136 lines
13 KiB
XML
136 lines
13 KiB
XML
<document id="77F9E46EBFEFE947C7E336B36D108D25" ID-CLB-Dataset="46369" ID-DOI="10.11646/zootaxa.3768.2.3" ID-GBIF-Dataset="c232760e-33e0-499b-adc1-ede336e64d29" ID-ISSN="1175-5326" ID-Zenodo-Dep="252057" ID-ZooBank="40B1B7AF-84E8-4EC0-97DA-68E4BD0C482A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460388281548" checkinUser="plazi" docAuthor="Maryan, Brad, Oliver, Paul M., Fitch, Alison J. & O’Connell, Morgan" docDate="2014" docId="039887CBFFADFF80F284F359FE95ABD5" docLanguage="en" docName="zt03768p158.pdf" docOrigin="Zootaxa 3768 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Varanus pilbarensis" docType="treatment" docVersion="9" lastPageNumber="145" masterDocId="FFA1FFB3FFA8FF86F213F602FFAFAB00" masterDocTitle="Molecular and morphological assessment of Varanus pilbarensis (Squamata: Varanidae), with a description of a new species from the southern Pilbara, Western Australia" masterLastPageNumber="158" masterPageNumber="139" pageNumber="144" updateTime="1698297920950" updateUser="ExternalLinkService">
|
||
<mods:mods id="1CE1B9260EBAAD6A5AE4BBD69329D579" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="7CD2F1A759C04E12790E614B948D2DDC">
|
||
<mods:title id="1D9A558B02C11EDE09E5C3351845DD19">Molecular and morphological assessment of Varanus pilbarensis (Squamata: Varanidae), with a description of a new species from the southern Pilbara, Western Australia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="ABDC45B7DFA40A1CAEECAEA6F7B3318A" type="personal">
|
||
<mods:role id="5F30F8D9C19D6510B663494CDCE1321E">
|
||
<mods:roleTerm id="C4A9173B591E8A34EA1FEDA4F03B96E7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7067D36839B80E4BEAB3E06221920A58">Maryan, Brad</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D013FD743C595F69D45C280A87651258" type="personal">
|
||
<mods:role id="F94CB70A21919F5D606E3BDBE6B4E6BF">
|
||
<mods:roleTerm id="D8785E7C20C2950B9344F393B4FEE8F2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="957AFF7E0473A2AE4BE31DA9CD1C66AB">Oliver, Paul M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="5D4018448F19C912B7C966029F10D021" type="personal">
|
||
<mods:role id="002CCA30271FE233C2B0C342DA0F9028">
|
||
<mods:roleTerm id="FEC09D4F33F567E3172B75396F4E42EF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AF26D63A296A4D4C2A68BB18535F1E76">Fitch, Alison J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1F68FE37F9B6AC841A87CC58932BFDF2" type="personal">
|
||
<mods:role id="3974A5A389CD4B81DC484B9928C3A7A5">
|
||
<mods:roleTerm id="774D98F4CEFA462351FC5ED87BAE92E3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="26F8B8F16FE87D246B08D4B19F5FAA5C">O’Connell, Morgan</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D70566E41E73C417586042B131F744CF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="05D1FDEB63022ADE37BED4CB3488D4C2" type="host">
|
||
<mods:titleInfo id="96D2229D3D3F292F54909A8255759DF3">
|
||
<mods:title id="E292CAB3D16EE51142D17A60ECAF1586">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="6F27BE7A54AF4772A54D98BC58D47038">
|
||
<mods:date id="8050FE158DF314316605EB00838D4811">2014</mods:date>
|
||
<mods:detail id="B781E4F98C7FC54684B0E4D2129C315B" type="volume">
|
||
<mods:number id="ED841E20A8207AED6496A8E8327B8CDE">3768</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="488B7BD07A8CECEAB033EB69F685E3AB" type="issue">
|
||
<mods:number id="52BC8E044490EB507E6EC01302009EEA">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7EA662CD1CF0A76A624CE2968C82FC79" unit="page">
|
||
<mods:start id="182834335A2EAFB37367A3D8DB94B757">139</mods:start>
|
||
<mods:end id="19B4E779E39D43E52A9C8B14FA7BF923">158</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BEF429C3796568A223A7E603D096D02A">journal article</mods:classification>
|
||
<mods:identifier id="F742A67F8E6BEC3F71B4F786B5BD93CA" type="CLB-Dataset">46369</mods:identifier>
|
||
<mods:identifier id="0B794C1A0242609F3A0C307C4B78572E" type="DOI">10.11646/zootaxa.3768.2.3</mods:identifier>
|
||
<mods:identifier id="E405D3D6E490E83BB56DE2BCB11FC8A1" type="GBIF-Dataset">c232760e-33e0-499b-adc1-ede336e64d29</mods:identifier>
|
||
<mods:identifier id="5D45C9D66E8F96C48DB12A67201AA1FB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="01A422AE8A990FEDCAD35DF79CDD1CEE" type="Zenodo-Dep">252057</mods:identifier>
|
||
<mods:identifier id="51F0040594E20153277F31D0B05FB428" type="ZooBank">40B1B7AF-84E8-4EC0-97DA-68E4BD0C482A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="039887CBFFADFF80F284F359FE95ABD5" ID-DOI="http://doi.org/10.5281/zenodo.5613435" ID-GBIF-Taxon="119416044" ID-Zenodo-Dep="5613435" LSID="urn:lsid:plazi:treatment:039887CBFFADFF80F284F359FE95ABD5" httpUri="http://treatment.plazi.org/id/039887CBFFADFF80F284F359FE95ABD5" lastPageId="6" lastPageNumber="145" pageId="5" pageNumber="144">
|
||
<subSubSection id="C32B6556FFADFF83F284F359FD97AE76" box="[151,568,1371,1398]" pageId="5" pageNumber="144" type="nomenclature">
|
||
<paragraph id="8B8E36DDFFADFF83F284F359FD97AE76" blockId="5.[151,568,1371,1398]" box="[151,568,1371,1398]" pageId="5" pageNumber="144">
|
||
<heading id="D0C681B1FFADFF83F284F359FD97AE76" bold="true" box="[151,568,1371,1398]" fontSize="11" level="1" pageId="5" pageNumber="144" reason="1">
|
||
<emphasis id="B945EACFFFADFF83F284F359FD97AE76" bold="true" box="[151,568,1371,1398]" pageId="5" pageNumber="144">
|
||
<taxonomicName id="4C314D5EFFADFF83F284F359FE28AE75" box="[151,391,1371,1397]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="pilbarensis">
|
||
<emphasis id="B945EACFFFADFF83F284F359FE28AE75" bold="true" box="[151,391,1371,1397]" italics="true" pageId="5" pageNumber="144">Varanus pilbarensis</emphasis>
|
||
</taxonomicName>
|
||
species-group
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6556FFADFF80F284F3A0FE95ABD5" lastPageId="6" lastPageNumber="145" pageId="5" pageNumber="144" type="diagnosis">
|
||
<paragraph id="8B8E36DDFFADFF83F284F3A0FF7AAD6F" blockId="5.[151,1436,1442,2007]" pageId="5" pageNumber="144">
|
||
<emphasis id="B945EACFFFADFF83F284F3A0FEBEAEBB" bold="true" box="[151,273,1442,1467]" pageId="5" pageNumber="144">Diagnosis.</emphasis>
|
||
Differs from all other Australian
|
||
<taxonomicName id="4C314D5EFFADFF83F09CF3A6FD44AEBB" box="[655,747,1444,1467]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="genus">
|
||
<emphasis id="B945EACFFFADFF83F09CF3A6FD44AEBB" box="[655,747,1444,1467]" italics="true" pageId="5" pageNumber="144">Varanus</emphasis>
|
||
</taxonomicName>
|
||
in the combination of small size (adult SVL up to
|
||
<quantity id="4CC99B38FFADFF83F73DF3A1FA20AEBB" box="[1326,1423,1443,1467]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="144" unit="mm" value="180.0">180 mm</quantity>
|
||
); slender build; tail long (170–210% length of body) and thin, circular in cross section at midpoint, midbody scales 118–128; dorsal and lateral scales weakly keeled, enlarged keeled ventrolateral scales on each side of and posterior to vent in 4–6 rows, scales on top of head smooth; supraoculars gradually merging with the larger interoculars; nostrils high and oriented dorsolaterally, and reddish base colouration of the dorsal and lateral surfaces of head and body.
|
||
</paragraph>
|
||
<paragraph id="8B8E36DDFFADFF83F2D4F078FD2DACD7" blockId="5.[151,1436,1442,2007]" pageId="5" pageNumber="144">
|
||
<emphasis id="B945EACFFFADFF83F2D4F078FDE3AD93" bold="true" box="[199,588,1658,1683]" pageId="5" pageNumber="144">Comparisons with other species.</emphasis>
|
||
The diagnosis above readily differentiates the
|
||
<taxonomicName id="4C314D5EFFADFF83F673F07EFB57AD93" box="[1120,1272,1658,1683]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="pilbarensis">
|
||
<emphasis id="B945EACFFFADFF83F673F07EFB57AD93" box="[1120,1272,1658,1683]" italics="true" pageId="5" pageNumber="144">V. pilbarensis</emphasis>
|
||
</taxonomicName>
|
||
species-group from all congeners, however a small number of species warrant additional comparison:
|
||
<taxonomicName id="4C314D5EFFADFF83F667F0A2FAB8ADB7" box="[1140,1303,1696,1719]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="tristis">
|
||
<emphasis id="B945EACFFFADFF83F667F0A2FAB8ADB7" box="[1140,1303,1696,1719]" italics="true" pageId="5" pageNumber="144">Varanus tristis</emphasis>
|
||
</taxonomicName>
|
||
also occurs in the Pilbara region but is larger (maximum SVL up to
|
||
<quantity id="4CC99B38FFADFF83F10BF0C1FCD9ADDA" box="[792,886,1731,1755]" metricMagnitude="-1" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="144" unit="mm" value="280.0">280 mm</quantity>
|
||
v.
|
||
<quantity id="4CC99B38FFADFF83F18AF0C1FC53ADDB" box="[921,1020,1731,1755]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="144" unit="mm" value="180.0">180 mm</quantity>
|
||
), has more midbody scales (between 119–152 v. 118–128), a less depressed head with a comparatively shorter snout, and a larger and more spinose cloacal scale cluster (
|
||
<bibRefCitation id="EFA04B2CFFADFF83F382F109FDF7AC23" author="Storr" box="[401,600,1802,1827]" pageId="5" pageNumber="144" refString="Storr, G. M., Smith, L. A. & Johnstone, R. E. (1983) Lizards of Western Australia. II. Dragons and Monitors. Western Australian Museum, Perth, Western Australia, Australia, 113 pp." type="book" year="1983">
|
||
Storr
|
||
<emphasis id="B945EACFFFADFF83F3C6F10EFDBDAC23" box="[469,530,1802,1827]" italics="true" pageId="5" pageNumber="144">et al.</emphasis>
|
||
1983
|
||
</bibRefCitation>
|
||
). Within the Pilbara,
|
||
<taxonomicName id="4C314D5EFFADFF83F145F10EFC5EAC23" box="[854,1009,1802,1827]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="pilbarensis">
|
||
<emphasis id="B945EACFFFADFF83F145F10EFC5EAC23" box="[854,1009,1802,1827]" italics="true" pageId="5" pageNumber="144">V. pilbarensis</emphasis>
|
||
</taxonomicName>
|
||
is also strictly saxicolous and has a reddish ground colour over most of the dorsum, whereas
|
||
<taxonomicName id="4C314D5EFFADFF83F10AF132FCDAAC47" box="[793,885,1840,1863]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="tristis">
|
||
<emphasis id="B945EACFFFADFF83F10AF132FCDAAC47" box="[793,885,1840,1863]" italics="true" pageId="5" pageNumber="144">V. tristis</emphasis>
|
||
</taxonomicName>
|
||
is largely arboreal and grey, dark brown to black over the dorsum.
|
||
<taxonomicName id="4C314D5EFFADFF83F363F156FD9DAC6B" box="[368,562,1874,1899]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="glauerti">
|
||
<emphasis id="B945EACFFFADFF83F363F156FD9DAC6B" box="[368,562,1874,1899]" italics="true" pageId="5" pageNumber="144">Varanus glauerti</emphasis>
|
||
</taxonomicName>
|
||
from the Kimberley is a saxicolous species similar in appearance to
|
||
<taxonomicName id="4C314D5EFFADFF83F794F156FEBCAC8F" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="pilbarensis">
|
||
<emphasis id="B945EACFFFADFF83F794F156FEBCAC8F" italics="true" pageId="5" pageNumber="144">V. pilbarensis</emphasis>
|
||
</taxonomicName>
|
||
(depressed body, long tail and dorsal ocelli); however,
|
||
<taxonomicName id="4C314D5EFFADFF83F195F17AFC54AC8F" box="[902,1019,1910,1935]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="144" phylum="Chordata" rank="species" species="glauerti">
|
||
<emphasis id="B945EACFFFADFF83F195F17AFC54AC8F" box="[902,1019,1910,1935]" italics="true" pageId="5" pageNumber="144">V. glauerti</emphasis>
|
||
</taxonomicName>
|
||
attains a larger size (maximum SVL up to
|
||
<quantity id="4CC99B38FFADFF83F2C6F199FE9DACB2" box="[213,306,1947,1971]" metricMagnitude="-1" metricUnit="m" metricValue="2.27" pageId="5" pageNumber="144" unit="mm" value="227.0">227 mm</quantity>
|
||
v.
|
||
<quantity id="4CC99B38FFADFF83F342F199FE1DACB3" box="[337,434,1947,1971]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="144" unit="mm" value="180.0">180 mm</quantity>
|
||
), has more midbody scales (between 122–151 v. 118–128) and a proportionally longer tail (between 183–268% of SVL v. 170–211%).
|
||
</paragraph>
|
||
<paragraph id="8B8E36DDFFAEFF80F2D4F695FE95ABD5" blockId="6.[151,1436,151,213]" pageId="6" pageNumber="145">
|
||
All subsequent morphological comparisons will involve only the two species in the
|
||
<taxonomicName id="4C314D5EFFAEFF80F6ABF69BFA33ABB0" box="[1208,1436,151,176]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="145" phylum="Chordata" rank="species" species="pilbarensis">
|
||
<emphasis id="B945EACFFFAEFF80F6ABF69BFA33ABB0" box="[1208,1436,151,176]" italics="true" pageId="6" pageNumber="145">Varanus pilbarensis</emphasis>
|
||
</taxonomicName>
|
||
species-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |