262 lines
31 KiB
XML
262 lines
31 KiB
XML
<document id="6A469795BB02B6E1A54B1325AFB9EF4C" ID-DOI="10.11646/zootaxa.4819.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4396744" ID-ZooBank="80258C53-F94D-46CB-9E86-AF0C3226DF9E" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1609199984737" checkinUser="plazi" docAuthor="Polizei, Thiago T. S., Barclay, Maxwell V. L. & Bispo, Pitágoras C." docDate="2020" docId="039887D2FF86AF782A8A3A91FEE9FA5B" docLanguage="en" docName="zootaxa.4819.1.2.pdf" docOrigin="Zootaxa 4819 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexacylloepus keitai Polizei & Barclay & Bispo 2020, sp. nov." docType="treatment" docVersion="1" lastPageNumber="24" masterDocId="FFA1FFAAFF97AF6A2A1D3A09FFC5FF9A" masterDocTitle="Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)" masterLastPageNumber="48" masterPageNumber="6" pageNumber="23" updateTime="1730286293773" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods id="959E3D27EF1C55D6C0C0A9FC12CE22E1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5B1863C97DA2847D711E46EBE1452263">
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<mods:title id="54E4F4DC4922B6A326952AAF9EF76998">Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="7EE8756BCD4EFC51AC677EDD52896B1B" type="personal">
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<mods:role id="767B0EEA54E76A17872C92D5FAD40CA0">
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<mods:roleTerm id="FD399AFFEB5BAD59F75E82FFB6F02CB4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5601FD80614B3EC05DBDC7D40FA37347">Polizei, Thiago T. S.</mods:namePart>
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<mods:nameIdentifier id="2A687E37C443150A0A8EB4302E2DE259" type="ORCID">0000-0003-3746-8865</mods:nameIdentifier>
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<mods:affiliation id="6626C22E9B49F2EAF3CD7CF191A8188E">Laboratório de Coleoptera (LaC) - Museu de Zoologia da Universidade de São Paulo (MZSP). São Paulo, São Paulo State, Brazil. thiagopolizei @ hotmail. com; https: // orcid. org / 0000 - 0003 - 3746 - 8865 & Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="DD497A80130F6AAB69C4FDED017EFB92" type="email">thiagopolizei@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="A140DCED05F37ABDA4B512AB109F8C7F" type="personal">
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<mods:role id="9FF480C175173675F1720380828AFCBD">
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<mods:roleTerm id="879348C499374697562239F34843B607">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F0C499362B132D75C387298798D2EC8A">Barclay, Maxwell V. L.</mods:namePart>
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<mods:affiliation id="8050371DE47F22B84A841C4CDDE77E53">Department of Life Sciences, Natural History Museum. London SW 7 5 BD, United Kingdom.</mods:affiliation>
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</mods:name>
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<mods:name id="0894D46B84850B3058B95CE6CAB35778" type="personal">
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<mods:role id="B23E7AAFD21E8926B27309CFF8254EB3">
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<mods:roleTerm id="DB22BC66326F6214BB46B81FCF01725C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A928D31C68967ED9A20CEE115D9D09FD">Bispo, Pitágoras C.</mods:namePart>
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<mods:affiliation id="46B396EDFE679BA03026680AF42B24E6">Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil. & Laboratório de Biologia Aquática (LABIA). Departamento de Ciências Biológicas - Faculdade de Ciências e Letras de Assis (FCL) - Universidade Estadual Paulista “ Júlio de Mesquita Filho ” (Unesp). Assis, São Paulo State, Brazil.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="5CEE8A24A6B455304F380CEF07D06A7A">text</mods:typeOfResource>
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<mods:titleInfo id="6F531388D776E8A1BDF5C83A5704ED4F">
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<mods:title id="46A9F31EC24FE780EEC7350FBB188D3D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B580A08BFAD535B53F846F515444B474">
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<mods:date id="51988392385DD2910941DD7C028113B1">2020</mods:date>
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<mods:detail id="6236D81620525D436998598FAE730825" type="pubDate">
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<mods:number id="5C46F0B2451FB079FF18066EAB7D5045">2020-07-22</mods:number>
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</mods:detail>
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<mods:detail id="65B72485D06170EE6A6FCCAE5F7360DF" type="volume">
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<mods:number id="95F14B264C5BBEA19CC44FEAC76C756F">4819</mods:number>
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</mods:detail>
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<mods:detail id="D85F4811E1A9A0EEDEE67000265A22D9" type="issue">
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<mods:number id="2D594CFC27A7F5AF8834B3D8477A2805">1</mods:number>
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<mods:start id="96C22FE947C094127B26C271A6D7DDDD">6</mods:start>
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<mods:end id="E7CB05DF23A37327B954A24C2530D5AF">48</mods:end>
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</mods:part>
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<mods:classification id="A93DDCB2223CA05313FD261DD2816C77">journal article</mods:classification>
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<mods:identifier id="373384C9F14EB050C80FFDA1EC806A50" type="DOI">10.11646/zootaxa.4819.1.2</mods:identifier>
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<mods:identifier id="C80F04BAA88DDC25C20A0FB1DBA4D0A4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4B314B7E476A5415A5C4729C00771151" type="Zenodo-Dep">4396744</mods:identifier>
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<mods:identifier id="8D62F6ADC555E13A92436C0C4191783B" type="ZooBank">80258C53-F94D-46CB-9E86-AF0C3226DF9E</mods:identifier>
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</mods:mods>
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<treatment id="039887D2FF86AF782A8A3A91FEE9FA5B" LSID="urn:lsid:plazi:treatment:039887D2FF86AF782A8A3A91FEE9FA5B" httpUri="http://treatment.plazi.org/id/039887D2FF86AF782A8A3A91FEE9FA5B" lastPageId="18" lastPageNumber="24" pageId="17" pageNumber="23">
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<subSubSection id="C32B654FFF86AF7B2A8A3A91FE34FF28" box="[151,497,152,178]" pageId="17" pageNumber="23" type="nomenclature">
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<paragraph id="8B8E36C4FF86AF7B2A8A3A91FE34FF28" blockId="17.[151,497,152,214]" box="[151,497,152,178]" pageId="17" pageNumber="23">
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<taxonomicName id="4C314D47FF86AF7B2A8A3A91FE49FF28" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[151,396,152,178]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="23" phylum="Arthropoda" rank="species" species="keitai" status="sp. nov.">
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<emphasis id="B945EAD6FF86AF7B2A8A3A91FE49FF28" box="[151,396,152,178]" italics="true" pageId="17" pageNumber="23">Hexacylloepus keitai</emphasis>
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</taxonomicName>
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<emphasis id="B945EAD6FF86AF7B2B893A91FE34FF28" bold="true" box="[404,497,152,178]" pageId="17" pageNumber="23">
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<taxonomicNameLabel id="A27657ADFF86AF7B2B893A91FE34FF28" box="[404,497,152,178]" pageId="17" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32B654FFF86AF7B2A8A3AB5FEE3FF4C" box="[151,294,188,214]" pageId="17" pageNumber="23" type="description">
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<paragraph id="8B8E36C4FF86AF7B2A8A3AB5FEE3FF4C" blockId="17.[151,497,152,214]" box="[151,294,188,214]" pageId="17" pageNumber="23">
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||
(
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<figureCitation id="130A2A41FF86AF7B2ABD3AB5FF27FF4C" box="[160,226,188,214]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="17" pageNumber="23">Fig. 8</figureCitation>
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A–B)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF86AF7B2A8A3B0DFCCDFE84" box="[151,776,259,286]" pageId="17" pageNumber="23" type="materials_examined">
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<paragraph id="8B8E36C4FF86AF7B2A8A3B0DFCCDFE84" blockId="17.[151,1437,259,862]" box="[151,776,259,286]" pageId="17" pageNumber="23">
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<emphasis id="B945EAD6FF86AF7B2A8A3B0DFEFCFE87" bold="true" box="[151,313,259,286]" pageId="17" pageNumber="23">
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<typeStatus id="548A8866FF86AF7B2A8A3B0DFF17FE84" box="[151,210,260,286]" pageId="17" pageNumber="23">Type</typeStatus>
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locality:
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</emphasis>
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<collectingCountry id="F3267654FF86AF7B2B5D3B0DFE5DFE84" box="[320,408,260,286]" name="Mexico" pageId="17" pageNumber="23">Mexico</collectingCountry>
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,
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<collectingRegion id="49F5F826FF86AF7B2BB93B0DFD53FE84" box="[420,662,260,286]" country="Mexico" name="San Luis Potosi" pageId="17" pageNumber="23">San Luis Potosi State</collectingRegion>
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,
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<collectorName id="26C45312FF86AF7B28BC3B0DFCC1FE84" box="[673,772,260,286]" pageId="17" pageNumber="23">La Cova</collectorName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF86AF7B2ADA3B2EFD05FE48" pageId="17" pageNumber="23" type="diagnosis">
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<paragraph id="8B8E36C4FF86AF7B2ADA3B2EFD05FE48" blockId="17.[151,1437,259,862]" pageId="17" pageNumber="23">
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<emphasis id="B945EAD6FF86AF7B2ADA3B2EFEFFFEDB" bold="true" box="[199,314,295,321]" pageId="17" pageNumber="23">Diagnosis</emphasis>
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: Antennae without setae. Frontoclypeal suture inconspicuous. Pronotum with a short and deep impression on disc. Hypomera with a belt of tomentum reaching the lateral and anterior margins. Epipleura with tomentum. Tibiae each with a row of setae on distal half; meso- and metatibiae each with a row of spines in distal half. Abdomen with the disc of the first ventrite without tomentum, and a deep depression with a pair of carinae extending towards and reaching the metaventrite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B654FFF86AF782ADA3BD2FED1FCA3" lastPageId="18" lastPageNumber="24" pageId="17" pageNumber="23" type="description">
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<paragraph id="8B8E36C4FF86AF7B2ADA3BD2FCD4FD80" blockId="17.[151,1437,259,862]" pageId="17" pageNumber="23">
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<emphasis id="B945EAD6FF86AF7B2ADA3BD2FE9EFE6F" bold="true" box="[199,347,475,501]" pageId="17" pageNumber="23">Description:</emphasis>
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Male. Body elongated, subparallel. Black colour, with the antenna, mouthparts and legs reddishbrown. Total length:
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<quantity id="4CC99B21FF86AF7B2B9C3BF6FE2DFD80" box="[385,488,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="17" pageNumber="23" unit="mm" value="1.61">1.61 mm</quantity>
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. Greatest width:
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<quantity id="4CC99B21FF86AF7B28BB3809FCC8FD80" box="[678,781,512,538]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="23" unit="mm" value="0.7">0.70 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B8E36C4FF86AF7B2ADA382DFE60FD68" blockId="17.[151,1437,259,862]" pageId="17" pageNumber="23">
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<emphasis id="B945EAD6FF86AF7B2ADA382DFEC8FDA4" box="[199,269,548,574]" italics="true" pageId="17" pageNumber="23">Head:</emphasis>
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surface with micropunctures separated from each other by their diameter and with short setae. Antenna inserted on the anterior margin of the eyes, filiform, with 11 antennomeres without setae. Frontoclypeal suture inconspicuous. Clypeus 5x as wide as long; anterior margin strongly concave and posterior sinuous; surface similar to that of head. Labrum twice as wide as long; antero-lateral angles rounded and surface with micropunctures. Maxillary palpus 4-segmented; terminal palpomere truncated apically. Labial palpus 3-segmented. Mentum and submentum rectangular.
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</paragraph>
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<paragraph id="8B8E36C4FF86AF7B2ADA38F5FCC1FCC4" blockId="17.[151,1437,259,862]" pageId="17" pageNumber="23">
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<emphasis id="B945EAD6FF86AF7B2ADA38F5FEE4FC8C" box="[199,289,764,790]" italics="true" pageId="17" pageNumber="23">Thorax:</emphasis>
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Pronotum as long as wide, arcuate anteriorly; sides sinuate and serrate, posterior margin smooth with two prescutellar foveae (
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<figureCitation id="130A2A41FF86AF7B2BB73929FDC1FCA0" box="[426,516,800,826]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="17" pageNumber="23">Fig. 8A</figureCitation>
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). Surface with micropunctures and setae. Sublateral carinae complete, sinuous and raised (
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<figureCitation id="130A2A41FF86AF7B2AF4394DFE86FCC4" box="[233,323,836,862]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="17" pageNumber="23">Fig. 8A</figureCitation>
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). Longitudinal impression deep on disc.
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</paragraph>
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<caption id="DF4E664CFF86AF7B2A8A3CDCFA49F974" ID-DOI="http://doi.org/10.5281/zenodo.4396760" ID-Zenodo-Dep="4396760" box="[151,1420,1749,1774]" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="17" pageNumber="23" startId="17.[151,250,1749,1773]" targetBox="[161,1424,896,1717]" targetPageId="17">
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<paragraph id="8B8E36C4FF86AF7B2A8A3CDCFA49F974" blockId="17.[151,1420,1749,1774]" box="[151,1420,1749,1774]" pageId="17" pageNumber="23">
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<emphasis id="B945EAD6FF86AF7B2A8A3CDCFED6F974" bold="true" box="[151,275,1749,1774]" pageId="17" pageNumber="23">FIGURE 8.</emphasis>
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<taxonomicName id="4C314D47FF86AF7B2B043CDFFE28F974" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[281,493,1750,1774]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="23" phylum="Arthropoda" rank="species" species="keitai" status="sp. nov.">
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<emphasis id="B945EAD6FF86AF7B2B043CDFFE28F974" box="[281,493,1750,1774]" italics="true" pageId="17" pageNumber="23">Hexacylloepus keitai</emphasis>
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</taxonomicName>
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<emphasis id="B945EAD6FF86AF7B2BEE3CDFFD81F974" bold="true" box="[499,580,1750,1774]" pageId="17" pageNumber="23">
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<taxonomicNameLabel id="A27657ADFF86AF7B2BEE3CDFFD81F974" box="[499,580,1750,1774]" pageId="17" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm.
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</paragraph>
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</caption>
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<paragraph id="8B8E36C4FF86AF7B2ADA3D15FD6AF85C" blockId="17.[151,1437,1820,2026]" pageId="17" pageNumber="23">
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Elytra moderately longer than wide; anterior margin arcuate, lateral margin moderately arcuate and serrate; rounded apex (
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<figureCitation id="130A2A41FF86AF7B2B5C3D49FE58F8C0" box="[321,413,1856,1882]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="17" pageNumber="23">Fig. 8A</figureCitation>
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). Humeri slightly prominent. Surface with puncture and setae rows. Elytral suture moderately elevated. Carina short and elevated on interval IV, extending from base to 1/5 of the elytral length; sublateral carinae on intervals VI and VIII, extending from base to 3/4 of the elytral length. Epipleura with tomentum (
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<figureCitation id="130A2A41FF86AF7B2F723D81FF7EF85C" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="17" pageNumber="23">Fig. 8B</figureCitation>
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). Hind wings macropterous. Scutellum oval.
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</paragraph>
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<paragraph id="8B8E36C4FF86AF782ADA3DD9FDACFE87" blockId="17.[151,1437,1820,2026]" lastBlockId="18.[151,1437,151,1473]" lastPageId="18" lastPageNumber="24" pageId="17" pageNumber="23">
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Prosternum narrow, strongly elevated; surface with micropunctures, anterior margin straight with many setae and lateral margin covered with tomentum. Disc moderately raised. Prosternal process long and narrow and beyond the procoxae; lateral margin elevated; rounded apex. Notosternal suture slightly sinuous. Hypomera wider posteriorly than anteriorly, lateral margin serrate; surface with micropunctures and with a thin belt of tomentum reaching the lateral and anterior margins (
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<figureCitation id="130A2A41FF85AF7828193B0AFD99FE87" box="[516,604,259,285]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="18" pageNumber="24">Fig. 8B</figureCitation>
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).
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</paragraph>
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<paragraph id="8B8E36C4FF85AF782ADA3B2EFE8CFE13" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">Mesoventrite with a groove to receive the prosternal process. Mesepimeron and mesepisternum covered with tomentum. Metaventrite broad with a longitudinal medial depression; surface with micropunctures, sides covered with tomentum.</paragraph>
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<paragraph id="8B8E36C4FF85AF782ADA3B9AFF0FFDA7" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
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Legs (except tarsi) covered with punctures and tomentum (
|
||
<figureCitation id="130A2A41FF85AF78296C3B9AFC0EFE37" box="[881,971,403,429]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="18" pageNumber="24">Fig. 8B</figureCitation>
|
||
). Coxae rounded; trochanters elongated; femora short and robust, 3x as long as wide, 1/4 of the elytra length; tibiae short and thin, 4x as long as wide, with 1/4 of the elytra length; tibiae each with a row of setae on distal half. Meso and metatibiae each with a row of spines on distal half. Tarsi elongate, with setae; apical tarsomere longer than the basal four combined. Claws long and thin.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF85AF782ADA384EFBB1FD6B" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA384EFEF4FDFB" box="[199,305,583,609]" italics="true" pageId="18" pageNumber="24">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="130A2A41FF85AF782B5C384EFE5EFDFB" box="[321,411,583,609]" captionStart="FIGURE 8" captionStartId="17.[151,250,1749,1773]" captionTargetBox="[161,1424,896,1717]" captionTargetId="figure-265@17.[151,1435,886,1726]" captionTargetPageId="17" captionText="FIGURE 8. Hexacylloepus keitai sp. nov.. Male. Holotype. Habitus. (A) Dorsal view. (B) Ventral view. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4396760" httpUri="https://zenodo.org/record/4396760/files/figure.png" pageId="18" pageNumber="24">Fig. 8B</figureCitation>
|
||
): five ventrites, as long as wide; strongly raised on middle. Anterior and posterior margins straight on the first and second ventrites and arcuate on the subsequent ventrites; lateral margins rounded. Surface densely covered with tomentum (excepted for disc of first ventrite), micropunctures and short setae. Disc of the first ventrite with a deep depression and a pair of carinae extending towards and reaching the metaventrite. Fifth ventrite with lateral margins slightly produced posterolaterad; and apex rounded with long setae.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF85AF782ADA38F2FED1FCA3" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA38F2FEEDFC8F" bold="true" box="[199,296,763,789]" pageId="18" pageNumber="24">Female:</emphasis>
|
||
External morphology similar to male, except for the absence of the row of spines on the meso- and metatibiae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF85AF782ADA394BFD8EFCC7" box="[199,587,834,861]" pageId="18" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8B8E36C4FF85AF782ADA394BFD8EFCC7" blockId="18.[151,1437,151,1473]" box="[199,587,834,861]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA394BFE76FCC6" bold="true" box="[199,435,834,860]" pageId="18" pageNumber="24">Material examined:</emphasis>
|
||
<specimenCount id="9D37FD4DFF85AF782BA7394AFD82FCC7" box="[442,583,835,861]" pageId="18" pageNumber="24" type="generic">3 specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF85AF782ADA396FFD01FC1B" box="[199,708,870,897]" pageId="18" pageNumber="24" type="distribution">
|
||
<paragraph id="8B8E36C4FF85AF782ADA396FFD01FC1B" blockId="18.[151,1437,151,1473]" box="[199,708,870,897]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA396FFEA6FC1A" bold="true" box="[199,355,870,896]" pageId="18" pageNumber="24">Distribution:</emphasis>
|
||
<collectingCountry id="F3267654FF85AF782B77396EFE01FC1B" box="[362,452,871,897]" name="Mexico" pageId="18" pageNumber="24">Mexico</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F5F826FF85AF782BD3396EFD04FC1B" box="[462,705,871,897]" country="Mexico" name="San Luis Potosi" pageId="18" pageNumber="24">San Luis Potosi State</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF85AF782ADA3982FBDDFB8B" pageId="18" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8B8E36C4FF85AF782ADA3982FBCAFC53" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<materialsCitation id="3B593C99FF85AF782ADA3982FBCAFC53" collectionCode="NHMUK" collectorName="H. E. Hinton" determinerName="Spangler" location="La Cova" pageId="18" pageNumber="24" specimenCode="B.M.1977-566" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B945EAD6FF85AF782ADA3982FEBBFC3F" bold="true" box="[199,382,907,933]" pageId="18" pageNumber="24">
|
||
<specimenCount id="9D37FD4DFF85AF782ADA3982FEC1FC3F" box="[199,260,907,933]" pageId="18" pageNumber="24" type="male">Male</specimenCount>
|
||
<typeStatus id="548A8866FF85AF782B133982FEBCFC3F" box="[270,377,907,933]" pageId="18" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
♂ |
|
||
<location id="8EEE601FFF85AF782BA53982FDDAFC3F" LSID="urn:lsid:plazi:treatment:039887D2FF86AF782A8A3A91FEE9FA5B:8EEE601FFF85AF782BA53982FDDAFC3F" box="[440,543,907,933]" name="La Cova" pageId="18" pageNumber="24">La Cova</location>
|
||
. SLP / x.70 Mex. /
|
||
<locationDeviation id="9A3C2FE6FF85AF7829113982FC7BFC3F" box="[780,958,907,933]" pageId="18" pageNumber="24">
|
||
<quantity id="4CC99B21FF85AF7829113982FC8EFC3F" box="[780,843,907,933]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="24" unit="km" value="23.0">23km</quantity>
|
||
C. Valles
|
||
</locationDeviation>
|
||
/ H.E.Hinton |
|
||
<collectorName id="26C45312FF85AF782E6A3982FB32FC3F" box="[1143,1271,907,933]" pageId="18" pageNumber="24">H.E.Hinton</collectorName>
|
||
/ collection. /
|
||
<specimenCode id="DB979EBFFF85AF782A8A39A7FE85FC53" box="[151,320,942,969]" pageId="18" pageNumber="24">B.M.1977-566</specimenCode>
|
||
. | Hexacylloepus
|
||
<taxonomicNameLabel id="A27657ADFF85AF78281939A6FD85FC53" box="[516,576,943,969]" pageId="18" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
|
||
/ det.
|
||
<determinerName id="7E52DA00FF85AF78289B39A6FD2FFC53" box="[646,746,943,969]" pageId="18" pageNumber="24">Spangler</determinerName>
|
||
| Holo- / type (
|
||
<collectionCode id="ED20AE01FF85AF78298E39A6FBC1FC53" box="[915,1028,943,969]" pageId="18" pageNumber="24">NHMUK</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF85AF782ADA39DAFBDDFB8B" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<materialsCitation id="3B593C99FF85AF782ADA39DAFBDDFB8B" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA39DAFE8DFC77" bold="true" box="[199,328,979,1005]" pageId="18" pageNumber="24">
|
||
<typeStatus id="548A8866FF85AF782ADA39DAFE86FC77" box="[199,323,979,1005]" pageId="18" pageNumber="24" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D37FD4DFF85AF782B5239DAFE31FC77" box="[335,500,979,1005]" pageId="18" pageNumber="24" type="female">2 specimens ♀</specimenCount>
|
||
| La Cova. SLP / x.70 Mex. /
|
||
<quantity id="4CC99B21FF85AF78292339DAFCB8FC77" box="[830,893,979,1005]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="24" unit="km" value="23.0">23km</quantity>
|
||
C. Valles / H.E.Hinton | H.E.Hinton / collection. / B.M.1977-566. | Hexacylloepus
|
||
<taxonomicNameLabel id="A27657ADFF85AF78280E39FEFD8AFB8B" box="[531,591,1015,1041]" pageId="18" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
|
||
/ det. Spangler | Para- / type (NHMUK).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF85AF782ADA3E12FECCFBC3" pageId="18" pageNumber="24" type="etymology">
|
||
<paragraph id="8B8E36C4FF85AF782ADA3E12FECCFBC3" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA3E12FE97FBAF" bold="true" box="[199,338,1051,1077]" pageId="18" pageNumber="24">Etymology:</emphasis>
|
||
<taxonomicName id="4C314D47FF85AF782B423E12FE58FBAF" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[351,413,1051,1077]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="24" phylum="Arthropoda" rank="species" species="keitai">
|
||
<emphasis id="B945EAD6FF85AF782B423E12FE58FBAF" box="[351,413,1051,1077]" italics="true" pageId="18" pageNumber="24">keitai</emphasis>
|
||
</taxonomicName>
|
||
is a tribute to Keita Matsumoto, Curatorial Assistant of the
|
||
<taxonomicName id="4C314D47FF85AF782E993E12FB3AFBAF" box="[1156,1279,1051,1077]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="24" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
|
||
collection at NHMUK.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B654FFF85AF782ADA3E6BFEE9FA5B" pageId="18" pageNumber="24" type="discussion">
|
||
<paragraph id="8B8E36C4FF85AF782ADA3E6BFB4CFA97" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA3E6BFE76FBE7" bold="true" box="[199,435,1122,1149]" pageId="18" pageNumber="24">Comparative notes:</emphasis>
|
||
<taxonomicName id="4C314D47FF85AF782BA73E6AFD60FBE7" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[442,677,1123,1149]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="24" phylum="Arthropoda" rank="species" species="keitai" status="sp. nov.">
|
||
<emphasis id="B945EAD6FF85AF782BA73E6AFD60FBE7" box="[442,677,1123,1149]" italics="true" pageId="18" pageNumber="24">Hexacylloepus keitai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EAD6FF85AF7828B13E6AFCC3FBE7" bold="true" box="[684,774,1123,1149]" pageId="18" pageNumber="24">
|
||
<taxonomicNameLabel id="A27657ADFF85AF7828B13E6AFCC3FBE7" box="[684,774,1123,1149]" pageId="18" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C314D47FF85AF7829983E6AFB00FBE7" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[901,1221,1123,1149]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="24" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B945EAD6FF85AF7829983E6AFBE6FBE7" box="[901,1059,1123,1149]" italics="true" pageId="18" pageNumber="24">H. granulosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B35FF85AF782E2E3E6AFB78FBE7" author="Sharp, D." box="[1075,1213,1123,1149]" pageId="18" pageNumber="24" pagination="116 - 141" refId="ref26633" refString="Sharp, D. (1882) Heteroceridae. Parnidae, Georissidae. Biologia Centrali-Americana, Coleoptera, 1 (2), 116 - 141." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by the inconspicuous frontoclypeal suture, pronotum with the longitudinal impression short and deep on disc, and short legs. However, the species can be differentiated by the epipleura in
|
||
<taxonomicName id="4C314D47FF85AF7829043EA2FCBCFB5F" authority="Polizei & Barclay & Bispo, 2020" authorityName="Polizei & Barclay & Bispo" authorityYear="2020" box="[793,889,1195,1221]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="24" phylum="Arthropoda" rank="species" species="keitai" status="sp. nov.">
|
||
<emphasis id="B945EAD6FF85AF7829043EA2FCBCFB5F" box="[793,889,1195,1221]" italics="true" pageId="18" pageNumber="24">H. keitai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EAD6FF85AF78299D3EA2FC1CFB5F" bold="true" box="[896,985,1195,1221]" pageId="18" pageNumber="24">
|
||
<taxonomicNameLabel id="A27657ADFF85AF78299D3EA2FC1CFB5F" box="[896,985,1195,1221]" pageId="18" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
completely covered with tomentum and hypomera with belt of tomentum reaching the anterior and posterior margin, while in
|
||
<taxonomicName id="4C314D47FF85AF782E7F3EC6FAC7FB73" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[1122,1282,1231,1257]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="24" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B945EAD6FF85AF782E7F3EC6FAC7FB73" box="[1122,1282,1231,1257]" italics="true" pageId="18" pageNumber="24">H. granulosus</emphasis>
|
||
</taxonomicName>
|
||
the epipleura lack tomentum and the belt of tomentum on the hypomera only reaches the lateral margin.
|
||
</paragraph>
|
||
<paragraph id="8B8E36C4FF85AF782ADA3F1FFEE9FA5B" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="24">
|
||
<emphasis id="B945EAD6FF85AF782ADA3F1FFE97FAAA" bold="true" box="[199,338,1302,1328]" pageId="18" pageNumber="24">Comments:</emphasis>
|
||
The new species was found in the NHMUK collection and had already been recognised as a new species by P. Spangler (no further information) but never published. Spangler studied and dissected the three specimens (one male and two females), but unfortunately, we were not able to find the aedeagus of the male. However, the species has several external morphological characters that clearly identify it as a new species and allowed it to be described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |