145 lines
17 KiB
XML
145 lines
17 KiB
XML
<document id="4045CA8C90D95A1DAA9A5FFEE3B85C8F" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A844FFCD0CBA1517FB7390A6" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus hellmani Perkins 2019, new species" docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="40" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:titleInfo id="A598335DE6C22733C12D6E15B5751091">
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<mods:title id="9BC6EB205293B0658E4E09691315ADA6">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="18C145AB8F7DBCF89E076E292DC4EFC7" type="personal">
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<mods:roleTerm id="17AD2BB16202EF423FEEC1BA813D1BAA">Author</mods:roleTerm>
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<mods:namePart id="6A602A3A7DC9018F3DC1B4CAD2699408">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="581F71D7CF938AF7134157B8BCC2AD3B">text</mods:typeOfResource>
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<mods:title id="FC713147716324E95353BB6F0AAAFBFD">Zootaxa</mods:title>
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<mods:part id="A4ECF7A7ABC81D65EF4053863B97A8BD">
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<mods:date id="1EAFA424B34B65D425844B3C1A7E533C">2019</mods:date>
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<mods:number id="9630C2F072D500657242C3E1BE75934B">2019-12-12</mods:number>
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<mods:number id="9125A94DBE278D5E06ACF037CF699BED">4708</mods:number>
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<mods:number id="C074427CD27506ECF094729F9D0C7A05">1</mods:number>
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<mods:start id="E1E4879A5EE7D6BE90629CB63F492253">1</mods:start>
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<mods:classification id="66150F257962E1033012E89744E19350">journal article</mods:classification>
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<mods:identifier id="16133389C9ECF1DBECF5893368134A67" type="CLB-Dataset">24540</mods:identifier>
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<mods:identifier id="CF54C6C8AA558D508C3F7C99742E6F35" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
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<mods:identifier id="347868040DDDF9FFEE36214E996C3DCC" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
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<mods:identifier id="A640F063B46E0E67FC43629693245F21" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A587E7A844FFCD0CBA1517FB7390A6" ID-DOI="http://doi.org/10.5281/zenodo.5941801" ID-GBIF-Taxon="160798421" ID-Zenodo-Dep="5941801" LSID="urn:lsid:plazi:treatment:03A587E7A844FFCD0CBA1517FB7390A6" httpUri="http://treatment.plazi.org/id/03A587E7A844FFCD0CBA1517FB7390A6" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40">
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<subSubSection id="C316657AA844FFC30CBA1517FDC69538" box="[151,551,1875,1902]" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="8BB336F1A844FFC30CBA1517FDC69538" blockId="39.[151,551,1875,1937]" box="[151,551,1875,1902]" pageId="39" pageNumber="40">
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<heading id="D0FB819DA844FFC30CBA1517FDC69538" bold="true" box="[151,551,1875,1902]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
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<emphasis id="B978EAE3A844FFC30CBA1517FDC69538" bold="true" box="[151,551,1875,1902]" pageId="39" pageNumber="40">
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<taxonomicName id="4C0C4D72A844FFC30CBA1517FE71953B" ID-CoL="8Y4QB" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,400,1875,1902]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hellmani" status="sp. nov.">
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<emphasis id="B978EAE3A844FFC30CBA1517FE71953B" bold="true" box="[151,400,1875,1902]" italics="true" pageId="39" pageNumber="40">Hydrochus hellmani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24B5798A844FFC30DBA1510FDC69538" box="[407,551,1875,1902]" pageId="39" pageNumber="40" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA844FFC30CBA1533FEEC95C4" box="[151,269,1911,1937]" pageId="39" pageNumber="40" type="description">
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<paragraph id="8BB336F1A844FFC30CBA1533FEEC95C4" blockId="39.[151,551,1875,1937]" box="[151,269,1911,1937]" pageId="39" pageNumber="40">
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<figureCitation id="13372A74A844FFC30CBA1533FF0295C4" box="[151,227,1911,1937]" captionStart="FIGURE 1" captionStartId="11.[151,250,1894,1919]" captionTargetBox="[156,1422,330,1824]" captionTargetId="figure@11.[151,1435,306,1866]" captionTargetPageId="11" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587183" httpUri="https://zenodo.org/record/3587183/files/figure.png" pageId="39" pageNumber="40">Figs. 1</figureCitation>
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,
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<figureCitation id="13372A74A844FFC30CDC1533FEEC95C4" box="[241,269,1911,1937]" captionStart="FIGURE 22" captionStartId="40.[151,250,1732,1757]" captionTargetBox="[174,1413,211,1635]" captionTargetId="figure@40.[151,1435,181,1706]" captionTargetPageId="40" captionText="FIGURE 22. Hydrochus hellmani n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587227" httpUri="https://zenodo.org/record/3587227/files/figure.png" pageId="39" pageNumber="40">22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA844FFCC0CBA1584FC0E95C4" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
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<paragraph id="8BB336F1A844FFC30CBA1584FEE695A8" blockId="39.[151,1437,1982,2045]" pageId="39" pageNumber="40">
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<emphasis id="B978EAE3A844FFC30CBA1584FEAC958C" bold="true" box="[151,333,1982,2010]" pageId="39" pageNumber="40">Type Material.</emphasis>
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<materialsCitation id="3B643CACA844FFC30D751584FEE695A8" ID-GBIF-Occurrence="2521263747" collectingDate="1982-10-31" collectionCode="NMNH" collectorName="John L. Hellman" country="Venezuela" location="Upata" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Bolivar" typeStatus="holotype">
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<emphasis id="B978EAE3A844FFC30D751584FD52958F" bold="true" box="[344,691,1982,2010]" pageId="39" pageNumber="40">
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<typeStatus id="54B78853A844FFC30D751584FE24958F" box="[344,453,1984,2010]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="F31B7661A844FFC30E1F1584FD4E958F" box="[562,687,1984,2010]" name="Venezuela" pageId="39" pageNumber="40">Venezuela</collectingCountry>
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:
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</emphasis>
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“
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<collectingRegion id="49C8F813A844FFC30EE415FBFCC5958C" box="[713,804,1983,2009]" country="Venezuela" name="Bolivar" pageId="39" pageNumber="40">Bolivar</collectingRegion>
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:
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<location id="8ED3602AA844FFC30F1E1584FC97958F" LSID="urn:lsid:plazi:treatment:03A587E7A844FFCD0CBA1517FB7390A6:8ED3602AA844FFC30F1E1584FC97958F" box="[819,886,1984,2010]" country="Venezuela" name="Upata" pageId="39" pageNumber="40" stateProvince="Bolivar">Upata</location>
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(
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<quantity id="4CF49B14A844FFC30FA415FBFC37958C" box="[905,982,1983,2009]" metricMagnitude="4" metricUnit="m" metricValue="6.9" pageId="39" pageNumber="40" unit="km" value="69.0">69 km</quantity>
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SE),
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<date id="FFB21031A844FFC3083315FBFB5D958C" box="[1054,1212,1983,2009]" pageId="39" pageNumber="40" value="1982-10-31">
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<collectingDate id="EFF6E9D9A844FFC3083315FBFB5D958C" box="[1054,1212,1983,2009]" pageId="39" pageNumber="40" value="1982-10-31">31 Oct. 1982</collectingDate>
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</date>
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,
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<collectorName id="26F95327A844FFC308E615FBFA73958C" box="[1227,1426,1983,2010]" pageId="39" pageNumber="40">John L. Hellman</collectorName>
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” (
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<collectionCode id="ED1DAE34A844FFC30CB215A7FF1A95A8" box="[159,251,2019,2045]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<caption id="DF736679A84BFFCC0CBA1480FC269488" ID-DOI="http://doi.org/10.5281/zenodo.3587227" ID-Zenodo-Dep="3587227" box="[151,967,1732,1757]" httpUri="https://zenodo.org/record/3587227/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1732,1757]" targetBox="[174,1413,211,1635]" targetPageId="40">
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<paragraph id="8BB336F1A84BFFCC0CBA1480FC269488" blockId="40.[151,967,1732,1757]" box="[151,967,1732,1757]" pageId="40" pageNumber="41">
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<emphasis id="B978EAE3A84BFFCC0CBA1480FEC19488" bold="true" box="[151,288,1732,1757]" pageId="40" pageNumber="41">FIGURE 22.</emphasis>
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<taxonomicName id="4C0C4D72A84BFFCC0D0B1480FE159489" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[294,500,1732,1756]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hellmani" status="sp. nov.">
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<emphasis id="B978EAE3A84BFFCC0D0B1480FE159489" box="[294,500,1732,1756]" italics="true" pageId="40" pageNumber="41">Hydrochus hellmani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24B5798A84BFFCC0DD71481FDD39489" box="[506,562,1733,1756]" pageId="40" pageNumber="41" rank="species">n. sp.</taxonomicNameLabel>
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, habitus and male genitalia of holotype.
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</paragraph>
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</caption>
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<paragraph id="8BB336F1A84BFFCC0CEA154EFC0E95C4" blockId="40.[151,1437,1802,2045]" pageId="40" pageNumber="41">
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<emphasis id="B978EAE3A84BFFCC0CEA154EFE379570" bold="true" box="[199,470,1802,1829]" pageId="40" pageNumber="41">Differential Diagnosis.</emphasis>
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Among Venezuelan species, recognized by the combination of small size (ca.
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<quantity id="4CF49B14A84BFFCC0946154FFF26951C" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="40" pageNumber="41" unit="mm" value="2.37">2.37 mm</quantity>
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), the pronotum with moderately deep impressions separated by narrow reliefs, the elytra with large and deep strial punctures, interstriae and spaces between punctures narrow ridges, and the male genitalia (
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<figureCitation id="13372A74A84BFFCC08FC1517FAC49538" box="[1233,1317,1875,1901]" captionStart="FIGURE 22" captionStartId="40.[151,250,1732,1757]" captionTargetBox="[174,1413,211,1635]" captionTargetId="figure@40.[151,1435,181,1706]" captionTargetPageId="40" captionText="FIGURE 22. Hydrochus hellmani n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587227" httpUri="https://zenodo.org/record/3587227/files/figure.png" pageId="40" pageNumber="41">Fig. 22</figureCitation>
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; described below). Completely reliable determinations will include dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA84BFFCD0CEA15DEFCA8909A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
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<paragraph id="8BB336F1A84BFFCD0CEA15DEFEDB92AE" blockId="40.[151,1437,1802,2045]" lastBlockId="41.[151,1437,153,755]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<emphasis id="B978EAE3A84BFFCC0CEA15DEFEB095E0" bold="true" box="[199,337,1946,1973]" pageId="40" pageNumber="41">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="13372A74A84BFFCC0E4D15DFFD5795E0" box="[608,694,1947,1973]" captionStart="FIGURE 22" captionStartId="40.[151,250,1732,1757]" captionTargetBox="[174,1413,211,1635]" captionTargetId="figure@40.[151,1435,181,1706]" captionTargetPageId="40" captionText="FIGURE 22. Hydrochus hellmani n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587227" httpUri="https://zenodo.org/record/3587227/files/figure.png" pageId="40" pageNumber="41">Fig. 22</figureCitation>
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). Size:
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<typeStatus id="54B78853A84BFFCC0F2615DFFC8C95E0" box="[779,877,1947,1973]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
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male (length/width, mm): body (length to elytral apices) 2.37/0.92; head width 0.60; pronotum l/w 0.53/0.57; PA 0.55; PB 0.47; elytra 1.50/0.92. Body size small (ca.
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<quantity id="4CF49B14A84BFFCC0CE515A7FED095A8" box="[200,305,2019,2045]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="40" pageNumber="41" unit="mm" value="2.37">2.37 mm</quantity>
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). Dorsum black, with slight iridescence, elytra without black spots. Legs dark brown. Punctation on elytra large, deep, ca. 2–
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<date id="FFB21031A84AFFCD0D8512DDFE2392E6" box="[424,450,153,179]" pageId="41" pageNumber="42">3x</date>
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that of pronotum. Elytra interstriae much narrower than strial puncture diameter, narrow ridges. Interstria 9
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<superScript id="7C799BB9A84AFFCD0D4A12F8FE95929F" attach="left" box="[359,372,188,202]" fontSize="6" pageId="41" pageNumber="42">th</superScript>
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more convex than others, weakly overhanging 10
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<superScript id="7C799BB9A84AFFCD0F8212F8FC5D929F" attach="left" box="[943,956,188,202]" fontSize="6" pageId="41" pageNumber="42">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C799BB9A84AFFCD097812F8FA83929F" attach="left" box="[1365,1378,188,202]" fontSize="6" pageId="41" pageNumber="42">th</superScript>
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very weakly raised.
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</paragraph>
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<paragraph id="8BB336F1A84AFFCD0CEA1341FAE29316" blockId="41.[151,1437,153,755]" pageId="41" pageNumber="42">Pronotum slightly wider than long (as ca. 53/57), widest at anterior margin, narrowed at base, sides very weakly arcuate, smooth; anterior margin very weakly bisinuate; depressions moderately deep, reliefs narrow.</paragraph>
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<paragraph id="8BB336F1A84AFFCD0CEA1309FCF293DE" blockId="41.[151,1437,153,755]" pageId="41" pageNumber="42">Elytra with spaces between strial punctures narrow ridges; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
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<paragraph id="8BB336F1A84AFFCD0CEA13D1FD629386" blockId="41.[151,1437,153,755]" pageId="41" pageNumber="42">Ventral characters: Mentum moderately coarsely and moderately densely punctate, with shallow median fovea. Submentum with two small shallow foveae.</paragraph>
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<paragraph id="8BB336F1A84AFFCD0CEA1399FC9C90D2" blockId="41.[151,1437,153,755]" pageId="41" pageNumber="42">
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Male genitalia (
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<figureCitation id="13372A74A84AFFCD0D561399FE2F93A2" box="[379,462,477,503]" captionStart="FIGURE 22" captionStartId="40.[151,250,1732,1757]" captionTargetBox="[174,1413,211,1635]" captionTargetId="figure@40.[151,1435,181,1706]" captionTargetPageId="40" captionText="FIGURE 22. Hydrochus hellmani n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587227" httpUri="https://zenodo.org/record/3587227/files/figure.png" pageId="41" pageNumber="42">Fig. 22</figureCitation>
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) general characters: genitalia stout; paramere tips tapering and arcing slightly toward midline, extending slightly beyond distal end of aedeagus, parameres in lateral view arcuate, and wider than dorsal/ ventral views; basal piece much shorter than parameres, tapering from distal end to base, orifice sclerotized, asymmetrical; aedeagus lanceolate, lateral margins moderately widely sclerotized, greatest width at about midlength, tapering apically, with internal lumen in basal ½, gonopore apical.
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</paragraph>
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<paragraph id="8BB336F1A84AFFCD0CEA10D5FA6F90FE" blockId="41.[151,1437,153,755]" box="[199,1422,657,683]" pageId="41" pageNumber="42">Dorsal surface: adtl subapical; pdmm widely overlapping alm, especially in distal 1/2; adbl moderately large.</paragraph>
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<paragraph id="8BB336F1A84AFFCD0CEA10F1FCA8909A" blockId="41.[151,1437,153,755]" box="[199,841,693,719]" pageId="41" pageNumber="42">Ventral surface: avtl subbasal; pvmm approximating alm.</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA84AFFCD0CEA109DFB7390A6" box="[199,1170,729,755]" pageId="41" pageNumber="42" type="etymology">
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<paragraph id="8BB336F1A84AFFCD0CEA109DFB7390A6" blockId="41.[151,1437,153,755]" box="[199,1170,729,755]" pageId="41" pageNumber="42">
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<emphasis id="B978EAE3A84AFFCD0CEA109DFEAF90A6" bold="true" box="[199,334,729,755]" pageId="41" pageNumber="42">Etymology.</emphasis>
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Named for John Leroy Hellman, who collected the only known specimen.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |