pageNumber="27">WAM</collectionCode>
|
||
R125521 from the ‘northern Pilbara’) the entire head and body are more intensely marked with variable sized spots; these are randomly distributed on the head and neck but are aligned into more or less regular transverse rows posterior to the level of attachment of the forelimbs. All specimens of
|
||
<emphasis id="B978EAF4FF8EFF92FC4DFB5DA224FB57" box="[951,1045,1204,1229]" italics="true" pageId="26" pageNumber="27">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8EFF92FBDFFB5DA2B9FB54" bold="true" box="[1061,1160,1204,1230]" pageId="26" pageNumber="27">sp. nov.</emphasis>
|
||
have ventral spotting on the throat, neck and body. In comparison with the
|
||
<typeStatus id="54B78844FF8EFF92FC15FB35A269FB6C" box="[1007,1112,1244,1270]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
, most individuals show more intense and evenly distributed spotting on the body. Some individuals also show a strong differentiation between abundant fine spotting on the throat and neck, with each spot consisting of a single pigmented primary scale, and less abundant but larger blotches on the body.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8EFF92FEC2FA4DA49EF9AC" blockId="26.[264,1324,284,1870]" pageId="26" pageNumber="27">
|
||
Dorsal ground colour in life is pale greybrown, typically with a reddish tinge on the upper surface, from the crown of the head to between the hind limbs (
|
||
<figureCitation id="13372A63FF8EFF92FBBBFA25A2B7FA7C" box="[1089,1158,1484,1510]" captionStart="FIGURE 1" captionStartId="2.[264,368,1268,1292]" captionTargetBox="[267,1319,526,1244]" captionTargetId="figure@2.[267,1319,526,1244]" captionTargetPageId="2" captionText="FIGURE 1. An adult Varanus bushi sp. nov. from near Roebourne, photographed in life by Robert BrowneCooper." httpUri="https://zenodo.org/record/173908/files/figure.png" pageId="26" pageNumber="27">Fig. 1</figureCitation>
|
||
). The ground colour of the tail is always paler than the body and generally has a cream wash on the distal one third of the upper surface.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8EFF93FEC2F9ADA2DDFE1C" blockId="26.[264,1324,284,1870]" lastBlockId="27.[264,1323,284,1870]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
<emphasis id="B978EAF4FF8EFF92FEC2F9ADA4ABF9C4" bold="true" box="[312,666,1604,1630]" pageId="26" pageNumber="27">Morphological comparisons</emphasis>
|
||
:
|
||
<taxonomicName id="4C0C4D65FF8EFF92FD57F9ADA568F9C7" box="[685,857,1604,1629]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8EFF92FD57F9ADA568F9C7" box="[685,857,1604,1629]" italics="true" pageId="26" pageNumber="27">Varanus bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8EFF92FC9FF9ADA5F6F9C4" box="[869,967,1604,1630]" pageId="26" pageNumber="27" rank="species">
|
||
<emphasis id="B978EAF4FF8EFF92FC9FF9ADA5F6F9C4" bold="true" box="[869,967,1604,1630]" pageId="26" pageNumber="27">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is most similar overall to
|
||
<taxonomicName id="4C0C4D65FF8EFF92FAEFF9ADA762F91F" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gilleni">
|
||
<emphasis id="B978EAF4FF8EFF92FAEFF9ADA762F91F" italics="true" pageId="26" pageNumber="27">V. gilleni</emphasis>
|
||
</taxonomicName>
|
||
, a phenetic resemblance that is consistent with the phylogenetic conclusions of the molecular analyses. These species share a number of morphological attributes including a relatively elongate body with proportionally shorter limbs, high midbody and ventral scale counts, high subdigital lamellar counts on the pes, a relatively nonspinose basal portion of the tail, and a tendency to include transverse bands in the dorsal patterning.
|
||
<collectingCountry id="F31B7676FF8EFF92FAF0F8E5A31AF8BC" box="[1290,1323,1804,1830]" name="American Samoa" pageId="26" pageNumber="27">As</collectingCountry>
|
||
indicated in the diagnosis,
|
||
<taxonomicName id="4C0C4D65FF8EFF92FDB9F8DDA491F8D7" box="[579,672,1844,1869]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8EFF92FDB9F8DDA491F8D7" box="[579,672,1844,1869]" italics="true" pageId="26" pageNumber="27">V. bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8EFF92FD5DF8DDA537F8D4" box="[679,774,1844,1870]" pageId="26" pageNumber="27" rank="species">
|
||
<emphasis id="B978EAF4FF8EFF92FD5DF8DDA537F8D4" bold="true" box="[679,774,1844,1870]" pageId="26" pageNumber="27">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C0C4D65FF8EFF92FC5BF8DDA238F8D7" box="[929,1033,1844,1869]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gilleni">
|
||
<emphasis id="B978EAF4FF8EFF92FC5BF8DDA238F8D7" box="[929,1033,1844,1869]" italics="true" pageId="26" pageNumber="27">V. gilleni</emphasis>
|
||
</taxonomicName>
|
||
in its lower average and maximum body size, its more elongate dorsal scales, its more densely spotted venter, its less conspicuous linear patterning on the head and neck, its more numerous presacral vertebrae, its simpler heimbacular morphology, and some minor meristic differences.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316656DFF8FFF94FEC2FE7DA7FCFE64" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8BB336E6FF8FFF93FEC2FE7DA2B6FC2C" blockId="27.[264,1323,284,1870]" pageId="27" pageNumber="28">
|
||
<emphasis id="B978EAF4FF8FFF93FEC2FE7DA7EFFE37" box="[312,478,404,429]" italics="true" pageId="27" pageNumber="28">
|
||
Var a nu s
|
||
<taxonomicName id="4C0C4D65FF8FFF93FE65FE7DA7EFFE37" box="[415,478,404,429]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">bushi</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A24B578FFF8FFF93FE1DFE7AA475FE37" box="[487,580,403,429]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="B978EAF4FF8FFF93FE1DFE7AA475FE37" bold="true" box="[487,580,403,429]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is more similar to
|
||
<taxonomicName id="4C0C4D65FF8FFF93FCDEFE7DA5D7FE37" box="[804,998,404,429]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FCDEFE7DA5D7FE37" box="[804,998,404,429]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
in body patterning and this probably explains the former confusion between these species. Both species share a spotted rather than banded pattern on the dorsum, and both are typically spotted on the throat and to some degree on the venter. However,
|
||
<taxonomicName id="4C0C4D65FF8FFF93FC91FDE5A222FDBF" box="[875,1043,524,549]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8FFF93FC91FDE5A222FDBF" box="[875,1043,524,549]" italics="true" pageId="27" pageNumber="28">Varanus bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8FFF93FBE6FDE2A24DFDBF" box="[1052,1148,523,549]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="B978EAF4FF8FFF93FBE6FDE2A24DFDBF" bold="true" box="[1052,1148,523,549]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is usually less heavily pigmented than typical
|
||
<taxonomicName id="4C0C4D65FF8FFF93FD85FDDDA570FDD7" box="[639,833,564,589]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FD85FDDDA570FDD7" box="[639,833,564,589]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
, with smaller and less intense spots. The lower back and basal portion of the tail are distinctly banded in
|
||
<emphasis id="B978EAF4FF8FFF93FC0DFDB5A261FDEF" box="[1015,1104,604,629]" italics="true" pageId="27" pageNumber="28">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8FFF93FBA6FDB2A288FDEF" bold="true" box="[1116,1209,603,629]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
but more irregularly patterned in
|
||
<taxonomicName id="4C0C4D65FF8FFF93FDCCFD6DA4CEFD07" box="[566,767,644,669]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FDCCFD6DA4CEFD07" box="[566,767,644,669]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
. Body proportions also distinguish the two species,
|
||
<emphasis id="B978EAF4FF8FFF93FE91FD45A7F3FD5F" box="[363,450,684,709]" italics="true" pageId="27" pageNumber="28">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8FFF93FE37FD45A41BFD5C" bold="true" box="[461,554,684,710]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
having a more elongate body form with more numerous presacral vertebrae, higher ventral scale counts, more elongate dorsal primary scales, and relatively shorter fore and hindlimbs. Although tail length is quite variable in both taxa, the majority of
|
||
<taxonomicName id="4C0C4D65FF8FFF93FE62FCCDA46AFCA7" box="[408,603,804,829]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FE62FCCDA46AFCA7" box="[408,603,804,829]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
have proportionally shorter tails than
|
||
<emphasis id="B978EAF4FF8FFF93FBD0FCCDA2B4FCA7" box="[1066,1157,804,829]" italics="true" pageId="27" pageNumber="28">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8FFF93FB69FCCDA2C2FCA4" bold="true" box="[1171,1267,804,830]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
The head is also shorter and stockier in
|
||
<taxonomicName id="4C0C4D65FF8FFF93FD55FCA5A540FCFF" box="[687,881,844,869]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FD55FCA5A540FCFF" box="[687,881,844,869]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
, produced mainly by a shortening of the rostrum. Midbody scale counts are typically higher in
|
||
<taxonomicName id="4C0C4D65FF8FFF93FC11FC9DA27FFC17" box="[1003,1102,884,909]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8FFF93FC11FC9DA27FFC17" box="[1003,1102,884,909]" italics="true" pageId="27" pageNumber="28">V. bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8FFF93FBA6FC9DA2F1FC14" box="[1116,1216,884,910]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="B978EAF4FF8FFF93FBA6FC9DA2F1FC14" bold="true" box="[1116,1216,884,910]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and the hemipenes differ in several important details from those of
|
||
<taxonomicName id="4C0C4D65FF8FFF93FC45FC75A2B1FC2F" box="[959,1152,924,949]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FC45FC75A2B1FC2F" box="[959,1152,924,949]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8FFF93FEC2FC2DA5E5FB54" blockId="27.[264,1323,284,1870]" pageId="27" pageNumber="28">
|
||
The evolutionary polarity of most of the morphological characteristics that distinguish each of the three members of the
|
||
<taxonomicName id="4C0C4D65FF8FFF93FD62FC05A56BFB9F" box="[664,858,1004,1029]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FD62FC05A56BFB9F" box="[664,858,1004,1029]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
species group is uncertain.
|
||
<bibRefCitation id="EF9D4B17FF8FFF93FB5DFC05A767FBB4" author="Sprackland" pageId="27" pageNumber="33" refString="Sprackland, R. G. (1991) The origin and zoogeography of monitor lizards of the subgenus Odatria Gray (Sauria: Varanidae): a re-evaluation. Mertensiella, 2, 240 - 252." type="journal article" year="1991">Sprackland (1991)</bibRefCitation>
|
||
suggests that more heavily rugose tails and smaller body size are probably apomorphic within
|
||
<taxonomicName id="4C0C4D65FF8FFF93FE15FBD5A47DFBCF" box="[495,588,1084,1109]" class="Squamata" family="Varanidae" genus="Odatria" higherTaxonomySource="CoL" kingdom="Animalia" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
|
||
<emphasis id="B978EAF4FF8FFF93FE15FBD5A47DFBCF" box="[495,588,1084,1109]" italics="true" pageId="27" pageNumber="28">Odatria</emphasis>
|
||
</taxonomicName>
|
||
. On both counts,
|
||
<taxonomicName id="4C0C4D65FF8FFF93FCE5FBD5A5D0FBCF" box="[799,993,1084,1109]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FCE5FBD5A5D0FBCF" box="[799,993,1084,1109]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
would rate as more derived than either of the other taxa.
|
||
<taxonomicName id="4C0C4D65FF8FFF93FD84FB8DA5A1FBE7" box="[638,912,1124,1149]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FD84FB8DA5A1FBE7" box="[638,912,1124,1149]" italics="true" pageId="27" pageNumber="28">Varanus caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
appears to be more primitive in hemipeneal morphology than either
|
||
<taxonomicName id="4C0C4D65FF8FFF93FD4AFB65A53CFB3F" box="[688,781,1164,1189]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8FFF93FD4AFB65A53CFB3F" box="[688,781,1164,1189]" italics="true" pageId="27" pageNumber="28">V. bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8FFF93FCE9FB65A541FB3C" box="[787,880,1164,1190]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="B978EAF4FF8FFF93FCE9FB65A541FB3C" bold="true" box="[787,880,1164,1190]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
or
|
||
<emphasis id="B978EAF4FF8FFF93FC6DFB65A5CBFB3F" box="[919,1018,1164,1189]" italics="true" pageId="27" pageNumber="28">V. g i l l e n i</emphasis>
|
||
; of the latter two species,
|
||
<taxonomicName id="4C0C4D65FF8FFF93FEF2FB5DA741FB57" box="[264,368,1204,1229]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gilleni">
|
||
<emphasis id="B978EAF4FF8FFF93FEF2FB5DA741FB57" box="[264,368,1204,1229]" italics="true" pageId="27" pageNumber="28">V. gilleni</emphasis>
|
||
</taxonomicName>
|
||
possesses a more derived hemibacular morphology.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8FFF93FEC2FB35A405FA7C" blockId="27.[264,1323,284,1870]" pageId="27" pageNumber="28">
|
||
<emphasis id="B978EAF4FF8FFF93FEC2FB35A40CFB6C" bold="true" box="[312,573,1244,1270]" pageId="27" pageNumber="28">Taxonomic remarks</emphasis>
|
||
: Although the
|
||
<typeStatus id="54B78844FF8FFF93FCE9FB35A577FB6C" box="[787,838,1244,1270]" pageId="27" pageNumber="28">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C0C4D65FF8FFF93FBE9FB35A7FCFA84" authority="Boulenger, 1885" authorityName="Boulenger" authorityYear="1885" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FBE9FB35A31BFB6F" box="[1043,1322,1244,1269]" italics="true" pageId="27" pageNumber="28">Varanus caudolineatus</emphasis>
|
||
<bibRefCitation id="EF9D4B17FF8FFF93FEF2FAEDA7FCFA84" author="Boulenger" box="[264,461,1284,1310]" pageId="27" pageNumber="32" refString="Boulenger, G. A. (1885) Catalogue of the lizards in the British Museum (Natural History). Volume 2. British Museum, London, xiii + 497 pp." type="book" year="1885">Boulenger, 1885</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0C4D65FF8FFF93FDF1FAEDA5F3FA84" authority="Lucas & Frost, 1895" authorityName="Lucas & Frost" authorityYear="1895" box="[523,962,1284,1310]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gilleni">
|
||
<emphasis id="B978EAF4FF8FFF93FDF1FAEDA4F1FA87" box="[523,704,1284,1309]" italics="true" pageId="27" pageNumber="28">Varanus gilleni</emphasis>
|
||
<bibRefCitation id="EF9D4B17FF8FFF93FD33FAEDA5F3FA84" author="Lucas" box="[713,962,1284,1310]" pageId="27" pageNumber="33" refString="Lucas, A. H. S. & Frost, C. (1895) Preliminary notice of certain new species of lizards from central Australia. Proceedings of the Royal Society of Victoria, 7, 264 - 269." type="journal article" year="1895">Lucas & Frost, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
were not examined as part of this study, the
|
||
<typeStatus id="54B78844FF8FFF93FE55FAC5A7D3FADC" box="[431,482,1324,1350]" pageId="27" pageNumber="28">type</typeStatus>
|
||
localities of each of these taxa [Champion Bay, W.A. (= Geraldton) and Charlotte Waters, N.T.; see
|
||
<figureCitation id="13372A63FF8FFF93FDB7FABDA4A2FAF4" box="[589,659,1364,1390]" captionStart="FIGURE 9" captionStartId="24.[264,368,1779,1803]" captionTargetBox="[465,1125,1158,1761]" captionTargetId="figure@24.[465,1125,1158,1766]" captionTargetPageId="24" captionText="FIGURE 9. Distribution of the three Varanus species based on examined materials and type localities. The approximate wider distribution of V. g i l l e n i is indicated by the dashed line. Symbols: Stars — V. caudolineatus; Triangles — V. gilleni; Circles — V. b u s h i sp. nov.; Circles with cross — s V. caudolineatus and V. b u s h i sp. nov. in local sympatry; Circles with dot – type localities." httpUri="https://zenodo.org/record/173916/files/figure.png" pageId="27" pageNumber="28">Fig. 9</figureCitation>
|
||
] is remote from the known geographic range of
|
||
<taxonomicName id="4C0C4D65FF8FFF93FB34FABDA31AFAF7" box="[1230,1323,1364,1389]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8FFF93FB34FABDA31AFAF7" box="[1230,1323,1364,1389]" italics="true" pageId="27" pageNumber="28">V. bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8FFF93FEF2FA95A759FA0C" box="[264,360,1404,1430]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="B978EAF4FF8FFF93FEF2FA95A759FA0C" bold="true" box="[264,360,1404,1430]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and well within the ranges of these other taxa as currently understood. We are therefore confident that the various names are correctly associated with the biological entities as defined herein.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8FFF93FEC2FA1DA7ACF91C" blockId="27.[264,1323,284,1870]" pageId="27" pageNumber="28">
|
||
<emphasis id="B978EAF4FF8FFF93FEC2FA1DA432F994" bold="true" box="[312,515,1524,1550]" pageId="27" pageNumber="28">Ecological notes</emphasis>
|
||
: Relatively little information is available on the ecology of Pilbara varanids and the new taxon is no exception. Most recent specimens of
|
||
<taxonomicName id="4C0C4D65FF8FFF93FB98F9F5A2F0F9AF" box="[1122,1217,1564,1589]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="bushi" status="sp. nov.">
|
||
<emphasis id="B978EAF4FF8FFF93FB98F9F5A2F0F9AF" box="[1122,1217,1564,1589]" italics="true" pageId="27" pageNumber="28">V. bushi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24B578FFF8FFF93FB30F9F5A31BF9AC" box="[1226,1322,1564,1590]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="B978EAF4FF8FFF93FB30F9F5A31BF9AC" bold="true" box="[1226,1322,1564,1590]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have been taken from fallen or standing hollow trees in mulga or eucalypt woodland associations.
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF8FFF94FEC2F97DA52BFEC4" blockId="27.[264,1323,284,1870]" lastBlockId="28.[264,1323,284,510]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
The sex ratio among the total sample of
|
||
<emphasis id="B978EAF4FF8FFF93FCDAF97DA54BF937" box="[800,890,1684,1709]" italics="true" pageId="27" pageNumber="28">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8FFF93FC7DF97DA5D6F934" bold="true" box="[903,999,1684,1710]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
is
|
||
<specimenCount id="9D0AFD6FFF8FFF93FBF7F97DA249F934" box="[1037,1144,1684,1710]" pageId="27" pageNumber="28" type="male">34 males</specimenCount>
|
||
to
|
||
<specimenCount id="9D0AFD6FFF8FFF93FB5BF97DA316F934" box="[1185,1319,1684,1710]" pageId="27" pageNumber="28" type="female">13 females</specimenCount>
|
||
. This situation is typical for opportunistically collected samples of varanids (
|
||
<bibRefCitation id="EF9D4B17FF8FFF93FB42F955A798F964" author="King" pageId="27" pageNumber="32" refString="King, D. J. & Rhodes, L. (1982) Sex ratio and breeding season of Varanus acanthurus. Copeia, 1984: 784 - 87." type="journal article" year="1982">King and Rhodes 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9D4B17FF8FFF93FE43F90DA472F964" author="Greer" box="[441,579,1764,1790]" pageId="27" pageNumber="32" refString="Greer, A. E., (1989) The Biology and Evolution of Australian Lizards. Surrey Beatty, Sydney, New South Wales, 264 pp." type="book" year="1989">Greer 1989</bibRefCitation>
|
||
) and probably reflects different activity patterns between the sexes. The lack of immature specimens of
|
||
<emphasis id="B978EAF4FF8FFF93FD07F8E5A564F8BF" box="[765,853,1804,1829]" italics="true" pageId="27" pageNumber="28">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF8FFF93FC9BF8E5A58FF8BC" bold="true" box="[865,958,1804,1830]" pageId="27" pageNumber="28">sp. nov.</emphasis>
|
||
and the small numbers of very young individuals in the large samples of
|
||
<taxonomicName id="4C0C4D65FF8FFF93FD0EF8DDA587F8D7" box="[756,950,1844,1869]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caudolineatus">
|
||
<emphasis id="B978EAF4FF8FFF93FD0EF8DDA587F8D7" box="[756,950,1844,1869]" italics="true" pageId="27" pageNumber="28">V. caudolineatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B978EAF4FF8FFF93FC15F8DDA262F8D7" box="[1007,1107,1844,1869]" italics="true" pageId="27" pageNumber="28">V. g i l l e n i</emphasis>
|
||
is consistent with previous comment that hatchling varanids are especially cryptic and poorly represented in wild caught samples (
|
||
<bibRefCitation id="EF9D4B17FF88FF94FDFCFEADA53CFEC4" author="Horn" box="[518,781,324,350]" pageId="28" pageNumber="32" refString="Horn, H. - G. & Visser, G. J. (1991) Basic data on the biology of monitors. Mertensiella, 2, 176 - 187." type="journal article" year="1991">Horn and Visser 1991</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB336E6FF88FF94FEC2FE85A7FCFE64" blockId="28.[264,1323,284,510]" pageId="28" pageNumber="29">
|
||
Identifiable stomach contents were observed in two of four
|
||
<emphasis id="B978EAF4FF88FF94FC0BFE85A278FE1F" box="[1009,1097,364,389]" italics="true" pageId="28" pageNumber="29">V. b u s h i</emphasis>
|
||
<emphasis id="B978EAF4FF88FF94FBAEFE82A280FE1F" bold="true" box="[1108,1201,363,389]" pageId="28" pageNumber="29">sp. nov.</emphasis>
|
||
examined by D.J. King. WAM R73142 (male, SVL
|
||
<quantity id="4CF49B03FF88FF94FCF0FE7DA552FE34" box="[778,867,404,430]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="28" pageNumber="29" unit="mm" value="96.0">96 mm</quantity>
|
||
) contained remains of a spider and a skink tail; WAM R54230 (male, SVL
|
||
<quantity id="4CF49B03FF88FF94FD11FE55A565FE4C" box="[747,852,444,470]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" pageId="28" pageNumber="29" unit="mm" value="135.0">135 mm</quantity>
|
||
) contained remains of a mole cricket (
|
||
<taxonomicName id="4C0C4D65FF88FF94FEEAFE0DA7F0FE64" box="[272,449,484,510]" class="Insecta" family="Gryllotalpidae" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Gryllotalpidae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |