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<document id="CA469B950BD5F7F075902D53DD2AFBF4" ID-DOI="10.1649/0010-065x-69.3.514" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106155" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175800600" checkinUser="diego" docAuthor="Chatzimanolis, Stylianos" docDate="2015" docId="03A58798FFAE391D506FFD63FCF7C181" docLanguage="en" docName="0010-065X-69.3.514.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.514" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Triacrus Nordmann. While I 1837" docType="treatment" docVersion="4" lastPageNumber="515" masterDocId="FF9CFFE0FFAF391F5174FFF3FFDFC548" masterDocTitle="New Records, Redescription, and Notes on Nomenclature forTriacrusNordmann (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="520" masterPageNumber="514" pageNumber="515" updateTime="1699636727904" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="88C55440FA97FB55A57FA13B17AB2E3D">New Records, Redescription, and Notes on Nomenclature forTriacrusNordmann (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:namePart id="B0D2ECCB4C20236D71AFB0FC810DA464">Chatzimanolis, Stylianos</mods:namePart>
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<mods:title id="90DC742AD419511B0EB2A0BD63BF2FC3">The Coleopterists Bulletin</mods:title>
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<mods:date id="9667B0936C8F03CAA70DAC629B2B8ADB">2015</mods:date>
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<mods:number id="54B4E2DE174C6671FD1F33D81991C3FB">2015-09-30</mods:number>
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<mods:number id="3865CDA76E40D3694D1E3D00BD20482B">69</mods:number>
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<mods:number id="0438015DEC51C83FE1A0AC10ED89FA01">3</mods:number>
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<mods:classification id="512B26FC0FA402597E207E662A0E78CC">journal article</mods:classification>
<mods:identifier id="2B9DFC69E18301506457726E4EFC0EA7" type="DOI">10.1649/0010-065x-69.3.514</mods:identifier>
<mods:identifier id="9D0CFB9B3F2A91AC03A36CF464E8E52C" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A58798FFAE391D506FFD63FCF7C181" ID-DOI="http://doi.org/10.5281/zenodo.5413346" ID-Zenodo-Dep="5413346" LSID="urn:lsid:plazi:treatment:03A58798FFAE391D506FFD63FCF7C181" httpUri="http://treatment.plazi.org/id/03A58798FFAE391D506FFD63FCF7C181" lastPageId="2" lastPageNumber="515" pageId="1" pageNumber="515">
<subSubSection id="C3166505FFAE391E506FFD63FDFAC7ED" box="[283,549,656,677]" pageId="1" pageNumber="515" type="nomenclature">
<paragraph id="8BB3368EFFAE391E506FFD63FDFAC7ED" blockId="1.[283,549,656,677]" box="[283,549,656,677]" pageId="1" pageNumber="515">
<heading id="D0FB81E2FFAE391E506FFD63FDFAC7ED" box="[283,549,656,677]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="515" reason="2">
<taxonomicName id="4C0C4D0DFFAE391E506FFD63FDFAC7ED" ID-CoL="8XQGD" authority="Nordmann, 1837" authorityName="Nordmann. While I" authorityYear="1837" box="[283,549,656,677]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E506FFD63FDFAC7ED" bold="true" box="[283,549,656,677]" pageId="1" pageNumber="515">
<emphasis id="B978EA9CFFAE391E506FFD63FEB1C7ED" bold="true" box="[283,366,656,677]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
<bibRefCitation id="EF9D4B7FFFAE391E5002FD63FDFAC7ED" author="Nordmann" box="[374,549,656,677]" pageId="1" pageNumber="515" pagination="1 - 167" refId="ref4561" refString="Nordmann, A. von. 1837. Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum 4: 1 - 167." type="journal article" year="1837">Nordmann, 1837</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3166505FFAE391E51B1FD48FD5EC61D" pageId="1" pageNumber="515" type="type_taxon">
<paragraph id="8BB3368EFFAE391E51B1FD48FD5EC61D" blockId="1.[173,659,699,1729]" pageId="1" pageNumber="515">
<emphasis id="B978EA9CFFAE391E51B1FD48FE92C798" bold="true" box="[197,333,699,720]" pageId="1" pageNumber="515">
<typeStatus id="54B7882CFFAE391E51B1FD48FF2AC798" box="[197,245,699,720]" pageId="1" pageNumber="515">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C0C4D0DFFAE391E502FFD48FDB2C798" ID-CoL="584V6" authority="Nordmann" authorityName="Nordmann" authorityYear="1837" box="[347,621,699,720]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="species" species="dilatus">
<emphasis id="B978EA9CFFAE391E502FFD48FE25C798" box="[347,506,699,720]" italics="true" pageId="1" pageNumber="515">Triacrus dilatus</emphasis>
Nordmann
</taxonomicName>
, by monotypy (but see “Nomenclature” below in the species treatment).
<bibRefCitation id="EF9D4B7FFFAE391E5004FD03FDFDC64D" author="Nordmann" box="[368,546,752,773]" pageId="1" pageNumber="515" pagination="1 - 167" refId="ref4561" refString="Nordmann, A. von. 1837. Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum 4: 1 - 167." type="journal article" year="1837">Nordmann (1837)</bibRefCitation>
mentioned that the name
<taxonomicName id="4C0C4D0DFFAE391E504DFCF8FE54C668" ID-CoL="8XQGD" authorityName="Nordmann. While I" authorityYear="1837" box="[313,395,779,800]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E504DFCF8FE54C668" box="[313,395,779,800]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
</taxonomicName>
means “three appendages” in Greek and can possibly refer to the structure of the left mandible (
<figureCitation id="13372A0BFFAE391E500FFCB3FE68C61D" box="[379,439,832,853]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="1" pageNumber="515">Fig. 2</figureCitation>
) or abdominal apex.
</paragraph>
</subSubSection>
<subSubSection id="C3166505FFAE391E51B1FCA9FEF8C020" pageId="1" pageNumber="515" type="diagnosis">
<paragraph id="8BB3368EFFAE391E51B1FCA9FEF8C020" blockId="1.[173,659,699,1729]" pageId="1" pageNumber="515">
<emphasis id="B978EA9CFFAE391E51B1FCA9FEF2C627" bold="true" box="[197,301,858,879]" pageId="1" pageNumber="515">Diagnosis.</emphasis>
Due to its large size, within
<taxonomicName id="4C0C4D0DFFAE391E5331FCA9FF31C6C2" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
<taxonomicName id="4C0C4D0DFFAE391E5183FC86FE96C6C2" authorityName="Nordmann. While I" authorityYear="1837" box="[247,329,885,906]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E5183FC86FE96C6C2" box="[247,329,885,906]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
</taxonomicName>
can only be confused with other similarly large genera such as
<taxonomicName id="4C0C4D0DFFAE391E536EFC63FEE3C6F7" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Darwinilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E536EFC63FD4CC6ED" box="[538,659,912,933]" italics="true" pageId="1" pageNumber="515">Darwinilus</emphasis>
Chatzimanolis
</taxonomicName>
or
<taxonomicName id="4C0C4D0DFFAE391E501FFC59FDC0C6F7" box="[363,543,938,959]" class="Insecta" family="Staphylinidae" genus="Trigonopselaphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E501FFC59FDC0C6F7" box="[363,543,938,959]" italics="true" pageId="1" pageNumber="515">Trigonopselaphus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C0C4D0DFFAE391E51D9FC36FEDDC692" authorityName="Nordmann. While I" authorityYear="1837" box="[173,258,965,986]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E51D9FC36FEDDC692" box="[173,258,965,986]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
</taxonomicName>
can be easily distinguished from these two genera by the shape of the head (rounded rectangle in
<taxonomicName id="4C0C4D0DFFAE391E5056FC09FEADC147" authorityName="Nordmann. While I" authorityYear="1837" box="[290,370,1018,1039]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E5056FC09FEADC147" box="[290,370,1018,1039]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
</taxonomicName>
; hexagonal in
<taxonomicName id="4C0C4D0DFFAE391E5375FC09FDB5C147" authorityName="Chatzimanolis" authorityYear="2014" box="[513,618,1018,1039]" class="Insecta" family="Staphylinidae" genus="Darwinilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E5375FC09FDB5C147" box="[513,618,1018,1039]" italics="true" pageId="1" pageNumber="515">Darwinilus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D0DFFAE391E51D9FBE7FEB6C161" box="[173,361,1044,1065]" class="Insecta" family="Staphylinidae" genus="Trigonopselaphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E51D9FBE7FEB6C161" box="[173,361,1044,1065]" italics="true" pageId="1" pageNumber="515">Trigonopselaphus</emphasis>
</taxonomicName>
), the shape of the antennae (antennomeres 6–11 emarginate in
<taxonomicName id="4C0C4D0DFFAE391E5364FBDCFDBAC10C" authorityName="Nordmann. While I" authorityYear="1837" box="[528,613,1071,1092]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E5364FBDCFDBAC10C" box="[528,613,1071,1092]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
</taxonomicName>
, not emarginate in
<taxonomicName id="4C0C4D0DFFAE391E504DFBB9FE7BC117" authorityName="Chatzimanolis" authorityYear="2014" box="[313,420,1098,1119]" class="Insecta" family="Staphylinidae" genus="Darwinilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E504DFBB9FE7BC117" box="[313,420,1098,1119]" italics="true" pageId="1" pageNumber="515">Darwinilus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D0DFFAE391E50ACFBB9FD5AC117" box="[472,645,1098,1119]" class="Insecta" family="Staphylinidae" genus="Trigonopselaphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E50ACFBB9FD5AC117" box="[472,645,1098,1119]" italics="true" pageId="1" pageNumber="515">Trigonopselaphus</emphasis>
</taxonomicName>
), and the shape of the pronotum (in
<taxonomicName id="4C0C4D0DFFAE391E5340FB97FD53C131" authorityName="Nordmann. While I" authorityYear="1837" box="[564,652,1124,1145]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E5340FB97FD53C131" box="[564,652,1124,1145]" italics="true" pageId="1" pageNumber="515">Triacrus</emphasis>
</taxonomicName>
, pronotum expanding from anterolateral corners to almost middle, then converging towards posterolateral corners and becoming almost explanate, with posterolateral corners expanded and broadly rounded; in
<taxonomicName id="4C0C4D0DFFAE391E505DFB1AFE4BC1B6" authorityName="Chatzimanolis" authorityYear="2014" box="[297,404,1257,1278]" class="Insecta" family="Staphylinidae" genus="Darwinilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E505DFB1AFE4BC1B6" box="[297,404,1257,1278]" italics="true" pageId="1" pageNumber="515">Darwinilus</emphasis>
</taxonomicName>
, posterolateral corners of pronotum not expanded and pronotum not explanate; in
<taxonomicName id="4C0C4D0DFFAE391E51BDFAEDFEACC07B" box="[201,371,1310,1331]" class="Insecta" family="Staphylinidae" genus="Trigonopselaphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA9CFFAE391E51BDFAEDFEACC07B" box="[201,371,1310,1331]" italics="true" pageId="1" pageNumber="515">Trigonopselaphus</emphasis>
</taxonomicName>
, pronotum with anterolateral corners bulging and clearly extending beyond anterior margin).
</paragraph>
</subSubSection>
<caption id="DF736606FFAE391E53AFFBBAFCC4C1C4" ID-DOI="http://doi.org/10.5281/zenodo.10106157" ID-Zenodo-Dep="10106157" httpUri="https://zenodo.org/record/10106157/files/figure.png" pageId="1" pageNumber="515" startId="1.[731,763,1097,1116]" targetBox="[708,1193,195,1073]" targetPageId="1">
<paragraph id="8BB3368EFFAE391E53AFFBBAFCC4C1C4" blockId="1.[707,1193,1097,1164]" pageId="1" pageNumber="515">
<emphasis id="B978EA9CFFAE391E53AFFBBAFCC8C114" bold="true" box="[731,791,1097,1116]" pageId="1" pageNumber="515">Fig. 1.</emphasis>
Habitus of
<taxonomicName id="4C0C4D0DFFAE391E52E5FBBAFBC1C113" authorityName="Nordmann" authorityYear="1837" box="[913,1054,1097,1115]" class="Insecta" family="Staphylinidae" genus="Triacrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="515" phylum="Arthropoda" rank="species" species="dilatus">
<emphasis id="B978EA9CFFAE391E52E5FBBAFBC1C113" box="[913,1054,1097,1115]" italics="true" pageId="1" pageNumber="515">Triacrus dilatus</emphasis>
</taxonomicName>
. Total length = 34 mm. Image copyright of the Natural History Museum (London).
</paragraph>
</caption>
<subSubSection id="C3166505FFAE391D51B1FA9DFCF7C181" lastPageId="2" lastPageNumber="516" pageId="1" pageNumber="515" type="description">
<paragraph id="8BB3368EFFAE391D51B1FA9DFCF7C181" blockId="1.[173,659,699,1729]" lastBlockId="2.[685,1172,964,1225]" lastPageId="2" lastPageNumber="516" pageId="1" pageNumber="515">
<emphasis id="B978EA9CFFAE391E51B1FA9DFE9EC0CB" bold="true" box="[197,321,1390,1411]" pageId="1" pageNumber="515">Description.</emphasis>
Size large, body robust, habitus as in
<figureCitation id="13372A0BFFAE391E51BDFA7BFED8C0D5" box="[201,263,1416,1437]" captionStart="Fig" captionStartId="1.[731,763,1097,1116]" captionTargetBox="[708,1193,195,1073]" captionTargetId="figure-492@1.[707,1193,194,1074]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Triacrus dilatus. Total length = 34 mm. Image copyright of the Natural History Museum (London)." figureDoi="http://doi.org/10.5281/zenodo.10106157" httpUri="https://zenodo.org/record/10106157/files/figure.png" pageId="1" pageNumber="515">Fig. 1</figureCitation>
. Head transverse, rounded rectangular, head size variable both in males and females; with microsculpture and small punctures. Clypeus not emarginate; anteclypeus expanded, weakly sclerotized. Eyes small to medium, positioned anteriorly in oblique angle to dorsal surface. Postoccipital suture and ventral basal ridge present; presence of infraorbital ridge not clear but ridge situated between postmandibular ridge and gular suture extends from posterior to middle part of gena; postmandibular ridge present and prominent; gular sutures converging medioposteriorly; with nuchal ridge and well-defined neck. Epicranium with large, prominent macrosetae around anterior and lateral margins. Antennae long, subserrate, with 11 antennomeres; antennomeres 1–4 with several rows of macrosetae; antennomeres 5–11 covered with microtrichiae; antennomeres 6–11 with small lateral emargination (visible in
<figureCitation id="13372A0BFFAE391E52D3FA9EFBDFC0CA" box="[935,1024,1389,1411]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="1" pageNumber="515">Figs. 2–3</figureCitation>
); antennomere 11 with 2 lobes protruding from base of antennomere (visible in
<figureCitation id="13372A0BFFAE391E5251FA50FCA3C0F0" box="[805,892,1442,1464]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="1" pageNumber="515">Figs. 2–3</figureCitation>
). Mouthparts with labrum medially emarginate; mandibles (
<figureCitation id="13372A0BFFAE391E5294FA4EFBE0C09A" box="[992,1087,1469,1491]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="1" pageNumber="515">Figs. 2–3</figureCitation>
) elongate, variable in size; mandibles with prominent fold extending from molar region to base; left mandible with 2 teeth, 1 with large, single molar extending laterally and smaller bicuspid molar posteriorly; right mandible with 1 bicuspid molar; with setose prostheca. Maxilla as in
<figureCitation id="13372A0BFFAE391E52CDF9AEFC2AC33A" box="[953,1013,1629,1650]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="1" pageNumber="515">Fig. 3</figureCitation>
; galea and lacinia densely setose; maxillary palpi 4-segmented; P
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short; P
<subScript id="178834CBFFAE391E5279F968FCC9C3E1" attach="left" box="[781,790,1691,1705]" fontSize="6" pageId="1" pageNumber="515">2</subScript>
curved, approximately 3 times as long as P
<subScript id="178834CBFFAE391E53A4F946FD06C38B" attach="left" box="[720,729,1717,1731]" fontSize="6" pageId="1" pageNumber="515">1</subScript>
; P
<subScript id="178834CBFFAE391E5380F946FD22C38B" attach="left" box="[756,765,1717,1731]" fontSize="6" pageId="1" pageNumber="515">3</subScript>
curved, slightly shorter than P
<subScript id="178834CBFFAE391E555CF946FBEEC38B" attach="left" box="[1064,1073,1717,1731]" fontSize="6" pageId="1" pageNumber="515">2</subScript>
; P
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straight, rounded apically, longer than P
<subScript id="178834CBFFAD391D50ACFC3DFE3EC694" attach="left" box="[472,481,974,988]" fontSize="6" pageId="2" pageNumber="516">3</subScript>
, as long as P
<subScript id="178834CBFFAD391D531AFC3DFDA8C694" attach="left" box="[622,631,974,988]" fontSize="6" pageId="2" pageNumber="516">2</subScript>
. Labium as in
<figureCitation id="13372A0BFFAD391D505DFC2CFEB6C6BC" box="[297,361,991,1012]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="2" pageNumber="516">Fig. 3</figureCitation>
; labial palpi 3-segmented; P
<subScript id="178834CBFFAD391D51D1FBF0FF71C159" attach="left" box="[165,174,1027,1041]" fontSize="6" pageId="2" pageNumber="516">1</subScript>
as long as P
<subScript id="178834CBFFAD391D5044FBF0FEE6C159" attach="left" box="[304,313,1027,1041]" fontSize="6" pageId="2" pageNumber="516">2</subScript>
; P
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slightly curved; P
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truncate, 1.5 times as long as P
<subScript id="178834CBFFAD391D500FFBEEFE5BC163" attach="right" box="[379,388,1053,1067]" fontSize="6" pageId="2" pageNumber="516">2</subScript>
. Pronotum (
<figureCitation id="13372A0BFFAD391D5371FBE7FD9CC161" box="[517,579,1044,1065]" captionStart="Figs" captionStartId="2.[175,215,884,903]" captionTargetBox="[155,1170,200,862]" captionTargetId="figure-374@2.[151,1171,194,862]" captionTargetPageId="2" captionText="Figs. 2–3. Triacrus dilatus. 2) Head and thorax, dorsal view; 3) Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106159" httpUri="https://zenodo.org/record/10106159/files/figure.png" pageId="2" pageNumber="516">Fig. 2</figureCitation>
) subquadrate to transverse; slightly wider than head; pronotum expanding from anterolateral corners to almost middle, then converging towards posterolateral corners, becoming almost explanate; posterolateral corners expanded and broadly rounded; with microsculpture and small punctures. Pronotal hypomeron expanded; superior and inferior marginal lines of hypomeron separate throughout their length and superior line fully visible from above; without postcoxal process. Basisternum with microsculpture, small punctures, and macrosetae; anterior marginal depression present; sternacostal ridge present; furcasternum with large, medial, elongate carina pointed vertically. Elytra wider and longer than pronotum; with large, prominent macrosetae around lateral and posterior margins. Elytral sutures slightly elevated; elytra becoming depressed near mesoscutellum. Mesoscutellum large, triangular, with dense punctation and setation similar to elytra. Hinds wings fully developed. Mesoventrite without median carina, with small mesoventral process; metaventrite disc with narrow, elongate, impunctate line; metaventrite with 2 small, round, triangular mesoventral processes. Leg with tarsal segmentation 5-5-5; covered with dense setae; protarsus enlarged in both sexes; meso- and metatarsi not enlarged but elongate; 2 empodial setae present. Abdomen with paired protergal glands present; abdominal tergites III–V with tergal basal carina and curved (arch-like) carina. Tergites and sternites with microsculpture. Abdominal sternite VII in males with porose structure (
<figureCitation id="13372A0BFFAD391D5228FB97FC43C131" box="[860,924,1124,1145]" captionStart="Fig" captionStartId="3.[197,229,915,934]" captionTargetBox="[184,649,194,886]" captionTargetId="figure-457@3.[173,659,194,892]" captionTargetPageId="3" captionText="Fig. 4. Triacrus dilatus, abdominal sternites VII–IX." figureDoi="http://doi.org/10.5281/zenodo.10106163" httpUri="https://zenodo.org/record/10106163/files/figure.png" pageId="2" pageNumber="516">Fig. 4</figureCitation>
). Male and female genitalia typical of
<taxonomicName id="4C0C4D0DFFAD391D5226FB8DFC3FC1DB" authorityName="Sharp" authorityYear="1884" box="[850,992,1150,1171]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="516" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
; spermatheca not sclerotized; aedeagus with long parameres and median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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