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<document id="E6186A9BD30FCD6DFA256140B484D2F5" ID-DOI="10.11646/zootaxa.3702.4.5" ID-GBIF-Dataset="d5a0e4f8-9331-4408-91a5-22dd70649743" ID-ISSN="1175-5326" ID-Zenodo-Dep="221020" ID-ZooBank="DAE56C35-EEAB-42B4-A36B-9674550945C6" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460671780265" checkinUser="plazi" docAuthor="Wheeler, Terry A. &amp; Solecki, Anna M." docDate="2013" docId="03A587EE5B01FFA296A1069493CAFCBB" docLanguage="en" docName="zt03702p385.pdf" docOrigin="Zootaxa 3702 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calamoncosis agricola Wheeler &amp; Solecki, sp. n." docType="treatment" docVersion="8" lastPageNumber="381" masterDocId="FF9CFF965B00FFA0963606039011FFE7" masterDocTitle="New species of Calamoncosis Enderlein (Diptera: Chloropidae) from South Africa" masterLastPageNumber="385" masterPageNumber="379" pageNumber="380" updateTime="1698348403470" updateUser="plazi">
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<mods:title id="846F90AAB209817DB80CBD4E33AB2EA9">New species of Calamoncosis Enderlein (Diptera: Chloropidae) from South Africa</mods:title>
</mods:titleInfo>
<mods:name id="EAE53DEF736466C6A59AD2DE4EBF17DB" type="personal">
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<mods:namePart id="92ECDDC6DE97F0F29D55514BFBFCC20D">Wheeler, Terry A.</mods:namePart>
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<mods:namePart id="B70D41E07CE0BC8797F1036700C16C41">Solecki, Anna M.</mods:namePart>
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<mods:title id="2D909F246F7DBB12F0225B10B4453A2A">Zootaxa</mods:title>
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<mods:date id="CB34062EE8E7DD3CD3C9F59C2AB0ECCC">2013</mods:date>
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<mods:number id="69A63900C018B5CD56448F15A0689803">3702</mods:number>
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<mods:identifier id="FAD37123DD45FCE0DD06CAE4D5A9B43B" type="DOI">10.11646/zootaxa.3702.4.5</mods:identifier>
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<treatment id="03A587EE5B01FFA296A1069493CAFCBB" ID-DOI="http://doi.org/10.5281/zenodo.6153216" ID-GBIF-Taxon="119576507" ID-Zenodo-Dep="6153216" LSID="urn:lsid:plazi:treatment:03A587EE5B01FFA296A1069493CAFCBB" httpUri="http://treatment.plazi.org/id/03A587EE5B01FFA296A1069493CAFCBB" lastPageId="2" lastPageNumber="381" pageId="1" pageNumber="380">
<subSubSection id="C31665735B01FFA196A106949100FF34" pageId="1" pageNumber="380" type="nomenclature">
<paragraph id="8BB336F85B01FFA196A1069492F7FF55" blockId="1.[151,742,151,211]" box="[151,742,151,178]" pageId="1" pageNumber="380">
<heading id="D0FB81945B01FFA196A1069492F7FF55" bold="true" box="[151,742,151,178]" fontSize="11" level="1" pageId="1" pageNumber="380" reason="1">
<emphasis id="B978EAEA5B01FFA196A1069492F7FF55" bold="true" box="[151,742,151,178]" pageId="1" pageNumber="380">
<taxonomicName id="4C0C4D7B5B01FFA196A10694928DFF55" authority="Wheeler &amp; Solecki" authorityName="Wheeler &amp; Solecki" box="[151,668,151,178]" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="380" phylum="Arthropoda" rank="species" species="agricola" status="sp. nov.">
<emphasis id="B978EAEA5B01FFA196A1069491BBFF56" bold="true" box="[151,426,151,177]" italics="true" pageId="1" pageNumber="380">Calamoncosis agricola</emphasis>
Wheeler &amp; Solecki
</taxonomicName>
<taxonomicNameLabel id="A24B57915B01FFA19492069B92F7FF55" box="[676,742,152,178]" pageId="1" pageNumber="380" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336F85B01FFA196A106B99100FF34" blockId="1.[151,742,151,211]" box="[151,273,186,211]" pageId="1" pageNumber="380">
(
<figureCitation id="13372A7D5B01FFA196A906B99119FF34" box="[159,264,186,211]" captionStart="FIGURES 1 3" captionStartId="1.[151,264,1561,1583]" captionTargetBox="[155,1429,781,1531]" captionTargetId="figure@1.[151,1436,770,1536]" captionTargetPageId="1" captionText="FIGURES 1 3. Calamoncosis agricola. 1. male genitalia, lateral. 2. male genitalia, posterior. 3. male genitalia, ventral. Abbreviations: cer—cerci, epd—epandrium, sur surstylus." httpUri="https://zenodo.org/record/221021/files/figure.png" pageId="1" pageNumber="380">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31665735B01FFA196A107019391FED8" pageId="1" pageNumber="380" type="diagnosis">
<paragraph id="8BB336F85B01FFA196A107019391FED8" blockId="1.[151,1436,258,715]" pageId="1" pageNumber="380">
<emphasis id="B978EAEA5B01FFA196A107019102FEFC" bold="true" box="[151,275,258,283]" pageId="1" pageNumber="380">Diagnosis:</emphasis>
This species may be distinguished from other described Afrotropical
<taxonomicName id="4C0C4D7B5B01FFA19269070094EFFEFC" box="[1119,1278,259,283]" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAEA5B01FFA19269070094EFFEFC" box="[1119,1278,259,283]" italics="true" pageId="1" pageNumber="380">Calamoncosis</emphasis>
</taxonomicName>
by the wing venation; the absence of crossvein dm-cu is atypical for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C31665735B01FFA296F107499198FE4A" lastPageId="2" lastPageNumber="381" pageId="1" pageNumber="380" type="description">
<paragraph id="8BB336F85B01FFA196F10749926BFD2C" blockId="1.[151,1436,258,715]" pageId="1" pageNumber="380">
<emphasis id="B978EAEA5B01FFA196F10749914AFE84" bold="true" box="[199,347,330,355]" pageId="1" pageNumber="380">Description:</emphasis>
Total length:
<quantity id="4CF49B1D5B01FFA194370748924BFE85" box="[513,602,331,355]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="380" unit="mm" value="1.4">1.4 mm</quantity>
(male),
<quantity id="4CF49B1D5B01FFA1948907499358FE85" box="[703,841,330,355]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="1" pageNumber="380" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.12.5 mm</quantity>
(female); head + thorax length 1.0 (male), 1.31.4 (female). Overall colour shining black; head higher than long, frons densely microtomentose, appearing velvety, interfrontal and fronto-orbital setulae less dense than setulae on scutum, fine, black; frontal triangle 0.40.5 times length of frons, shining but striated, with setulae in irregular punctate row near lateral margin, slightly longer posteriorly, stronger than interfrontals; inner vertical seta barely differentiated from interfrontals, outer vertical seta, ocellars and postocellars stronger, black, ocellars reclinate, postocellars slightly convergent; eye with sparse ommatrichia, each hair approximately as long as diameter of 2 ommatidia; gena black, vibrissal angle obtuse, not extending beyond anterior margin of eye, genal height 0.20.3 times eye height; face black to dark brown, carina narrow, complete, antennae in deep foveae; antenna with pedicel dark yellow-brown, first flagellomere smaller than genal height, with angulate apex, dark yellow-brown, arista short, brown, pubescent with thick base; palpus small, dark-brown; proboscis brown, short.
</paragraph>
<caption id="DF7366705B01FFA196A1001A92EBF9AA" httpUri="https://zenodo.org/record/221021/files/figure.png" pageId="1" pageNumber="380" targetBox="[155,1429,781,1531]" targetPageId="1">
<paragraph id="8BB336F85B01FFA196A1001A92EBF9AA" blockId="1.[151,1436,1561,1613]" pageId="1" pageNumber="380">
<emphasis id="B978EAEA5B01FFA196A1001A912EF9C8" bold="true" box="[151,319,1561,1583]" pageId="1" pageNumber="380">FIGURES 13.</emphasis>
<taxonomicName id="4C0C4D7B5B01FFA1977C001A9228F9C8" box="[330,569,1561,1583]" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="380" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="B978EAEA5B01FFA1977C001A9228F9C8" box="[330,569,1561,1583]" italics="true" pageId="1" pageNumber="380">Calamoncosis agricola</emphasis>
</taxonomicName>
. 1. male genitalia, lateral. 2. male genitalia, posterior. 3. male genitalia, ventral. Abbreviations: cer—cerci, epd—epandrium, sur surstylus.
</paragraph>
</caption>
<paragraph id="8BB336F85B01FFA196F100769170F811" blockId="1.[151,1436,1652,2038]" pageId="1" pageNumber="380">
Scutum shining but densely punctate, evenly covered with long, pale setulae; thoracic setae: 1 anterior and 2 posterior notopleural, 1 postalar, 1 postsutural intra-alar, 1 dorsocentral stronger and darker than surrounding setulae; scutellum round, slightly wider than long, punctate and rugose dorsally, apical scutellar setae dark, strong, slightly shorter than scutellum, arising from small but distinct tubercles, one pair of shorter and weaker subapical setae, arising from small but distinct tubercles, other scutellar setae as strong as scutal setulae; thoracic pleurites mostly bare and shining, katepisternum with sparse, fine, pale setulae ventrally. Legs: femora and tibiae dark brown with yellow apices (fore tibia more extensively yellow in females), tarsi yellow; male mid-femoral organ a dense patch of 2530 small tubercles with minute setulae, hind-tibial organ indistinct, apparent as an oval, velvety patch. Wing faintly whitish in males, with darker veins anteriorly, faintly brownish in females with brown veins, a pronounced ventral bulge in cu
<emphasis id="B978EAEA5B01FFA197C501B99210F836" box="[499,513,1978,2001]" italics="true" pageId="1" pageNumber="380">p</emphasis>
area, crossvein dm-cu absent; ratio of costal sectors C1: C2: C3: C40.8: 1.0: 0.6: 0.4; halter yellow.
</paragraph>
<paragraph id="8BB336F85B02FFA296F1069492ACFF57" blockId="2.[151,1436,151,861]" box="[199,701,151,176]" pageId="2" pageNumber="381">Abdomen slightly paler than thorax, shining.</paragraph>
<paragraph id="8BB336F85B02FFA296F106BF9588FE82" blockId="2.[151,1436,151,861]" pageId="2" pageNumber="381">
Male genitalia (
<figureCitation id="13372A7D5B02FFA2974C06BF91F6FF33" box="[378,487,188,213]" captionStart="FIGURES 1 3" captionStartId="1.[151,264,1561,1583]" captionTargetBox="[155,1429,781,1531]" captionTargetId="figure@1.[151,1436,770,1536]" captionTargetPageId="1" captionText="FIGURES 1 3. Calamoncosis agricola. 1. male genitalia, lateral. 2. male genitalia, posterior. 3. male genitalia, ventral. Abbreviations: cer—cerci, epd—epandrium, sur surstylus." httpUri="https://zenodo.org/record/221021/files/figure.png" pageId="2" pageNumber="381">Figs. 13</figureCitation>
): epandrium higher than long in lateral view, broad in posterior view, with setae on posterior half; surstylus quadrate, broader at ventral apex with jagged margin, with sparse, weak setae; hypandrium robust anteriorly; pregonites, postgonites and phallic guide all well-sclerotized; postgonites with weak setulae posteriorly; basiphallus well-sclerotized, distiphallus longer than basiphallus, membranous; cerci fused medially, not projecting ventrally as in many
<taxonomicName id="4C0C4D7B5B02FFA29413074F92A5FE82" box="[549,692,332,357]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="381" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
, indistinctly sclerotized; subepandrial sclerite not well-developed.
</paragraph>
<paragraph id="8BB336F85B02FFA296F1076C9198FE4A" blockId="2.[151,1436,151,861]" pageId="2" pageNumber="381">Female postabdomen with segments 810 laterally compressed and sclerotized, cerci laterally compressed, sclerotized, knifelike.</paragraph>
</subSubSection>
<subSubSection id="C31665735B02FFA296F107B49480FE13" pageId="2" pageNumber="381" type="materials_examined">
<paragraph id="8BB336F85B02FFA296F107B49480FE13" blockId="2.[151,1436,151,861]" pageId="2" pageNumber="381">
<emphasis id="B978EAEA5B02FFA296F107B4916CFE37" bold="true" box="[199,381,439,464]" pageId="2" pageNumber="381">
<typeStatus id="54B7885A5B02FFA296F107B49113FE37" box="[199,258,439,464]" pageId="2" pageNumber="381">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54B7885A5B02FFA297BC07B491E3FE37" box="[394,498,439,464]" pageId="2" pageNumber="381" type="holotype">Holotype</typeStatus>
♂:
<collectingCountry id="F31B76685B02FFA2941F07BB92EFFE37" box="[553,766,440,464]" name="South Africa" pageId="2" pageNumber="381">SOUTH AFRICA</collectingCountry>
: East Transvaal:
<collectingCountry id="F31B76685B02FFA295F807BB942BFE37" box="[974,1082,440,464]" name="Guernsey" pageId="2" pageNumber="381">Guernsey</collectingCountry>
Farm,
<quantity id="4CF49B1D5B02FFA292A007B494C7FE37" box="[1174,1238,439,464]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="381" unit="km" value="15.0">15km</quantity>
E Klaserie,
<date id="FFB210385B02FFA2934407BB9108FE12" pageId="2" pageNumber="381" value="1985-12-19" valueMax="1985-12-31" valueMin="1985-12-19">19 31.xii.1985</date>
, P.M. Sanborne, Malaise (LEM).
<typeStatus id="54B7885A5B02FFA294A407DE9312FE12" box="[658,771,477,501]" pageId="2" pageNumber="381" type="paratype">Paratypes</typeStatus>
: same data as
<typeStatus id="54B7885A5B02FFA2959407DF9415FE12" box="[930,1028,476,501]" pageId="2" pageNumber="381" type="holotype">holotype</typeStatus>
(3♀, LEM).
</paragraph>
</subSubSection>
<subSubSection id="C31665735B02FFA296F107FC90E0FDDA" pageId="2" pageNumber="381" type="etymology">
<paragraph id="8BB336F85B02FFA296F107FC90E0FDDA" blockId="2.[151,1436,151,861]" pageId="2" pageNumber="381">
<emphasis id="B978EAEA5B02FFA296F107FC9140FDFF" bold="true" box="[199,337,511,536]" pageId="2" pageNumber="381">Etymology:</emphasis>
The species name is a masculine noun from the Latin
<taxonomicName id="4C0C4D7B5B02FFA295EC04039428FDFF" box="[986,1081,512,536]" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="381" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="B978EAEA5B02FFA295EC04039428FDFF" box="[986,1081,512,536]" italics="true" pageId="2" pageNumber="381">agricola</emphasis>
</taxonomicName>
(farmer), referring to the
<typeStatus id="54B7885A5B02FFA2935D0403958AFDFF" box="[1387,1435,512,536]" pageId="2" pageNumber="381">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C31665735B02FFA296F1044493CAFCBB" pageId="2" pageNumber="381" type="discussion">
<paragraph id="8BB336F85B02FFA296F1044491B0FD62" blockId="2.[151,1436,151,861]" pageId="2" pageNumber="381">
<emphasis id="B978EAEA5B02FFA296F10444912CFD87" bold="true" box="[199,317,583,608]" pageId="2" pageNumber="381">Remarks:</emphasis>
Based on the structure of the female ovipositor, this species would be assigned to the subgenus
<emphasis id="B978EAEA5B02FFA296A1046F913DFD63" box="[151,300,620,644]" italics="true" pageId="2" pageNumber="381">Rhaphiopyga</emphasis>
Nartshuk.
</paragraph>
<paragraph id="8BB336F85B02FFA296F1048C93CAFCBB" blockId="2.[151,1436,151,861]" pageId="2" pageNumber="381">
Three additional specimens with the same collecting data (
<specimenCount id="9D0AFD715B02FFA295650493936EFD40" box="[851,895,656,680]" pageId="2" pageNumber="381" type="generic">2 ♂</specimenCount>
, 1 ♀, LEM) are similar to
<taxonomicName id="4C0C4D7B5B02FFA2929004939534FD4F" box="[1190,1317,656,680]" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="381" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="B978EAEA5B02FFA2929004939534FD4F" box="[1190,1317,656,680]" italics="true" pageId="2" pageNumber="381">C. agricola</emphasis>
</taxonomicName>
, including male genitalia, but vary in multiple characters including the presence of crossvein dm-cu, the colour of the halteres, and/or the shape of the antennae. Based on these differences, we have not included them as
<typeStatus id="54B7885A5B02FFA292DE04DB9542FD17" box="[1256,1363,728,752]" pageId="2" pageNumber="381" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C0C4D7B5B02FFA293B404DB90E7FCF3" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="381" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="B978EAEA5B02FFA293B404DB90E7FCF3" italics="true" pageId="2" pageNumber="381">C. agricola</emphasis>
</taxonomicName>
; however, there is insufficient material to describe any of them as new species. In the absence of additional specimens from additional collecting events, we cannot determine whether
<taxonomicName id="4C0C4D7B5B02FFA292BA05239500FCDF" box="[1164,1297,800,824]" class="Insecta" family="Chloropidae" genus="Calamoncosis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="381" phylum="Arthropoda" rank="species" species="agricola">
<emphasis id="B978EAEA5B02FFA292BA05239500FCDF" box="[1164,1297,800,824]" italics="true" pageId="2" pageNumber="381">C. agricola</emphasis>
</taxonomicName>
is a highly variable species, or whether there are additional sympatric species present.
</paragraph>
</subSubSection>
</treatment>
</document>