treatments-xml/data/03/98/87/039887D2FFB7AF482A8A3813FB9EFC3F.xml

<document id="6915CE508CD66FC3C2BC68575531116B" ID-DOI="10.11646/zootaxa.4819.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4396744" ID-ZooBank="80258C53-F94D-46CB-9E86-AF0C3226DF9E" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1609199984737" checkinUser="plazi" docAuthor="Polizei, Thiago T. S., Barclay, Maxwell V. L. &amp; Bispo, Pitágoras C." docDate="2020" docId="039887D2FFB7AF482A8A3813FB9EFC3F" docLanguage="en" docName="zootaxa.4819.1.2.pdf" docOrigin="Zootaxa 4819 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexacylloepus thoracica Polizei &amp; Barclay &amp; Bispo 2020, sp. nov." docType="treatment" docVersion="1" lastPageNumber="40" masterDocId="FFA1FFAAFF97AF6A2A1D3A09FFC5FF9A" masterDocTitle="Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)" masterLastPageNumber="48" masterPageNumber="6" pageNumber="38" updateTime="1730286293773" updateUser="diego" zenodo-license-document="CLOSED">
<mods:mods id="672CFDAE302C27350DF6E9DB4E6E7E62" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C778C93EDD78324E6A687ACFD6DD2E7F">
<mods:title id="8B6870D783B01B0BFE8A2E91E3F5EA5A">Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)</mods:title>
</mods:titleInfo>
<mods:name id="D459C6101D0F3E06C31D762677C4FD78" type="personal">
<mods:role id="1864F9FB9F7EDB4A79F82572E678839D">
<mods:roleTerm id="AFA760F88683A4C96361F6CCB9699A40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB03C9CE37F1DE7469A67C6828507946">Polizei, Thiago T. S.</mods:namePart>
<mods:nameIdentifier id="3C96ABA3CD8CD45605C53920BEB1EA7A" type="ORCID">0000-0003-3746-8865</mods:nameIdentifier>
<mods:affiliation id="F98E9C7DFF5AE33CD36E6C98C8A4537B">Laboratório de Coleoptera (LaC) - Museu de Zoologia da Universidade de São Paulo (MZSP). São Paulo, São Paulo State, Brazil. thiagopolizei @ hotmail. com; https: // orcid. org / 0000 - 0003 - 3746 - 8865 &amp; Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil.</mods:affiliation>
<mods:nameIdentifier id="E4073860036C90BAA5ACF8C58F818A05" type="email">thiagopolizei@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7B14DF43393C37AFF0F4A94D4429F6D2" type="personal">
<mods:role id="062976625FFE9E7E8A790F5BB1E21F83">
<mods:roleTerm id="F083A7A5E87064A90FF216C279FD8C3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5049F032AC1B9596C63B739FA5CA21E6">Barclay, Maxwell V. L.</mods:namePart>
<mods:affiliation id="A4762D51F509D685607D09CFD18B7667">Department of Life Sciences, Natural History Museum. London SW 7 5 BD, United Kingdom.</mods:affiliation>
</mods:name>
<mods:name id="6BDA3D3D4A4F961EECC679E6106657FF" type="personal">
<mods:role id="76B3B79AE7175C171637117EEE8BBDC7">
<mods:roleTerm id="BBAFB7881AE3D7D008407708A40B9B12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08612FA8134194EB817B8B81A0179BF5">Bispo, Pitágoras C.</mods:namePart>
<mods:affiliation id="1C6A4687B83D4D43102A26A611AAA56A">Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil. &amp; Laboratório de Biologia Aquática (LABIA). Departamento de Ciências Biológicas - Faculdade de Ciências e Letras de Assis (FCL) - Universidade Estadual Paulista “ Júlio de Mesquita Filho ” (Unesp). Assis, São Paulo State, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9DEFC8385626A7C205768490F7CB0FB4">text</mods:typeOfResource>
<mods:relatedItem id="DD1AD01BC892D3F167D375D790E7F5FD" type="host">
<mods:titleInfo id="8BB26C12BBA0DA58103FA5A2B604A26A">
<mods:title id="651820B061EFA9F72943933BCBB71201">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4234E415AECFE36E79D62F18782AC826">
<mods:date id="3C0E5B84BD9022A7419D656F3A9AAD98">2020</mods:date>
<mods:detail id="4F37E5F190041013EF2142BD56148322" type="pubDate">
<mods:number id="6932440AB794D2D0A3F1C8E33E678473">2020-07-22</mods:number>
</mods:detail>
<mods:detail id="E8737C7FCD77953B4DD6FDB3DB7E33CD" type="volume">
<mods:number id="5EC0F17ED7F9ADD71783361D84ACEB7E">4819</mods:number>
</mods:detail>
<mods:detail id="D4FA7AC1DBD2A68A166323478A91BE59" type="issue">
<mods:number id="E698BD8DE1FACC876686B15683A26493">1</mods:number>
</mods:detail>
<mods:extent id="CA5FEA82AB932B7658BA599D54836ECE" unit="page">
<mods:start id="C5207F2C26B5D6B0F24E24801AB3BD29">6</mods:start>
<mods:end id="9287A79E78452B9EC122394AB546449D">48</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="074A6293182B4886F1EF84AFA499810D">journal article</mods:classification>
<mods:identifier id="8F3398E90E814B6BA9F0B0D15F508EBC" type="DOI">10.11646/zootaxa.4819.1.2</mods:identifier>
<mods:identifier id="30E0EF56D9940CBEE05863E80808CA21" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="703A37C1E5D2F1F8573B7CFEA4760142" type="Zenodo-Dep">4396744</mods:identifier>
<mods:identifier id="BFF10DA3CA03247BDEDCD04C93FBB614" type="ZooBank">80258C53-F94D-46CB-9E86-AF0C3226DF9E</mods:identifier>
</mods:mods>
<treatment id="039887D2FFB7AF482A8A3813FB9EFC3F" LSID="urn:lsid:plazi:treatment:039887D2FFB7AF482A8A3813FB9EFC3F" httpUri="http://treatment.plazi.org/id/039887D2FFB7AF482A8A3813FB9EFC3F" lastPageId="34" lastPageNumber="40" pageId="32" pageNumber="38">
<subSubSection id="C32B654FFFB7AF4A2A8A3813FDE4FDAE" box="[151,545,538,564]" pageId="32" pageNumber="38" type="nomenclature">
<paragraph id="8B8E36C4FFB7AF4A2A8A3813FDE4FDAE" blockId="32.[151,545,538,600]" box="[151,545,538,564]" pageId="32" pageNumber="38">
<taxonomicName id="4C314D47FFB7AF4A2A8A3813FE7EFDAE" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[151,443,538,564]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="38" phylum="Arthropoda" rank="species" species="thoracica" status="sp. nov.">
<emphasis id="B945EAD6FFB7AF4A2A8A3813FE7EFDAE" box="[151,443,538,564]" italics="true" pageId="32" pageNumber="38">Hexacylloepus thoracica</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FFB7AF4A2BDE3813FDE4FDAE" bold="true" box="[451,545,538,564]" pageId="32" pageNumber="38">
<taxonomicNameLabel id="A27657ADFFB7AF4A2BDE3813FDE4FDAE" box="[451,545,538,564]" pageId="32" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB7AF4A2A8A3834FEF1FDCD" box="[151,308,573,600]" pageId="32" pageNumber="38" type="description">
<paragraph id="8B8E36C4FFB7AF4A2A8A3834FEF1FDCD" blockId="32.[151,545,538,600]" box="[151,308,573,600]" pageId="32" pageNumber="38">
(
<figureCitation id="130A2A41FFB7AF4A2ABD3834FF35FDC2" box="[160,240,573,600]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB7AF4A2A8A388DFDB8FD07" box="[151,637,643,670]" pageId="32" pageNumber="38" type="materials_examined">
<paragraph id="8B8E36C4FFB7AF4A2A8A388DFDB8FD07" blockId="32.[151,1437,643,2008]" box="[151,637,643,670]" pageId="32" pageNumber="38">
<materialsCitation id="3B593C99FFB7AF4A2A8A388DFDBCFD07" box="[151,633,643,670]" country="Ecuador" location="Orellana Province" pageId="32" pageNumber="38" specimenCount="1" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="B945EAD6FFB7AF4A2A8A388DFEFCFD07" bold="true" box="[151,313,643,670]" pageId="32" pageNumber="38">
<typeStatus id="548A8866FFB7AF4A2A8A388DFF17FD04" box="[151,210,644,670]" pageId="32" pageNumber="38">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3267654FFB7AF4A2B5D388AFE5AFD07" box="[320,415,643,669]" name="Ecuador" pageId="32" pageNumber="38">Ecuador</collectingCountry>
, 
<collectingRegion id="49F5F826FFB7AF4A2BB7388AFDBCFD07" box="[426,633,643,669]" country="Ecuador" name="Orellana" pageId="32" pageNumber="38">Orellana Province</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB7AF4A2ADA38AFFDBDFC22" pageId="32" pageNumber="38" type="diagnosis">
<paragraph id="8B8E36C4FFB7AF4A2ADA38AFFDBDFC22" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
<emphasis id="B945EAD6FFB7AF4A2ADA38AFFEFFFD5A" bold="true" box="[199,314,678,704]" pageId="32" pageNumber="38">Diagnosis</emphasis>
: Antennae without setae. Frontoclypeal suture inconspicuous. Pronotum with a long and deep longitudinal impression on disc, reaching the anterior and posterior margins. Hypomera with a narrow belt of tomentum reaching the anterior and lateral margins. Epipleura without tomentum. Metaventrite with a longitudinal medial depression, very strongly raised towards the mesoventrite, forming a tubercle. Tibiae each with a row of setae on distal half; meso- and metatibiae each with a row of spines on distal half. Abdomen with the disc of the first ventrite broad and without tomentum. Male genitalia elongated; phallobase twice as long as wide, length similar to the penis; parameres reaching 2/3 of the penis length, 2x wider than the penis; penis elongated with anterior half sinuous and posterior half straight, apex strongly acute.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB7AF482ADA39C9FE60FEDB" lastPageId="34" lastPageNumber="40" pageId="32" pageNumber="38" type="description">
<paragraph id="8B8E36C4FFB7AF4A2ADA39C9FCD4FC64" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
<emphasis id="B945EAD6FFB7AF4A2ADA39C9FE9EFC40" bold="true" box="[199,347,960,986]" pageId="32" pageNumber="38">Description:</emphasis>
Male. Body elongated, subparallel. Brown colour, with the antenna, mouthparts and legs reddishbrown. Total length: 
<quantity id="4CC99B21FFB7AF4A2B9C39EDFE2DFC64" box="[385,488,996,1022]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="32" pageNumber="38" unit="mm" value="1.81">1.81 mm</quantity>
. Greatest width: 
<quantity id="4CC99B21FFB7AF4A28BB39EDFCC8FC64" box="[678,781,996,1022]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="32" pageNumber="38" unit="mm" value="0.78">0.78 mm</quantity>
.
</paragraph>
<paragraph id="8B8E36C4FFB7AF4A2ADA3E0EFC8FFB4B" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
<emphasis id="B945EAD6FFB7AF4A2ADA3E0EFEC8FBBB" box="[199,269,1031,1057]" italics="true" pageId="32" pageNumber="38">Head:</emphasis>
surface with micropunctures separated from each other by their diameter. Antenna inserted on the anterior margin of the eyes, filiform, with 11 antennomeres each one with long thin setae at apex. Frontoclypeal suture inconspicuous. Clypeus 4x as wide as long; anterior margin concave and posterior sinuous; surface similar to that of head. Labrum twice as wide as long; antero-lateral angles rounded and alutaceous; surface without setae. Maxillary palpus 4-segmented; terminal palpomere truncated apically. Labial palpus 3-segmented. Mentum and submentum rectangular. Gula rectangular. Genae covered with tomentum.
</paragraph>
<paragraph id="8B8E36C4FFB7AF4A2ADA3ED3FED0FAC4" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
<emphasis id="B945EAD6FFB7AF4A2ADA3ED3FEE4FB6E" box="[199,289,1242,1268]" italics="true" pageId="32" pageNumber="38">Thorax:</emphasis>
Pronotum as long as wide, arcuate anteriorly; sides sinuate and serrate, posterior margin smooth with two prescutellar foveae (
<figureCitation id="130A2A41FFB7AF4A2BAF3EF4FDD9FA82" box="[434,540,1277,1304]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15A</figureCitation>
). Surface with micropunctures and setae. Sublateral carinae complete, sinuous and raised (
<figureCitation id="130A2A41FFB7AF4A2B063F29FE40FAA1" box="[283,389,1312,1339]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15A</figureCitation>
). Longitudinal impression long and deep on disc, reaching the anterior and posterior margin (
<figureCitation id="130A2A41FFB7AF4A2ABD3F4AFECDFAC4" box="[160,264,1347,1374]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15A</figureCitation>
).
</paragraph>
<paragraph id="8B8E36C4FFB7AF4A2AEA3F6EFED0F9AB" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
Elytra 3x as long as wide; anterior margin arcuate, lateral margin moderately arcuate and serrate; raised apex (
<figureCitation id="130A2A41FFB7AF4A2AC33F83FE8CFA3F" box="[222,329,1418,1445]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15A</figureCitation>
). Humeri moderately prominent. Surface with puncture rows separated from each other by their diameter. Elytral suture moderately elevated. Carina composed by two rows of punctures. Carina short and elevated on interval IV, extending from base to 1/4 of the elytral length; sublateral carinae on intervals VI and VIII, extending from base to 3/4 of the elytral length. Epipleura without tomentum (
<figureCitation id="130A2A41FFB7AF4A299B3FFAFC29F994" box="[902,1004,1523,1550]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15B</figureCitation>
). Hind wings macropterous. Scutellum shield-like.
</paragraph>
<paragraph id="8B8E36C4FFB7AF4A2ADA3C33FAD5F924" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
Prosternum narrow, strongly elevated, surface with micropunctures; anterior margin straight and sides covered with tomentum. Prosternal process long, narrow, and beyond the procoxae; lateral margin, concave and elevated arrow-like apex. Notosternal suture sinuous. Hypomera narrow, wider posteriorly than anteriorly; surface with micropunctures and with a narrow belt of tomentum reaching the anterior and lateral margins (
<figureCitation id="130A2A41FFB7AF4A2E803CAAFAC6F924" box="[1181,1283,1699,1726]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15B</figureCitation>
).
</paragraph>
<paragraph id="8B8E36C4FFB7AF4A2ADA3CCEFB5BF8B2" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">Mesoventrite with a deep groove to receive the prosternal process. Mesepimeron and mesepisternum covered with tomentum. Metaventrite broad with a longitudinal medial depression, very strongly raised towards the mesoventrite and forming a tubercle; surface with micropunctures, sides covered with tomentum.</paragraph>
<paragraph id="8B8E36C4FFB7AF4A2ADA3D38FEAEF842" blockId="32.[151,1437,643,2008]" pageId="32" pageNumber="38">
Legs (except tarsi) covered with punctures and tomentum (
<figureCitation id="130A2A41FFB7AF4A29723D39FC1DF8D1" box="[879,984,1840,1867]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="32" pageNumber="38">Fig. 15B</figureCitation>
). Coxae rounded; trochanter elongated; femora elongated and thin, 8x longer than wide, 1/3 of the length of the elytra; tibiae long and thin, 10x as long as wide; with the half of the elytra length, with a row of setae on distal half. Meso- and metatibiae each with a row of spines on distal half. Tarsi elongate, with setae; apical tarsomere with the length of the basal four combined. Claws very long and thin.
</paragraph>
<caption id="DF4E664CFFB6AF4B2A8A3CBFFCF0F968" ID-DOI="http://doi.org/10.5281/zenodo.4396776" ID-Zenodo-Dep="4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="33" pageNumber="39" startId="33.[151,250,1718,1742]" targetBox="[189,1377,196,1681]" targetPageId="33">
<paragraph id="8B8E36C4FFB6AF4B2A8A3CBFFCF0F968" blockId="33.[151,1436,1718,1778]" pageId="33" pageNumber="39">
<emphasis id="B945EAD6FFB6AF4B2A8A3CBFFEDAF954" bold="true" box="[151,287,1718,1742]" pageId="33" pageNumber="39">FIGURE 15.</emphasis>
<taxonomicName id="4C314D47FFB6AF4B2B383CBFFDE5F954" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[293,544,1718,1742]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="39" phylum="Arthropoda" rank="species" species="thoracica" status="sp. nov.">
<emphasis id="B945EAD6FFB6AF4B2B383CBFFDE5F954" box="[293,544,1718,1742]" italics="true" pageId="33" pageNumber="39">Hexacylloepus thoracica</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FFB6AF4B283B3CBFFDB3F954" bold="true" box="[550,630,1718,1742]" pageId="33" pageNumber="39">
<taxonomicNameLabel id="A27657ADFFB6AF4B283B3CBFFDB3F954" box="[550,630,1718,1742]" pageId="33" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm.
</paragraph>
</caption>
<paragraph id="8B8E36C4FFB6AF4B2ADA3D29FBFDF851" blockId="33.[151,1436,1824,2031]" pageId="33" pageNumber="39">
<emphasis id="B945EAD6FFB6AF4B2ADA3D29FEF4F8A0" box="[199,305,1824,1850]" italics="true" pageId="33" pageNumber="39">Abdomen</emphasis>
(
<figureCitation id="130A2A41FFB6AF4B2B5F3D29FE6EF8A1" box="[322,427,1824,1851]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="33" pageNumber="39">Fig. 15B</figureCitation>
): five ventrites; as long as wide. Anterior and posterior margins straight on the first and second ventrites and arcuate on the subsequent ventrites, lateral margins rounded. Surface densely covered with tomentum (excepted for disc of first ventrite), micropunctures and short setae. Disc of the first ventrite broad with a deep depression, and with a pair of carinae extending towards and reaching the metaventrite. Fifth ventrite with lateral margins slightly produced posterolaterad; apex tapered and with short setae.
</paragraph>
<paragraph id="8B8E36C4FFB6AF482ADA3DDDFF1FFE87" blockId="33.[151,1436,1824,2031]" lastBlockId="34.[151,1437,151,933]" lastPageId="34" lastPageNumber="40" pageId="33" pageNumber="39">
<emphasis id="B945EAD6FFB6AF4B2ADA3DDDFEAEF874" box="[199,363,2004,2030]" italics="true" pageId="33" pageNumber="39">Male genitalia</emphasis>
(
<figureCitation id="130A2A41FFB6AF4B2B643DDDFE1BF875" box="[377,478,2004,2031]" captionStart="FIGURE 15" captionStartId="33.[151,250,1718,1742]" captionTargetBox="[189,1377,196,1681]" captionTargetId="figure-24@33.[185,1402,181,1693]" captionTargetPageId="33" captionText="FIGURE 15. Hexacylloepus thoracica sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396776" httpUri="https://zenodo.org/record/4396776/files/figure.png" pageId="33" pageNumber="39">Fig. 15C</figureCitation>
): (Total length: 
<quantity id="4CC99B21FFB6AF4B28933DDDFD31F875" box="[654,756,2004,2031]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="33" pageNumber="39" unit="mm" value="0.59">0.59 mm</quantity>
. Greatest width: 
<quantity id="4CC99B21FFB6AF4B29B23DDDFBD3F875" box="[943,1046,2004,2031]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="33" pageNumber="39" unit="mm" value="0.16">0.16 mm</quantity>
), symmetrical and elongated. Phallobase twice as long as wide, length similar to the penis. Parameres articulated with the phallobase, reaching 2/3 of the penis length; 2x wider than the penis and twice as long as wide, apex slightly sharp. Penis elongated, anterior half arcuate and posterior half straight, apex strongly sharp; fibula elongated approximately the same length as the penis.
</paragraph>
<paragraph id="8B8E36C4FFB5AF482ADA3B2EFE60FEDB" blockId="34.[151,1437,151,933]" box="[199,421,295,321]" pageId="34" pageNumber="40">
<emphasis id="B945EAD6FFB5AF482ADA3B2EFEEDFEDB" bold="true" box="[199,296,295,321]" pageId="34" pageNumber="40">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB5AF482ADA3B43FDDDFEFF" box="[199,536,330,357]" pageId="34" pageNumber="40" type="materials_examined">
<paragraph id="8B8E36C4FFB5AF482ADA3B43FDDDFEFF" blockId="34.[151,1437,151,933]" box="[199,536,330,357]" pageId="34" pageNumber="40">
<emphasis id="B945EAD6FFB5AF482ADA3B43FE76FEFE" bold="true" box="[199,435,330,356]" pageId="34" pageNumber="40">Material examined:</emphasis>
<specimenCount id="9D37FD4DFFB5AF482BA73B42FDD1FEFF" box="[442,532,331,357]" pageId="34" pageNumber="40" type="male">2 males</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB5AF482ADA3B67FCA3FE13" box="[199,870,366,393]" pageId="34" pageNumber="40" type="distribution">
<paragraph id="8B8E36C4FFB5AF482ADA3B67FCA3FE13" blockId="34.[151,1437,151,933]" box="[199,870,366,393]" pageId="34" pageNumber="40">
<emphasis id="B945EAD6FFB5AF482ADA3B67FEA6FE12" bold="true" box="[199,355,366,392]" pageId="34" pageNumber="40">Distribution:</emphasis>
<collectingCountry id="F3267654FFB5AF482B773B66FE09FE13" box="[362,460,367,393]" name="Ecuador" pageId="34" pageNumber="40">Ecuador</collectingCountry>
: 
<collectingRegion id="49F5F826FFB5AF482BCB3B66FD97FE13" box="[470,594,367,393]" country="Ecuador" name="Sucumbios" pageId="34" pageNumber="40">Sucumbíos</collectingRegion>
and 
<collectingRegion id="49F5F826FFB5AF4828943B66FD2CFE13" box="[649,745,367,393]" country="Ecuador" name="Orellana" pageId="34" pageNumber="40">Orellana</collectingRegion>
Provinces.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB5AF482ADA3B9AFE7EFDA7" pageId="34" pageNumber="40" type="materials_examined">
<paragraph id="8B8E36C4FFB5AF482ADA3B9AFC42FE4B" blockId="34.[151,1437,151,933]" pageId="34" pageNumber="40">
<emphasis id="B945EAD6FFB5AF482ADA3B9AFEC1FE37" bold="true" box="[199,260,403,429]" pageId="34" pageNumber="40">Male</emphasis>
<materialsCitation id="3B593C99FFB5AF482B103B9AFC46FE4B" collectingDate="2007-07-30" country="Ecuador" elevation="300" latitude="-0.49333608" location="Tropical Rainforest" longLatPrecision="1" longitude="-77.29143" municipality="Payamino Research Station" pageId="34" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="B945EAD6FFB5AF482B103B9AFEB8FE37" bold="true" box="[269,381,403,429]" pageId="34" pageNumber="40">
<typeStatus id="548A8866FFB5AF482B103B9AFEBDFE37" box="[269,376,403,429]" pageId="34" pageNumber="40" type="holotype">holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D37FD4DFFB5AF482B9B3B9AFE5EFE37" box="[390,411,403,429]" pageId="34" pageNumber="40" type="male">♂</specimenCount>
| 
<collectingCountry id="F3267654FFB5AF482BAE3B9AFDFFFE37" box="[435,570,403,429]" name="Ecuador" pageId="34" pageNumber="40">ECUADOR</collectingCountry>
(
<collectingRegion id="49F5F826FFB5AF4828563B9AFD6AFE34" box="[587,687,403,430]" country="Ecuador" name="Orellana" pageId="34" pageNumber="40">Orellana</collectingRegion>
) / 
<collectingMunicipality id="6BEAACBEFFB5AF4828CD3B9AFBCDFE37" box="[720,1032,403,429]" pageId="34" pageNumber="40">Payamino Research Station</collectingMunicipality>
/ 
<geoCoordinate id="EE055003FFB5AF482E3E3B9AFB06FE37" box="[1059,1219,403,429]" degrees="0" direction="south" minutes="29" orientation="latitude" pageId="34" pageNumber="40" precision="1" seconds="36.01" value="-0.49333608">0º 29’36.01”S</geoCoordinate>
<geoCoordinate id="EE055003FFB5AF482ED13B9AFA49FE37" box="[1228,1420,402,429]" degrees="77" direction="west" minutes="17" orientation="longitude" pageId="34" pageNumber="40" precision="1" seconds="29.15" value="-77.29143">77º 17’ 29.15”W</geoCoordinate>
/ 
<quantity id="4CC99B21FFB5AF482A8A3BBEFF12FE4B" box="[151,215,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="40" unit="m" value="300.0">
<elevation id="001CD1F7FFB5AF482A8A3BBEFF12FE4B" box="[151,215,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="40" unit="m" value="300.0">300m</elevation>
</quantity>
<location id="8EEE601FFFB5AF482AC03BBEFE72FE4B" LSID="urn:lsid:plazi:treatment:039887D2FFB7AF482A8A3813FB9EFC3F:8EEE601FFFB5AF482AC03BBEFE72FE4B" box="[221,439,439,465]" country="Ecuador" latitude="-0.49333608" longLatPrecision="1" longitude="-77.29143" municipality="Payamino Research Station" name="Tropical Rainforest" pageId="34" pageNumber="40" stateProvince="Orellana">Tropical Rainforest</location>
/ at. M.V. light / 
<date id="FF8F1004FFB5AF48286A3BBEFD3FFE4B" box="[631,762,439,465]" pageId="34" pageNumber="40" value="2007-07-30">
<collectingDate id="EFCBE9ECFFB5AF48286A3BBEFD3FFE4B" box="[631,762,439,465]" pageId="34" pageNumber="40" value="2007-07-30">30.vii.2007</collectingDate>
</date>
| (NHMUK)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8E36C4FFB5AF482ADA3BD2FE7EFDA7" blockId="34.[151,1437,151,933]" pageId="34" pageNumber="40">
<materialsCitation id="3B593C99FFB5AF482ADA3BD2FE72FDA7" collectingDate="1975-08-28" collectorName="Langley &amp; Cohen &amp; Cantor &amp; Yingling" country="Ecuador" determinerName="P. J. Spangler" location="Rio Aguarico" municipality="Lago Agrio" pageId="34" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="B945EAD6FFB5AF482ADA3BD2FEF8FE6F" bold="true" box="[199,317,475,501]" pageId="34" pageNumber="40">
<typeStatus id="548A8866FFB5AF482ADA3BD2FEFDFE6F" box="[199,312,475,501]" pageId="34" pageNumber="40" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D37FD4DFFB5AF482B5A3BD2FE99FE6F" box="[327,348,475,501]" pageId="34" pageNumber="40" type="male">♂</specimenCount>
| 
<collectingCountry id="F3267654FFB5AF482B6B3BD2FDC5FE6F" box="[374,512,475,501]" name="Ecuador" pageId="34" pageNumber="40">ECUADOR</collectingCountry>
, 
<collectingRegion id="49F5F826FFB5AF4828133BD2FD8EFE6F" box="[526,587,475,501]" country="Ecuador" name="Napo" pageId="34" pageNumber="40">Napo</collectingRegion>
[nowadays 
<collectingRegion id="49F5F826FFB5AF4828CA3BD3FC92FE6F" box="[727,855,474,501]" country="Ecuador" name="Sucumbios" pageId="34" pageNumber="40">Sucumbíos</collectingRegion>
] / 
<collectingMunicipality id="6BEAACBEFFB5AF4829643BD2FC39FE6F" box="[889,1020,475,501]" pageId="34" pageNumber="40">Lago Agrio</collectingMunicipality>
/ (18KmE) / 
<date id="FF8F1004FFB5AF482E803BD2FAFFFE6F" box="[1181,1338,474,501]" pageId="34" pageNumber="40" value="1975-08-28">
<collectingDate id="EFCBE9ECFFB5AF482E803BD2FAFFFE6F" box="[1181,1338,474,501]" pageId="34" pageNumber="40" value="1975-08-28">28 Aug. 1975</collectingDate>
</date>
at / 
<location id="8EEE601FFFB5AF482F693BD2FF3AFD83" LSID="urn:lsid:plazi:treatment:039887D2FFB7AF482A8A3813FB9EFC3F:8EEE601FFFB5AF482F693BD2FF3AFD83" country="Ecuador" municipality="Lago Agrio" name="Rio Aguarico" pageId="34" pageNumber="40" stateProvince="Napo">Rio Aguarico</location>
| Collected on / gravel bar / at blacklight / 
<collectorName id="26C45312FFB5AF4828F23BF6FC89FD83" box="[751,844,511,537]" pageId="34" pageNumber="40">Langley</collectorName>
, 
<collectorName id="26C45312FFB5AF4829443BF6FC60FD83" box="[857,933,511,537]" pageId="34" pageNumber="40">Cohen</collectorName>
, 
<collectorName id="26C45312FFB5AF4829AF3BF6FC3BFD83" box="[946,1022,511,537]" pageId="34" pageNumber="40">Cantor</collectorName>
&amp; 
<collectorName id="26C45312FFB5AF482E393BF6FB43FD83" box="[1060,1158,511,537]" pageId="34" pageNumber="40">Yingling</collectorName>
| HEXACYLLOEPUS / det. 
<determinerName id="7E52DA00FFB5AF482AA2382AFE8CFDA7" box="[191,329,547,573]" pageId="34" pageNumber="40">P.J.Spangler</determinerName>
(USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB5AF482ADA384EFC1CFDFB" box="[199,985,583,609]" pageId="34" pageNumber="40" type="etymology">
<paragraph id="8B8E36C4FFB5AF482ADA384EFC1CFDFB" blockId="34.[151,1437,151,933]" box="[199,985,583,609]" pageId="34" pageNumber="40">
<emphasis id="B945EAD6FFB5AF482ADA384EFE97FDFB" bold="true" box="[199,338,583,609]" pageId="34" pageNumber="40">Etymology:</emphasis>
<taxonomicName id="4C314D47FFB5AF482B44384EFE01FDFB" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[345,452,583,609]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="B945EAD6FFB5AF482B44384EFE01FDFB" box="[345,452,583,609]" italics="true" pageId="34" pageNumber="40">thoracica</emphasis>
</taxonomicName>
is a reference to a tubercle on the metaventrite.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFFB5AF482ADA3863FB9EFC3F" pageId="34" pageNumber="40" type="discussion">
<paragraph id="8B8E36C4FFB5AF482ADA3863FB9EFC3F" blockId="34.[151,1437,151,933]" pageId="34" pageNumber="40">
<emphasis id="B945EAD6FFB5AF482ADA3863FE70FD1F" bold="true" box="[199,437,618,645]" pageId="34" pageNumber="40">Comparative notes:</emphasis>
<taxonomicName id="4C314D47FFB5AF482BA23862FD1CFD1F" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[447,729,619,645]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="thoracica" status="sp. nov.">
<emphasis id="B945EAD6FFB5AF482BA23862FD1CFD1F" box="[447,729,619,645]" italics="true" pageId="34" pageNumber="40">Hexacylloepus thoracica</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FFB5AF4828FE3862FCFAFD1F" bold="true" box="[739,831,619,645]" pageId="34" pageNumber="40">
<taxonomicNameLabel id="A27657ADFFB5AF4828FE3862FCFAFD1F" box="[739,831,619,645]" pageId="34" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C314D47FFB5AF4829DE3862FAC2FD1F" authority="Hinton, 1969" authorityName="Hinton" authorityYear="1969" box="[963,1287,619,645]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="bassindalei">
<emphasis id="B945EAD6FFB5AF4829DE3862FBA2FD1F" box="[963,1127,619,645]" italics="true" pageId="34" pageNumber="40">H. bassindalei</emphasis>
<bibRefCitation id="EFA04B35FFB5AF482E6C3862FAC2FD1F" author="Hinton, H. E." box="[1137,1287,619,645]" pageId="34" pageNumber="40" pagination="125 - 130" refId="ref25979" refString="Hinton, H. E. (1969) Structure of the plastron of Hexacylloepus, with a description of a new species (Coleoptera, Elminthidae). Journal of Natural History, 3 (1), 125 - 130. https: // doi. org / 10.1080 / 00222936900770131" type="journal article" year="1969">Hinton, 1969</bibRefCitation>
</taxonomicName>
with a similar belt of tomentum on hypomera, long legs, and epipleura without tomentum. Nevertheless, the species can be differentiated by 
<taxonomicName id="4C314D47FFB5AF482B4638BAFE2EFD57" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[347,491,691,717]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="thoracica" status="sp. nov.">
<emphasis id="B945EAD6FFB5AF482B4638BAFE2EFD57" box="[347,491,691,717]" italics="true" pageId="34" pageNumber="40">H. thoracica</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FFB5AF482BE938BAFD8AFD57" bold="true" box="[500,591,691,717]" pageId="34" pageNumber="40">
<taxonomicNameLabel id="A27657ADFFB5AF482BE938BAFD8AFD57" box="[500,591,691,717]" pageId="34" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
having a long, deep longitudinal impression on the disc of the pronotum, reaching the anterior and posterior margins; antennae with all antennomeres with setae; and metacoxae without a spur. 
<taxonomicName id="4C314D47FFB5AF482AC838F2FDC7FC8F" authorityName="Hinton" authorityYear="1969" box="[213,514,763,789]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="bassindalei">
<emphasis id="B945EAD6FFB5AF482AC838F2FDC7FC8F" box="[213,514,763,789]" italics="true" pageId="34" pageNumber="40">Hexacylloepus bassindalei</emphasis>
</taxonomicName>
has a short longitudinal impression on the disc of the pronotum, not reaching the anterior and posterior margins; antennae with only the six apical antennomeres with setae; and a long and robust spur on the metacoxae. The male genitalia are also different: in 
<taxonomicName id="4C314D47FFB5AF482974394AFC32FCC7" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[873,1015,835,861]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="thoracica" status="sp. nov.">
<emphasis id="B945EAD6FFB5AF482974394AFC32FCC7" box="[873,1015,835,861]" italics="true" pageId="34" pageNumber="40">H. thoracica</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FFB5AF482E1D394AFB9FFCC7" bold="true" box="[1024,1114,835,861]" pageId="34" pageNumber="40">
<taxonomicNameLabel id="A27657ADFFB5AF482E1D394AFB9FFCC7" box="[1024,1114,835,861]" pageId="34" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the parameres extend 2/3 of the penis length, and the penis is elongated, with anterior half arcuate and posterior half straight; in 
<taxonomicName id="4C314D47FFB5AF482EE7396EFA58FC1B" authorityName="Hinton" authorityYear="1969" box="[1274,1437,871,897]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="40" phylum="Arthropoda" rank="species" species="bassindalei">
<emphasis id="B945EAD6FFB5AF482EE7396EFA58FC1B" box="[1274,1437,871,897]" italics="true" pageId="34" pageNumber="40">H. bassindalei</emphasis>
</taxonomicName>
the parameres reach 3/4 of the penis length, and the penis is pointed towards the apex.
</paragraph>
</subSubSection>
</treatment>
</document>