treatments-xml/data/03/A5/87/03A587D1FFAE58451FB310531B6EFAC5.xml
2024-06-21 12:22:17 +02:00

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<document id="AE2F64BDE0ACD7644249DBA11E9C90FA" ID-CLB-Dataset="22914" ID-DOI="10.5281/zenodo.3758425" ID-GBIF-Dataset="7ac1366d-0143-441f-89f5-4a0bbba2a48a" ID-ISSN="0253-116X" ID-Zenodo-Dep="3758425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587401510267" checkinUser="valdenar" docAuthor="Schmid-Egger, Christian" docDate="2019" docId="03A587D1FFAE58451FB310531B6EFAC5" docLanguage="en" docName="Linzer.biol.Beitr.51.1.459-471.pdf.imf" docOrigin="Linzer biologische Beiträge 51 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Sphex afer Schmid-Egger 2019" docType="treatment" docVersion="6" lastPageNumber="462" masterDocId="FF9CFFA9FFAD58461F01114A197BFFED" masterDocTitle="A review oI the Sphex flavipennis species group (Hymenoptera, Sphecidae) in northwest AIrica with description oI two new species and a discussion oI a species Irom Sardinia" masterLastPageNumber="471" masterPageNumber="459" pageNumber="462" updateTime="1698834177235" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="51CE8AC88466EB77E277286F7D649486">A review oI the Sphex flavipennis species group (Hymenoptera, Sphecidae) in northwest AIrica with description oI two new species and a discussion oI a species Irom Sardinia</mods:title>
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<mods:namePart id="0BBA9B3F947488F6F89AAF56F0E8C9BE">Schmid-Egger, Christian</mods:namePart>
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<mods:title id="60F3C5FCBE62F3036A6C01CD48518AE6">Linzer biologische Beiträge</mods:title>
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<subSubSection id="C316654CFFAE58451FB31053180AFE69" pageId="3" pageNumber="462" type="nomenclature">
<paragraph id="8BB336C7FFAE58451FB310531ABFFEC3" blockId="3.[178,964,280,303]" box="[178,964,280,303]" pageId="3" pageNumber="462">
<heading id="D0FB81ABFFAE58451FB310531ABFFEC3" box="[178,964,280,303]" fontSize="8" level="2" pageId="3" pageNumber="462" reason="2">
<emphasis id="B978EAD5FFAE58451FB310531BFBFEC0" bold="true" box="[178,640,280,303]" pageId="3" pageNumber="462">
<taxonomicName id="4C0C4D44FFAE58451FB310531858FEC2" authority="LEPELETIER de SAINT FARGEAU, 1845" authorityName="Schmid-Egger" authorityYear="2019" box="[178,291,281,303]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451FB310531858FEC2" bold="true" box="[178,291,281,303]" italics="true" pageId="3" pageNumber="462">Sphex afer</emphasis>
</taxonomicName>
LEPELETIER de SAINT FARGEAU
</emphasis>
,
<emphasis id="B978EAD5FFAE58451D8D10531ABFFEC3" bold="true" box="[652,964,280,302]" pageId="3" pageNumber="462">1845, stat.rest. (figs 5, 15, 20)</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336C7FFAE58451FB3100A180AFE69" blockId="3.[178,1086,318,469]" pageId="3" pageNumber="462">
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<treatmentCitation id="0AAD10D6FFAE58451FB3100A1BD4FEB9" box="[178,687,318,340]" pageId="3" pageNumber="462">
<taxonomicName id="4C0C4D44FFAE58451FB3100A1868FEB9" authorityName="Schmid-Egger" authorityYear="2019" box="[178,275,320,340]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451FB3100A1868FEB9" box="[178,275,320,340]" italics="true" pageId="3" pageNumber="462">Sphex afer</emphasis>
</taxonomicName>
LEPELETIER DE SAINT FARGEAU, 1845: 350
</treatmentCitation>
,
<specimenCount id="9D0AFD4EFFAE58451DB810751BB1FEB9" box="[697,714,319,340]" pageId="3" pageNumber="462" type="female"></specimenCount>
(as
<emphasis id="B978EAD5FFAE58451DF0100A1A6CFEB9" box="[753,791,320,340]" italics="true" pageId="3" pageNumber="462">afra</emphasis>
, incorrect original termination).
<typeStatus id="54B78865FFAE58451FE6101D183EFE81" box="[231,325,343,364]" pageId="3" pageNumber="462" type="lectotype">Lectotype</typeStatus>
: female,
<collectingCountry id="F31B7657FFAE58451E9C101C189EFE81" box="[413,485,342,364]" name="Algeria" pageId="3" pageNumber="462">Algeria</collectingCountry>
: Oran (MNHN).
<location id="8ED3601CFFAE58451D88101C1BA1FE81" LSID="urn:lsid:plazi:treatment:03A587D1FFAE58451FB310531B6EFAC5:8ED3601CFFAE58451D88101C1BA1FE81" box="[649,730,342,364]" country="Algeria" name="Restored" pageId="3" pageNumber="462">Restored</location>
from synonymy of
<taxonomicName id="4C0C4D44FFAE58451C9A10121D46FE81" authority="BRULLÉ, 1833" box="[923,1085,344,364]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451C9A10121D46FE81" box="[923,1085,344,364]" italics="true" pageId="3" pageNumber="462">Sphex leuconotus</emphasis>
</taxonomicName>
BRULLÉ, 1833.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C316654CFFAE58451FB310C51B6EFAC5" pageId="3" pageNumber="462" type="description">
<paragraph id="8BB336C7FFAE58451FB310C51BACFE38" blockId="3.[178,1086,318,469]" pageId="3" pageNumber="462">
M a t e r i a l e x a m i n e d:
<taxonomicName id="4C0C4D44FFAE58451EF110DA1B53FE49" authorityName="Schmid-Egger" authorityYear="2019" box="[496,552,400,420]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451EF110DA1B53FE49" box="[496,552,400,420]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
:
<collectingCountry id="F31B7657FFAE58451D3210C51BF9FE49" box="[563,642,399,420]" name="Morocco" pageId="3" pageNumber="462">Morocco</collectingCountry>
<specimenCount id="9D0AFD4EFFAE58451D8910C41BBFFE49" box="[648,708,398,420]" pageId="3" pageNumber="462" type="male">1 male</specimenCount>
,
<specimenCount id="9D0AFD4EFFAE58451DCC10C41A62FE49" box="[717,793,398,420]" pageId="3" pageNumber="462" type="female">1 female</specimenCount>
<date id="FFB21007FFAE58451C1F10C41A08FE49" box="[798,883,398,420]" pageId="3" pageNumber="462" value="2013-06-03">3.vi.2013</date>
. S Ait Mzizoui
<geoCoordinate id="EE385000FFAE58451CFD10C41D46FE4E" box="[1020,1085,398,419]" degrees="34,47" direction="north" orientation="latitude" pageId="3" pageNumber="462" precision="555" value="34.47">34,47N</geoCoordinate>
<geoCoordinate id="EE385000FFAE58451FDE10EC185EFE51" box="[223,293,422,444]" degrees="13,35" direction="west" orientation="longitude" pageId="3" pageNumber="462" precision="555" value="-13.35">13,35W</geoCoordinate>
(leg. et coll. T. Ljubomirov).
<taxonomicName id="4C0C4D44FFAE58451D4110E21BC3FE51" box="[576,696,424,444]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451D4110E21BC3FE51" box="[576,696,424,444]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
was examined from
<collectingCountry id="F31B7657FFAE58451C7F10ED1AC5FE51" box="[894,958,423,444]" name="Greece" pageId="3" pageNumber="462">Greece</collectingCountry>
, Kazsachstan,
<collectingCountry id="F31B7657FFAE58451FDE10F51838FE39" box="[223,323,447,468]" name="Kyrgyzstan" pageId="3" pageNumber="462">Kyrgyzstan</collectingCountry>
and
<collectingCountry id="F31B7657FFAE58451E6C10F418D7FE39" box="[365,428,446,468]" name="Turkey" pageId="3" pageNumber="462">Turkey</collectingCountry>
(
<specimenCount id="9D0AFD4EFFAE58451EB910F51B74FE39" box="[440,527,446,468]" pageId="3" pageNumber="462" type="female">2 females</specimenCount>
,
<specimenCount id="9D0AFD4EFFAE58451D1910F51B26FE39" box="[536,605,446,468]" pageId="3" pageNumber="462" type="male">4 males</specimenCount>
, in coll CSE).
</paragraph>
<paragraph id="8BB336C7FFAE58451FB310A9185EFD45" blockId="3.[178,1085,483,681]" pageId="3" pageNumber="462">
R e m a r k: MENKE &amp; PULAWSKI (2000) synonymised
<taxonomicName id="4C0C4D44FFAE58451C1C10AE1A1CFE14" authorityName="Schmid-Egger" authorityYear="2019" box="[797,871,484,505]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451C1C10AE1A1CFE14" box="[797,871,484,505]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
with
<taxonomicName id="4C0C4D44FFAE58451CAF10AE1D46FE14" box="[942,1085,484,505]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451CAF10AE1D46FE14" box="[942,1085,484,505]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
BRULLÉ, 1833, described from
<collectingCountry id="F31B7657FFAE58451D07134A1B2AFDF8" box="[518,593,512,533]" name="Greece" pageId="3" pageNumber="462">Greece</collectingCountry>
. However, the present investigation revealed distinct morphological differences in a male from
<collectingCountry id="F31B7657FFAE58451DC013541A67FDDE" box="[705,796,542,563]" name="Morocco" pageId="3" pageNumber="462">Morocco</collectingCountry>
compared with males from
<collectingCountry id="F31B7657FFAE58451FB313711981FDBD" box="[178,250,571,592]" name="Greece" pageId="3" pageNumber="462">Greece</collectingCountry>
and Central Asia. Apex of S8 is clearly different in these males. For that reason, I consider the male from
<collectingCountry id="F31B7657FFAE58451EAF13121B72FD80" box="[430,521,600,621]" name="Morocco" pageId="3" pageNumber="462">Morocco</collectingCountry>
as belonging to a different species.
<taxonomicName id="4C0C4D44FFAE58451C8A13131A81FD83" authorityName="Schmid-Egger" authorityYear="2019" box="[907,1018,601,622]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451C8A13131A81FD83" box="[907,1018,601,622]" italics="true" pageId="3" pageNumber="462">Sphex afer</emphasis>
</taxonomicName>
is the next available name for it, and the taxon has to be restored from synonymy of
<taxonomicName id="4C0C4D44FFAE58451B2B133D185BFD44" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451B2B133D185BFD44" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB336C7FFAE58451FB313F21AA8FC8C" blockId="3.[178,1086,696,865]" pageId="3" pageNumber="462">
The results are also seen against the background that form of S8 is an important character for species recognition in the
<taxonomicName id="4C0C4D44FFAE58451EE0139D1B12FD01" authorityName="Fabricius" authorityYear="1793" box="[481,617,727,748]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="flavipennis">
<emphasis id="B978EAD5FFAE58451EE0139D1B12FD01" box="[481,617,727,748]" italics="true" pageId="3" pageNumber="462">S. flavipennis</emphasis>
</taxonomicName>
species group, clearly different in all species. Another important hint for valuation of the taxonomic rank of these taxa is the large genetic gap between specimens from Europe and from Africa in many
<taxonomicName id="4C0C4D44FFAE58451C92125A1D6BFCCB" authorityName="Latreille" authorityYear="1802" box="[915,1040,784,806]" class="Insecta" family="Crabronidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="family">Crabronidae</taxonomicName>
and
<taxonomicName id="4C0C4D44FFAE58451FB312671861FCAE" authorityName="Latreille" authorityYear="1802" box="[178,282,813,835]" class="Insecta" family="Sphecidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="family">Sphecidae</taxonomicName>
species which makes a state as valid species of the African populations in most of these taxa probable (see discussion under SCHMID- EGGER et. al. 2018).
</paragraph>
<paragraph id="8BB336C7FFAE58451FB3123B1BB5FB9C" blockId="3.[178,1085,880,1137]" pageId="3" pageNumber="462">
R e c o g n i t i o n: The male of
<taxonomicName id="4C0C4D44FFAE58451D1F12381B18FC6B" authorityName="Schmid-Egger" authorityYear="2019" box="[542,611,881,903]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451D1F12381B18FC6B" box="[542,611,881,903]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
has S8 with an obtuse apical angle (fig. 15), whereas the apical angle is acute in
<taxonomicName id="4C0C4D44FFAE58451D2012C51BD2FC49" box="[545,681,911,932]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451D2012C51BD2FC49" box="[545,681,911,932]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
(fig. 14,
<specimenCount id="9D0AFD4EFFAE58451C0612C41A2DFC4E" box="[775,854,909,931]" pageId="3" pageNumber="462" type="male">4 males</specimenCount>
examined). Wings are dark in
<taxonomicName id="4C0C4D44FFAE58451E0012E6183FFC2C" authorityName="Schmid-Egger" authorityYear="2019" box="[257,324,940,961]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451E0012E6183FFC2C" box="[257,324,940,961]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
, and are somewhat yellowish in the
<taxonomicName id="4C0C4D44FFAE58451DBA12E61A3FFC2C" box="[699,836,940,961]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451DBA12E61A3FFC2C" box="[699,836,940,961]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
males. The female of
<taxonomicName id="4C0C4D44FFAE58451B2B12E619A0FC33" authorityName="Schmid-Egger" authorityYear="2019" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451B2B12E619A0FC33" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
is larger (body length
<quantity id="4CF49B22FFAE58451ED512821B5BFC33" box="[468,544,968,990]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="462" unit="mm" value="30.0">30 mm</quantity>
) than both examined
<taxonomicName id="4C0C4D44FFAE58451C0A12801AEDFC33" box="[779,918,969,991]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451C0A12801AEDFC33" box="[779,918,969,991]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
females (
<quantity id="4CF49B22FFAE58451B01128219A0FC16" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="3" pageNumber="462" unit="mm" value="24.5" valueMax="25.0" valueMin="24.0">24-25 mm</quantity>
) from Central Asia, wings are dark and only slightly yellow in colour, whereas they are distinctly yellow in
<taxonomicName id="4C0C4D44FFAE58451EAF154E1B42FBF4" box="[430,569,1028,1049]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451EAF154E1B42FBF4" box="[430,569,1028,1049]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
. Propodeal setae are clearly white and gaster all black in
<taxonomicName id="4C0C4D44FFAE58451E11156818E1FBDB" box="[272,410,1057,1079]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451E11156818E1FBDB" box="[272,410,1057,1079]" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
, and greyish in
<taxonomicName id="4C0C4D44FFAE58451D4A15681BEBFBDB" authorityName="Schmid-Egger" authorityYear="2019" box="[587,656,1057,1079]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451D4A15681BEBFBDB" box="[587,656,1057,1079]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
with a red gaster base in
<taxonomicName id="4C0C4D44FFAE58451CAB15681A8BFBDB" authorityName="Schmid-Egger" authorityYear="2019" box="[938,1008,1057,1079]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451CAB15681A8BFBDB" box="[938,1008,1057,1079]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
. These characters has to be verified with a larger sample, and especially the tergal colour is variable in both species (MENKE &amp; PULAWSKI 2000).
</paragraph>
<paragraph id="8BB336C7FFAE58451FB315CA1B6EFAC5" blockId="3.[178,1085,1152,1321]" pageId="3" pageNumber="462">
G e o g r a f i c d i s t r i b u t i o n: The general distribution of
<taxonomicName id="4C0C4D44FFAE58451C7F15C81ABEFB7B" authorityName="Schmid-Egger" authorityYear="2019" box="[894,965,1153,1175]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451C7F15C81ABEFB7B" box="[894,965,1153,1175]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
is unclear, because I could only examine a single male and female from
<collectingCountry id="F31B7657FFAE58451C2E15D41AF5FB5E" box="[815,910,1182,1203]" name="Morocco" pageId="3" pageNumber="462">Morocco</collectingCountry>
. However, information from literature suggests that
<taxonomicName id="4C0C4D44FFAE58451D3615F61B05FB3C" authorityName="Schmid-Egger" authorityYear="2019" box="[567,638,1212,1233]" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B978EAD5FFAE58451D3615F61B05FB3C" box="[567,638,1212,1233]" italics="true" pageId="3" pageNumber="462">S. afer</emphasis>
</taxonomicName>
occurs in whole North Africa apart from
<collectingCountry id="F31B7657FFAE58451FB315931994FB03" box="[178,239,1241,1262]" name="Egypt" pageId="3" pageNumber="462">Egypt</collectingCountry>
(MENKE &amp; PULAWSKI 2000), and the European and Asian populations belong to
<taxonomicName id="4C0C4D44FFAE58451B2B1590185BFAE1" class="Insecta" family="Pompilidae" genus="Sphex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="leuconotus">
<emphasis id="B978EAD5FFAE58451B2B1590185BFAE1" italics="true" pageId="3" pageNumber="462">S. leuconotus</emphasis>
</taxonomicName>
. The distribution area of the latter reaches
<collectingCountry id="F31B7657FFAE58451DEE15BF1A52FAE6" box="[751,809,1269,1291]" name="Spain" pageId="3" pageNumber="462">Spain</collectingCountry>
in the West,
<collectingCountry id="F31B7657FFAE58451CBF15BF1A8DFAE6" box="[958,1014,1269,1291]" name="Israel" pageId="3" pageNumber="462">Israel</collectingCountry>
in the South and Kazsachstan in the East.
</paragraph>
</subSubSection>
</treatment>
</document>