treatments-xml/data/03/A5/87/03A587E7A84EFFCB0CBA1528FCA5910A.xml
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<document id="8F5A428B6B9D6B772446E1B6AC0BBCEE" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A84EFFCB0CBA1528FCA5910A" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus denarius Perkins 2019, new species" docType="treatment" docVersion="7" lastPageNumber="48" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="46" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:title id="3C67B33796A06FB136D3A77886080876">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="7504021C9578B7FFE31AA3DE9FE770D5">Perkins, Philip D.</mods:namePart>
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<mods:title id="411B8BDDB197C42A9345FECA3DAB3F0A">Zootaxa</mods:title>
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<mods:date id="2715A16FEA99B86813FEDD58289FEEF1">2019</mods:date>
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<mods:number id="04C5236AEC11A96FD4898192CE443B3D">2019-12-12</mods:number>
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<mods:number id="F2F8AF2903EC34226442FC7497E7216F">4708</mods:number>
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<mods:identifier id="40D0A1C9E30CDCD643AC5B7A7C113BCF" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
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<treatment id="03A587E7A84EFFCB0CBA1528FCA5910A" ID-GBIF-Taxon="160798417" LSID="urn:lsid:plazi:treatment:03A587E7A84EFFCB0CBA1528FCA5910A" httpUri="http://treatment.plazi.org/id/03A587E7A84EFFCB0CBA1528FCA5910A" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
<subSubSection id="C316657AA84EFFC90CBA1528FDC495D2" box="[151,549,1900,1927]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BB336F1A84EFFC90CBA1528FDC495D2" blockId="45.[151,549,1900,1963]" box="[151,549,1900,1927]" pageId="45" pageNumber="46">
<heading id="D0FB819DA84EFFC90CBA1528FDC495D2" bold="true" box="[151,549,1900,1927]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis id="B978EAE3A84EFFC90CBA1528FDC495D2" bold="true" box="[151,549,1900,1927]" pageId="45" pageNumber="46">
<taxonomicName id="4C0C4D72A84EFFC90CBA1528FE6F95D2" ID-CoL="8Y4PV" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,398,1900,1927]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="denarius" status="sp. nov.">
<emphasis id="B978EAE3A84EFFC90CBA1528FE6F95D2" bold="true" box="[151,398,1900,1927]" italics="true" pageId="45" pageNumber="46">Hydrochus denarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A84EFFC90DB81529FDC495D2" box="[405,549,1901,1927]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA84EFFC90CBA15D5FEEC95FE" box="[151,269,1937,1963]" pageId="45" pageNumber="46" type="description">
<paragraph id="8BB336F1A84EFFC90CBA15D5FEEC95FE" blockId="45.[151,549,1900,1963]" box="[151,269,1937,1963]" pageId="45" pageNumber="46">
<figureCitation id="13372A74A84EFFC90CBA15D5FF0495FE" box="[151,229,1937,1963]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="45" pageNumber="46">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A84EFFC90CDC15D5FEEC95FE" box="[241,269,1937,1963]" captionStart="FIGURE 26" captionStartId="46.[151,250,2006,2031]" captionTargetBox="[202,1421,436,1967]" captionTargetId="figure@46.[151,1435,417,1981]" captionTargetPageId="46" captionText="FIGURE 26. Hydrochus denarius n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587235" httpUri="https://zenodo.org/record/3587235/files/figure.png" pageId="45" pageNumber="46">26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA84EFFCA0CBA159DFED293DE" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BB336F1A84EFFC90CBA159DFA7D95A6" blockId="45.[151,1436,2008,2035]" box="[151,1436,2008,2035]" pageId="45" pageNumber="46">
<emphasis id="B978EAE3A84EFFC90CBA159DFEA695A6" bold="true" box="[151,327,2008,2035]" pageId="45" pageNumber="46">Type Material.</emphasis>
<materialsCitation id="3B643CACA84EFFC90D61159DFA7D95A6" ID-GBIF-Occurrence="2521263750" box="[332,1436,2008,2035]" country="Venezuela" elevation="80" latitude="6.5936165" location="Bolivar State" longLatPrecision="1" longitude="-66.82063" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Bolivar" typeStatus="holotype">
<emphasis id="B978EAE3A84EFFC90D61159DFDF595A6" bold="true" box="[332,532,2008,2035]" pageId="45" pageNumber="46">
<typeStatus id="54B78853A84EFFC90D61159DFE5895A6" box="[332,441,2009,2035]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F31B7661A84EFFC90E09159DFD7F95A6" box="[548,670,2009,2035]" name="Venezuela" pageId="45" pageNumber="46">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A84EFFC90E8A159DFCDC95A6" box="[679,829,2009,2035]" country="Venezuela" name="Bolivar" pageId="45" pageNumber="46">Bolivar State</collectingRegion>
:
<geoCoordinate id="EE385036A84EFFC90F6B159DFC3795A6" box="[838,982,2008,2035]" degrees="6" direction="north" minutes="35.617" orientation="latitude" pageId="45" pageNumber="46" precision="1" value="6.5936165">6° 35.617 N</geoCoordinate>
,
<geoCoordinate id="EE385036A84EFFC90FCD159DFB6395A6" box="[992,1154,2009,2035]" degrees="66" direction="west" minutes="49.238" orientation="longitude" pageId="45" pageNumber="46" precision="1" value="-66.82063">66° 49.238 W</geoCoordinate>
,
<quantity id="4CF49B14A84EFFC908A6159DFB2195A6" box="[1163,1216,2009,2035]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="45" pageNumber="46" unit="m" value="80.0">
<elevation id="0021D1C2A84EFFC908A6159DFB2195A6" box="[1163,1216,2009,2035]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="45" pageNumber="46" unit="m" value="80.0">80m</elevation>
</quantity>
, Los Pijiguaos: out-
</materialsCitation>
</paragraph>
<paragraph id="8BB336F1A84DFFCA0CBA12DDFBF79282" blockId="46.[151,1437,152,395]" pageId="46" pageNumber="47">
crop/morichal,
<date id="FFB21031A84DFFCA0D6F12DDFE4F92E6" box="[322,430,153,179]" pageId="46" pageNumber="47" value="2009-01-12">12.i.2009</date>
, leg. Miller &amp; Short, detrital pools VZ09-0112-01C” (
<collectionCode id="ED1DAE34A84DFFCA082112DDFBBB92E6" box="[1036,1114,153,179]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="46" pageNumber="47" type="Museum">MIZA</collectionCode>
).
<materialsCitation id="3B643CACA84DFFCA084112DDFD319282" ID-GBIF-Occurrence="2521263667" collectionCode="SEMC, MIZA" country="Venezuela" location="Same" pageId="46" pageNumber="47" specimenCount="2" stateProvince="Bolivar" typeStatus="paratype">
<emphasis id="B978EAE3A84DFFCA084112DDFF189282" bold="true" pageId="46" pageNumber="47">
<typeStatus id="54B78853A84DFFCA084112DDFB0592E6" box="[1132,1252,153,179]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
(8):
<collectingCountry id="F31B7661A84DFFCA093612DDFA7992E6" box="[1307,1432,153,179]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A84DFFCA0CBA12F8FF159282" box="[151,244,188,215]" country="Venezuela" name="Bolivar" pageId="46" pageNumber="47">Bolivar</collectingRegion>
:
</emphasis>
<location id="8ED3602AA84DFFCA0D2D12F9FEDF9282" LSID="urn:lsid:plazi:treatment:03A587E7A84EFFCB0CBA1528FCA5910A:8ED3602AA84DFFCA0D2D12F9FEDF9282" box="[256,318,189,215]" country="Venezuela" name="Same" pageId="46" pageNumber="47" stateProvince="Bolivar">Same</location>
data as
<typeStatus id="54B78853A84DFFCA0DB412F9FE1A9282" box="[409,507,189,215]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
(7
<collectionCode id="ED1DAE34A84DFFCA0E0C12F9FD919282" box="[545,624,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="47" type="Museum">SEMC</collectionCode>
,
<collectionCode id="ED1DAE34A84DFFCA0E5612F9FD299282" box="[635,712,189,215]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="46" pageNumber="47" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA84DFFCA0EF712F9FBF39282" ID-GBIF-Occurrence="2521263666" box="[730,1042,189,215]" collectionCode="SEMC" country="Venezuela" pageId="46" pageNumber="47" specimenCode="VZ09-0112, 02" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<specimenCode id="DBAA9E8AA84DFFCA0EF712F9FCBF9282" box="[730,862,189,215]" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="46" pageNumber="47" type="Museum">VZ09-0112</specimenCode>
-
<specimenCode id="DBAA9E8AA84DFFCA0F4B12F9FC639282" box="[870,898,189,215]" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="46" pageNumber="47" type="Museum">02</specimenCode>
B (1
<collectionCode id="ED1DAE34A84DFFCA0F9412F9FBEB9282" box="[953,1034,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="46" pageNumber="47" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A84DFFCA0CEA12A4FED293DE" blockId="46.[151,1437,152,395]" pageId="46" pageNumber="47">
<emphasis id="B978EAE3A84DFFCA0CEA12A4FE3092AE" bold="true" box="[199,465,224,251]" pageId="46" pageNumber="47">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of small size (ca.
<quantity id="4CF49B14A84DFFCA090112A5FF26934A" metricMagnitude="-3" metricUnit="m" metricValue="2.32" metricValueMax="2.46" metricValueMin="2.18" pageId="46" pageNumber="47" unit="mm" value="2.32" valueMax="2.46" valueMin="2.18">2.182.46 mm</quantity>
), the elytra having 10 costate interstriae and with unusual, large translucent punctures at apices, the densely punctate pronotum with deep depressions and convex reliefs, the black dorsal color, and the male genitalia (
<figureCitation id="13372A74A84DFFCA0968136DFA769316" box="[1349,1431,297,323]" captionStart="FIGURE 26" captionStartId="46.[151,250,2006,2031]" captionTargetBox="[202,1421,436,1967]" captionTargetId="figure@46.[151,1435,417,1981]" captionTargetPageId="46" captionText="FIGURE 26. Hydrochus denarius n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587235" httpUri="https://zenodo.org/record/3587235/files/figure.png" pageId="46" pageNumber="47">Fig. 26</figureCitation>
; described below). Differentiation from other similarly small sized Venezuelan species easily made because of the costate elytra.
</paragraph>
</subSubSection>
<caption id="DF736679A84DFFCA0CBA1592FC2695BA" ID-DOI="http://doi.org/10.5281/zenodo.3587235" ID-Zenodo-Dep="3587235" box="[151,967,2006,2031]" httpUri="https://zenodo.org/record/3587235/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,2006,2031]" targetBox="[202,1421,436,1967]" targetPageId="46">
<paragraph id="8BB336F1A84DFFCA0CBA1592FC2695BA" blockId="46.[151,967,2006,2031]" box="[151,967,2006,2031]" pageId="46" pageNumber="47">
<emphasis id="B978EAE3A84DFFCA0CBA1592FEC195BA" bold="true" box="[151,288,2006,2031]" pageId="46" pageNumber="47">FIGURE 26.</emphasis>
<taxonomicName id="4C0C4D72A84DFFCA0D0B1593FE1595BA" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[294,500,2007,2031]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="denarius" status="sp. nov.">
<emphasis id="B978EAE3A84DFFCA0D0B1593FE1595BA" box="[294,500,2007,2031]" italics="true" pageId="46" pageNumber="47">Hydrochus denarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A84DFFCA0DD71593FDD395BB" box="[506,562,2007,2030]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<subSubSection id="C316657AA84CFFCB0CEA12DCFBEB916F" pageId="47" pageNumber="48" type="description">
<paragraph id="8BB336F1A84CFFCB0CEA12DCFCAC9317" blockId="47.[151,1437,152,863]" pageId="47" pageNumber="48">
<emphasis id="B978EAE3A84CFFCB0CEA12DCFEB092E6" bold="true" box="[199,337,152,179]" pageId="47" pageNumber="48">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A84CFFCB0E4912DCFD5B92E6" box="[612,698,152,179]" captionStart="FIGURE 26" captionStartId="46.[151,250,2006,2031]" captionTargetBox="[202,1421,436,1967]" captionTargetId="figure@46.[151,1435,417,1981]" captionTargetPageId="46" captionText="FIGURE 26. Hydrochus denarius n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587235" httpUri="https://zenodo.org/record/3587235/files/figure.png" pageId="47" pageNumber="48">Fig. 26</figureCitation>
). Size:
<typeStatus id="54B78853A84CFFCB0F3D12DCFC9392E7" box="[784,882,152,178]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.18 /0.88; head width 0.58; pronotum l/w 0.46/0.50; PA 0.50; PB 0.40; elytra 1.38 /0.88. Body size small (ca.
<quantity id="4CF49B14A84CFFCB0CBA12A4FEDC92AE" box="[151,317,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.32" metricValueMax="2.46" metricValueMin="2.18" pageId="47" pageNumber="48" unit="mm" value="2.32" valueMax="2.46" valueMin="2.18">2.182.46 mm</quantity>
). Dorsum black, with very weak iridescence. Legs brown, with tibiofemoral joints darker. Punctation on elytra ca. 3
<date id="FFB21031A84CFFCB0D691341FE81934A" box="[324,352,261,287]" pageId="47" pageNumber="48">4x</date>
that of pronotum. Elytra with odd numbered interstriae costate, costae ca. 0.5
<date id="FFB21031A84CFFCB08F11340FB15934A" box="[1244,1268,260,287]" pageId="47" pageNumber="48">1x</date>
strial puncture diameter; 5
<superScript id="7C799BB9A84CFFCB0D3B1363FEC29360" attach="left" box="[278,291,295,309]" fontSize="6" pageId="47" pageNumber="48">th</superScript>
costate, usual area of callus not otherwise raised.
</paragraph>
<paragraph id="8BB336F1A84CFFCB0CEA1309FE719387" blockId="47.[151,1437,152,863]" pageId="47" pageNumber="48">Pronotum slightly wider than long (as ca. 46/50), widest at anterior margin, markedly narrowed at base, sides very slightly sinuate, minutely denticulate, each denticle with a short seta; anterior margin weakly bisinuate, and with border of short, closely spaced setae; depressions deep and densely punctate, pits near anterior and posterior angles large and deep.</paragraph>
<paragraph id="8BB336F1A84CFFCB0CEA1398FF0C906A" blockId="47.[151,1437,152,863]" pageId="47" pageNumber="48">Elytra with spaces between strial punctures ca. 0.5x their diameter, to narrow ridges; apices widened and with submarginal groove having row of large, translucent punctures, another row of translucent punctures above groove.</paragraph>
<paragraph id="8BB336F1A84CFFCB0CEA100CFDDC90D3" blockId="47.[151,1437,152,863]" pageId="47" pageNumber="48">Ventral characters: Mentum coarsely punctate, with shallow median fovea. Submentum with two very large, confluent or nearly confluent, foveae.</paragraph>
<paragraph id="8BB336F1A84CFFCB0CEA10D4FB8F90A7" blockId="47.[151,1437,152,863]" pageId="47" pageNumber="48">
Male genitalia (
<figureCitation id="13372A74A84CFFCB0D5A10D4FE2890FE" box="[375,457,656,683]" captionStart="FIGURE 26" captionStartId="46.[151,250,2006,2031]" captionTargetBox="[202,1421,436,1967]" captionTargetId="figure@46.[151,1435,417,1981]" captionTargetPageId="46" captionText="FIGURE 26. Hydrochus denarius n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587235" httpUri="https://zenodo.org/record/3587235/files/figure.png" pageId="47" pageNumber="48">Fig. 26</figureCitation>
) general characters: paramere tips pointed, straight in dorsal and ventral views, slightly arcuate in lateral view, extending well beyond aedeagal apex; basal piece long, in dorsal/ventral views oriented at an angle to parameres, very arcuate in lateral view; aedeagus simple, lanceolate shaped.
</paragraph>
<paragraph id="8BB336F1A84CFFCB0CEA10B8FB7D9143" blockId="47.[151,1437,152,863]" box="[199,1180,764,791]" pageId="47" pageNumber="48">Dorsal surface: pdmm very slightly overlapping aedeagus lateral margins; adbl lacking.</paragraph>
<paragraph id="8BB336F1A84CFFCB0CEA1164FBEB916F" blockId="47.[151,1437,152,863]" box="[199,1034,800,827]" pageId="47" pageNumber="48">Ventral surface: pvmm very slightly overlapping aedeagus lateral margins.</paragraph>
</subSubSection>
<subSubSection id="C316657AA84CFFCB0CEA1101FCA5910A" box="[199,836,836,863]" pageId="47" pageNumber="48" type="etymology">
<paragraph id="8BB336F1A84CFFCB0CEA1101FCA5910A" blockId="47.[151,1437,152,863]" box="[199,836,836,863]" pageId="47" pageNumber="48">
<emphasis id="B978EAE3A84CFFCB0CEA1101FEAF910A" bold="true" box="[199,334,837,863]" pageId="47" pageNumber="48">Etymology.</emphasis>
Named in reference to the ten elytral costae.
</paragraph>
</subSubSection>
</treatment>
</document>