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<document id="1B4B3F23288F0B0D31511B17C0753B3F" ID-DOI="10.5281/zenodo.174076" ID-GBIF-Dataset="50e26ef8-c2f5-47ff-a6bd-b6b34323800b" ID-ISSN="1175­5326" ID-Zenodo-Dep="174076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459789464809" checkinUser="plazi" docAuthor="Schwendinger, Peter J." docDate="2006" docId="03A5879D7538FF9BFE985FE5DE332DC4" docLanguage="en" docName="zt01325p266.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Martensiellus" docType="treatment" docVersion="9" lastPageNumber="263" masterDocId="FF9CFFE5753FFF93FF905E66DD6C2F5D" masterDocTitle="A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores)" masterLastPageNumber="266" masterPageNumber="255" pageNumber="262" updateTime="1698219045570" updateUser="plazi">
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<mods:title id="DF0EFC314434BA2F01E53D9C28624451">A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores)</mods:title>
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<mods:namePart id="7FC67243BC138436AE3FEF15B26B18D4">Schwendinger, Peter J.</mods:namePart>
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<subSubSection id="C31665007538FF94FE985FE5DCE62E9C" pageId="7" pageNumber="262" type="nomenclature">
<paragraph id="8BB3368B7538FF94FE985FE5DF5F2EC0" blockId="7.[264,563,387,449]" box="[264,563,387,414]" pageId="7" pageNumber="262">
<heading id="D0FB81E77538FF94FE985FE5DF5F2EC0" bold="true" box="[264,563,387,414]" fontSize="11" level="1" pageId="7" pageNumber="262" reason="1">
<emphasis id="B978EA997538FF94FE985FE5DF5F2EC0" bold="true" box="[264,563,387,414]" pageId="7" pageNumber="262">
?
<taxonomicName id="4C0C4D087538FF94FE875FE2DCD12EC3" ID-CoL="7PNJ7" box="[279,445,388,414]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997538FF94FE875FE2DCD12EC3" bold="true" box="[279,445,388,414]" italics="true" pageId="7" pageNumber="262">Martensiellus</emphasis>
</taxonomicName>
n. g. spp.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB3368B7538FF94FE985FC1DCE62E9C" blockId="7.[264,563,387,449]" box="[264,394,423,449]" pageId="7" pageNumber="262">
<figureCitation id="13372A0E7538FF94FE985FC1DCE62E9C" box="[264,394,423,449]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="7" pageNumber="262">Figs 1822</figureCitation>
</paragraph>
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<subSubSection id="C31665007538FF9BFE985F97DE332DC4" lastPageId="8" lastPageNumber="263" pageId="7" pageNumber="262" type="description">
<paragraph id="8BB3368B7538FF94FE985F97DCC62DA6" blockId="7.[264,1323,497,763]" pageId="7" pageNumber="262">
Two females from other parts of Borneo possess the same characteristics in somatic morphology (tarsal formula 1­1­2­2, leg tarsi/tarsomeres fairly large, a tarsal pore present on legs I and II) that place
<taxonomicName id="4C0C4D087538FF94FDD75C27DF942D07" box="[583,760,577,602]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA997538FF94FDD75C27DF942D07" box="[583,760,577,602]" italics="true" pageId="7" pageNumber="262">M. tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997538FF94FD6F5C27DE2E2D06" bold="true" box="[767,834,577,603]" pageId="7" pageNumber="262">
<taxonomicNameLabel id="A24B57E27538FF94FD6F5C27DE2E2D06" box="[767,834,577,603]" pageId="7" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
apart from all other described oncopodid species. I assume that these females also belong to
<taxonomicName id="4C0C4D087538FF94FCFD5C0FD9622DDF" box="[877,1038,617,642]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997538FF94FCFD5C0FD9622DDF" box="[877,1038,617,642]" italics="true" pageId="7" pageNumber="262">Martensiellus</emphasis>
</taxonomicName>
<emphasis id="B978EA997538FF94FB875C0FD9222DDE" bold="true" box="[1047,1102,617,643]" pageId="7" pageNumber="262">n. g.</emphasis>
and represent two undescribed species distinct from
<taxonomicName id="4C0C4D087538FF94FD225CF7DE052DF7" box="[690,873,657,682]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA997538FF94FD225CF7DE052DF7" box="[690,873,657,682]" italics="true" pageId="7" pageNumber="262">M. tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997538FF94FCE75CF7DEAD2DF6" bold="true" box="[887,961,657,683]" pageId="7" pageNumber="262">
<taxonomicNameLabel id="A24B57E27538FF94FCE75CF7DEAD2DF6" box="[887,961,657,683]" pageId="7" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
However, generic placement without conspecific males is uncertain and I therefore refrain from formally describing and naming them.
</paragraph>
<caption id="DF7366037538FF94FE985B8DD84C293E" httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="7" pageNumber="262" targetBox="[271,1312,809,1489]" targetPageId="7">
<paragraph id="8BB3368B7538FF94FE985B8DD84C293E" blockId="7.[264,1323,1515,1635]" pageId="7" pageNumber="262">
<emphasis id="B978EA997538FF94FE985B8DDCBA295E" bold="true" box="[264,470,1515,1539]" pageId="7" pageNumber="262">FIGURES 1822.</emphasis>
?
<taxonomicName id="4C0C4D087538FF94FE7F5B8ADFEE295E" box="[495,642,1516,1539]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997538FF94FE7F5B8ADFEE295E" box="[495,642,1516,1539]" italics="true" pageId="7" pageNumber="262">Martensiellus</emphasis>
</taxonomicName>
<emphasis id="B978EA997538FF94FD1F5B8DDF90295E" bold="true" box="[655,764,1515,1539]" pageId="7" pageNumber="262">n. g. spp</emphasis>
., female from Mulu N.P. (18, 19), female from Kaharian (2022). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (1819, 2022 same scale).
</paragraph>
</caption>
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1. female (MHNG): Eastern
<collectingCountry id="F31B761B7538FF94FD5658E8DE5829F5" box="[710,820,1678,1704]" name="Malaysia" pageId="7" pageNumber="262">Malaysia</collectingCountry>
,
<collectingCountry id="F31B761B7538FF94FCDF58E8DEDB29F5" box="[847,951,1678,1704]" name="Malaysia" pageId="7" pageNumber="262">Sarawak</collectingCountry>
, Mulu National Park, near headquarters (
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,
<geoCoordinate id="EE38504C7538FF94FD8258D0DFE9298D" box="[530,645,1718,1744]" direction="east" orientation="longitude" pageId="7" pageNumber="262" precision="925" value="114.816666">114°49E</geoCoordinate>
),
<quantity id="4CF49B6E7538FF94FD0A58D0DF8A298D" box="[666,742,1718,1744]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="262" unit="m" value="200.0">200 m</quantity>
, collected on
<date id="FFB2104B7538FF94FC1558D0D907298D" box="[901,1131,1718,1744]" pageId="7" pageNumber="262" value="2003-08-19" valueMax="2003-08-24" valueMin="2003-08-19">1924 August 2003</date>
from rain forest leaf litter by A. Schulz. Different from the males of
<taxonomicName id="4C0C4D087538FF94FC3258B8D93429AA" box="[930,1112,1758,1783]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA997538FF94FC3258B8D93429AA" box="[930,1112,1758,1783]" italics="true" pageId="7" pageNumber="262">M. tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997538FF94FBF458BBD9C029AA" bold="true" box="[1124,1196,1757,1783]" pageId="7" pageNumber="262">
<taxonomicNameLabel id="A24B57E27538FF94FBF458BBD9C029AA" box="[1124,1196,1757,1783]" pageId="7" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by: Body distinctly smaller (
<quantity id="4CF49B6E7538FF94FE745963DF232842" box="[484,591,1797,1823]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="7" pageNumber="262" unit="mm" value="2.75">2.75 mm</quantity>
long); eye tubercle more conical (
<figureCitation id="13372A0E7538FF94FC785963D92C2842" box="[1000,1088,1797,1823]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="7" pageNumber="262">Fig. 18</figureCitation>
); legs more slender and palpal tibia thicker; carapace­opisthosoma bridge present, divided; tiny process on palpal trochanter; distinct ventral process on proximal article of chelicera (
<figureCitation id="13372A0E7538FF94FBE55932D9A12833" box="[1141,1229,1876,1902]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="7" pageNumber="262">Fig. 18</figureCitation>
); tarsus of leg II relatively longer (
<figureCitation id="13372A0E7537FF9BFDAE5F7ADFF92E6B" box="[574,661,284,310]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="8" pageNumber="263">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BB3368B7537FF9BFEA85F25DE332DC4" blockId="8.[264,1323,284,665]" pageId="8" pageNumber="263">
2. female (collection of C. Deeleman­Reinhold; mentioned in
<bibRefCitation id="EF9D4B7A7537FF9BFB0B5F25DC972ED8" author="Martens" pageId="8" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: 548):
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, Central Kalimantan Province, Kaharian (
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,
<geoCoordinate id="EE38504C7537FF9BFE985FF4DC162EF1" box="[264,378,402,428]" direction="east" orientation="longitude" pageId="8" pageNumber="263" precision="925" value="113.666664">113°40E</geoCoordinate>
), collected from the leaf litter of a swampy primary forest on
<date id="FFB2104B7537FF9BFBD95FF4D8472EF1" box="[1097,1323,402,428]" pageId="8" pageNumber="263" value="1985-10-02" valueMax="1985-10-16" valueMin="1985-10-02">216 October 1985</date>
by S. Djojosudharmo. Different from males of
<taxonomicName id="4C0C4D087537FF9BFCBC5FDCDEB02E8E" box="[812,988,442,467]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA997537FF9BFCBC5FDCDEB02E8E" box="[812,988,442,467]" italics="true" pageId="8" pageNumber="263">M. tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997537FF9BFC735FDFD9492E8E" bold="true" box="[995,1061,441,467]" pageId="8" pageNumber="263">
<taxonomicNameLabel id="A24B57E27537FF9BFC735FDFD9492E8E" box="[995,1061,441,467]" pageId="8" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by: Body much larger (
<quantity id="4CF49B6E7537FF9BFE805F87DCED2EA6" box="[272,385,481,507]" metricMagnitude="-3" metricUnit="m" metricValue="5.19" pageId="8" pageNumber="263" unit="mm" value="5.19">5.19 mm</quantity>
long); eye tubercle broadly rounded; carapace­opisthosoma bridge present, undivided (
<figureCitation id="13372A0E7537FF9BFE1D5C6FDC882D7E" box="[397,484,521,547]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="8" pageNumber="263">Fig. 20</figureCitation>
); legs more slender and papal tibia thicker; process on palpal trochanter large, bifid (
<figureCitation id="13372A0E7537FF9BFE0E5C56DC952D17" box="[414,505,560,586]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="8" pageNumber="263">Fig. 20</figureCitation>
); chelicera more robust, with distad­directed dorsodistal process on proximal article and ventrodistal process on second article (
<figureCitation id="13372A0E7537FF9BFC575C3ED9732D2F" box="[967,1055,600,626]" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="8" pageNumber="263">Fig. 21</figureCitation>
). Tarsus of leg II (
<figureCitation id="13372A0E7537FF9BFB695C3EDC4B2DC4" captionStart="FIGURES 18 22" captionStartId="7.[264,382,1515,1539]" captionTargetBox="[271,1312,809,1489]" captionTargetId="figure@7.[264,1323,798,1500]" captionTargetPageId="7" captionText="FIGURES 18 22.? Martensiellus n. g. spp., female from Mulu N. P. (18, 19), female from Kaharian (20 22). 18. Anterior part of body and proximal part of palpus and of chelicera, lateral view. 19, 22. Distal part of leg II, lateral view. 20. Anterior part of body and proximal part of palpus, lateral view; 21. Chelicera, retrolateral view. Scale bars: 1 mm (18 19, 20 22 same scale)." httpUri="https://zenodo.org/record/174080/files/figure.png" pageId="8" pageNumber="263">Fig. 22</figureCitation>
) similar to that of
<taxonomicName id="4C0C4D087537FF9BFD905C19DFDD2DC5" box="[512,689,639,664]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA997537FF9BFD905C19DFDD2DC5" box="[512,689,639,664]" italics="true" pageId="8" pageNumber="263">M. tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997537FF9BFD275C19DF962DC4" bold="true" box="[695,762,639,665]" pageId="8" pageNumber="263">
<taxonomicNameLabel id="A24B57E27537FF9BFD275C19DF962DC4" box="[695,762,639,665]" pageId="8" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13372A0E7537FF9BFC995C19DE3D2DC4" box="[777,849,639,665]" captionStart="FIGURES 4 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 10. Martensiellus tenuipalpus n. g., n. sp., male (4 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="8" pageNumber="263">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>