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<document id="41873BBADAAB920CB28F5D310A674844" ID-DOI="10.11646/zootaxa.4019.1.18" ID-GBIF-Dataset="17696c34-012b-4e06-bd03-8516d44a6814" ID-ISSN="1175-5326" ID-Zenodo-Dep="289527" ID-ZooBank="906BB67C-F137-4CDA-A26B-77A025725800" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461264687039" checkinUser="plazi" docAuthor="Nogueira, João Miguel Matos, Hutchings, Pat &amp; Carrerette, Orlemir" docDate="2015" docId="039887C6FFD89E3AFF31451AFD190904" docLanguage="en" docName="zt04019p576.pdf" docOrigin="Zootaxa 4019 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pistella franciscana Nogueira, Hutchings &amp; Carrerette, 2015, n. sp." docType="treatment" docVersion="14" lastPageNumber="539" masterDocId="FFA1FFBEFFEA9E0DFFA64624FFFD0B64" masterDocTitle="Terebellidae (Annelida, Terebelliformia) from Lizard Island, Great Barrier Reef, Australia" masterLastPageNumber="576" masterPageNumber="484" pageNumber="534" updateTime="1698610256753" updateUser="plazi">
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<mods:title id="C3713AA2760008BC6D5333648B9A8D10">Terebellidae (Annelida, Terebelliformia) from Lizard Island, Great Barrier Reef, Australia</mods:title>
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<mods:namePart id="D5C3A33A538FD93016F1818044D8F083">Nogueira, João Miguel Matos</mods:namePart>
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<mods:namePart id="458ED521F1E63981403B8901C1A6973C">Hutchings, Pat</mods:namePart>
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<mods:namePart id="8227F354750D6D60596032080BF81EFF">Carrerette, Orlemir</mods:namePart>
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<mods:title id="68F8C96B909B5AB6AE699B39393F4D06">Zootaxa</mods:title>
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<mods:date id="4F65976212B45F2E6AF77DDD1C15AACB">2015</mods:date>
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<mods:identifier id="74C8C0D736F08F30770053E35EB8C5B5" type="DOI">10.11646/zootaxa.4019.1.18</mods:identifier>
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<treatment id="039887C6FFD89E3AFF31451AFD190904" ID-DOI="http://doi.org/10.5281/zenodo.5313623" ID-GBIF-Taxon="119637659" ID-Zenodo-Dep="5313623" LSID="urn:lsid:plazi:treatment:039887C6FFD89E3AFF31451AFD190904" httpUri="http://treatment.plazi.org/id/039887C6FFD89E3AFF31451AFD190904" lastPageId="55" lastPageNumber="539" pageId="50" pageNumber="534">
<subSubSection id="C32B655BFFD89E3FFF31451AFE93081E" pageId="50" pageNumber="534" type="nomenclature">
<paragraph id="8B8E36D0FFD89E3FFF31451AFE2C083C" blockId="50.[151,465,830,890]" box="[151,465,830,856]" pageId="50" pageNumber="534">
<heading id="D0C681BCFFD89E3FFF31451AFE2C083C" bold="true" box="[151,465,830,856]" fontSize="11" level="1" pageId="50" pageNumber="534" reason="1">
<emphasis id="B945EAC2FFD89E3FFF31451AFE2C083C" bold="true" box="[151,465,830,856]" pageId="50" pageNumber="534">
<taxonomicName id="4C314D53FFD89E3FFF31451AFE7A083C" ID-CoL="77MLC" box="[151,391,830,856]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="50" pageNumber="534" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFD89E3FFF31451AFE7A083C" bold="true" box="[151,391,830,856]" italics="true" pageId="50" pageNumber="534">Pistella franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFD89E3FFE28451AFE2C083C" box="[398,465,830,856]" pageId="50" pageNumber="534" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36D0FFD89E3FFF314545FE93081E" blockId="50.[151,465,830,890]" box="[151,366,865,890]" pageId="50" pageNumber="534">
(
<figureCitation id="130A2A55FFD89E3FFF394545FF18081E" box="[159,229,865,890]" captionStart="FIGURE 2" captionStartId="3.[151,250,1615,1637]" captionTargetBox="[173,1421,183,1594]" captionTargetId="figure@3.[166,1426,179,1607]" captionTargetPageId="3" captionText="FIGURE 2. Live specimens. A B. Pista kristiani n. sp. AM W. 45451, MI QLD 2445; C D. Lanicides rubra n. sp. AM W. 44272, MI QLD 2344; E G. Eupolymnia chlorobranchiata n. sp. AM W. 45455, MI QLD 2446; H I. Pistella franciscana n. sp. H. Discarded specimen; I. AM W. 44593, MI QLD 2394; J L. Loimia tuberculata n. sp; J K. Spec. AM W. 44280, MI QLD 2359; L. Discarded specimen. Photos: A K—Alexander Semenov, L—Gary Cranitch." httpUri="https://zenodo.org/record/289529/files/figure.png" pageId="50" pageNumber="534">Figs 2</figureCitation>
HI, 3235)
</paragraph>
</subSubSection>
<subSubSection id="C32B655BFFD89E3FFF314586FD7C0E23" pageId="50" pageNumber="534" type="materials_examined">
<paragraph id="8B8E36D0FFD89E3FFF314586FEB60F2F" blockId="50.[151,1437,930,2035]" pageId="50" pageNumber="534">
<emphasis id="B945EAC2FFD89E3FFF314586FEB808DF" bold="true" box="[151,325,930,955]" pageId="50" pageNumber="534">
<typeStatus id="548A8872FFD89E3FFF314586FF2F08DF" box="[151,210,930,955]" pageId="50" pageNumber="534">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548A8872FFD89E3FFEEA4586FE4408DF" box="[332,441,930,955]" pageId="50" pageNumber="534" type="holotype">Holotype</typeStatus>
: AM W.45445, MI QLD 2444, north of North Point,
<geoCoordinate id="EE055017FFD89E3FFBAD4586FB7008DF" box="[1035,1165,930,955]" direction="south" orientation="latitude" pageId="50" pageNumber="534" precision="15" value="-14.643333">14°38'36&quot;S</geoCoordinate>
,
<geoCoordinate id="EE055017FFD89E3FFB3E4586FAD408DF" box="[1176,1321,930,955]" direction="east" orientation="longitude" pageId="50" pageNumber="534" precision="15" value="145.4525">145°27'09&quot;E</geoCoordinate>
, complete specimen, in excellent state of preservation, ~
<quantity id="4CC99B35FFD89E3FFD3345E2FD1908BA" box="[661,740,966,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="50" pageNumber="534" unit="mm" value="15.0">15 mm</quantity>
long,
<quantity id="4CC99B35FFD89E3FFC8F45E2FC8308BA" box="[809,894,966,991]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="534" unit="mm" value="0.6">0.6 mm</quantity>
wide.
<typeStatus id="548A8872FFD89E3FFC6F45E3FBC708BB" box="[969,1082,967,991]" pageId="50" pageNumber="534" type="paratype">Paratypes</typeStatus>
: AM W.47715, CReefs, LI10 0 33, MI QLD 2198, Watsons Bay,
<geoCoordinate id="EE055017FFD89E3FFDFE45CFFD240F67" box="[600,729,1003,1027]" direction="south" orientation="latitude" pageId="50" pageNumber="534" precision="15" value="-14.658333">14°39'30&quot;S</geoCoordinate>
,
<geoCoordinate id="EE055017FFD89E3FFD4945CEFC7D0F67" box="[751,896,1002,1027]" direction="east" orientation="longitude" pageId="50" pageNumber="534" precision="15" value="145.44888">145°26'56&quot;E</geoCoordinate>
, incomplete specimen, in excellent state of preservation; AM W.44593, MI QLD 2394, mounted on 2 SEM pins; AM W.44964, MI QLD 2435 (photographed).
</paragraph>
<paragraph id="8B8E36D0FFD89E3FFF614272FCF60FD3" blockId="50.[151,1437,930,2035]" pageId="50" pageNumber="534">
<emphasis id="B945EAC2FFD89E3FFF614272FDFD0F0B" bold="true" box="[199,512,1110,1135]" pageId="50" pageNumber="534">Other material examined.</emphasis>
AM W.44957, MI QLD 2429; AM W.44626, MI QLD 2414; AM W.44628, MI QLD 2416; AM W.45441, MI QLD 2444; AM W.45145, MI QLD 2440; AM W.45447, MI QLD 2444 (3); AM W.45146, MI QLD 2441; AM W.45446, MI QLD 2444.
</paragraph>
<paragraph id="8B8E36D0FFD89E3FFF6142E6FD7C0E23" blockId="50.[151,1437,930,2035]" pageId="50" pageNumber="534">
<emphasis id="B945EAC2FFD89E3FFF6142E6FDA80FBF" bold="true" box="[199,597,1218,1243]" pageId="50" pageNumber="534">Comparative material examined.</emphasis>
<typeStatus id="548A8872FFD89E3FFDF842E6FD3B0FBF" box="[606,710,1218,1243]" pageId="50" pageNumber="534" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C314D53FFD89E3FFD5642E6FB220FBF" authority="Hutchings &amp; Glasby, 1990" authorityName="Hutchings &amp; Glasby" authorityYear="1990" box="[752,1247,1218,1243]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="50" pageNumber="534" phylum="Annelida" rank="species" species="anthela">
<emphasis id="B945EAC2FFD89E3FFD5642E6FC5E0FBF" box="[752,931,1218,1243]" italics="true" pageId="50" pageNumber="534">Pistella anthela</emphasis>
<bibRefCitation id="EFA04B21FFD89E3FFC0A42E6FB220FBF" author="Hutchings" box="[940,1247,1218,1243]" pageId="50" pageNumber="534" refString="Hutchings, P. A. &amp; Glasby, C. J. (1990) Additional new species of the family Terebellidae (Polychaeta) from Western Australia, with a key to all described species of the region. In: Wells, F. E., Walker, D. I., Kirkman, H. &amp; Lethbridge, R. (Eds.), Proceedings of the Third International Marine Biological Workshop: the marine flora and fauna of Albany, Western Australia. Western Australian Museum, Perth, pp. 251 - 289." type="book chapter" year="1990">Hutchings &amp; Glasby, 1990</bibRefCitation>
</taxonomicName>
, AM W.203515.
<typeStatus id="548A8872FFD89E3FFF3142C3FEF90F9B" box="[151,260,1255,1279]" pageId="50" pageNumber="534" type="paratype">Paratypes</typeStatus>
of
<taxonomicName id="4C314D53FFD89E3FFE9742C2FE180F9B" box="[305,485,1254,1279]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="50" pageNumber="534" phylum="Annelida" rank="species" species="anthela">
<emphasis id="B945EAC2FFD89E3FFE9742C2FE180F9B" box="[305,485,1254,1279]" italics="true" pageId="50" pageNumber="534">Pistella anthela</emphasis>
</taxonomicName>
, AM W.203516, AM W.203517, AM W.203520, AM W.203521, AM W.203523.
<typeStatus id="548A8872FFD89E3FFF31432EFF020E47" box="[151,255,1290,1315]" pageId="50" pageNumber="534" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C314D53FFD89E3FFE8F432EFD530E47" authority="Pearson, 1969" authorityName="Pearson" authorityYear="1969" box="[297,686,1290,1315]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="50" pageNumber="534" phylum="Annelida" rank="species" species="lornensis">
<emphasis id="B945EAC2FFD89E3FFE8F432EFE120E47" box="[297,495,1290,1315]" italics="true" pageId="50" pageNumber="534">Pistella lornensis</emphasis>
(
<bibRefCitation id="EFA04B21FFD89E3FFE59432FFD5B0E47" author="Pearson" box="[511,678,1290,1315]" pageId="50" pageNumber="534" refString="Pearson, T. H. (1969) Scionella lornensis sp. nov., a new terebellid (Polychaeta: Annelida) from the west coast of Scotland, with notes on the genus Scionella Moore, and a key to the genera of the Terebellidae recorded from European waters. Journal of Natural History, 3, 509 - 516. http: // dx. doi. org / 10.1080 / 00222936900770441" type="journal article" year="1969">Pearson, 1969</bibRefCitation>
)
</taxonomicName>
, ZB 1968: 15.
<typeStatus id="548A8872FFD89E3FFCF8432FFC3D0E47" box="[862,960,1291,1315]" pageId="50" pageNumber="534" type="paratype">Paratype</typeStatus>
of
<taxonomicName id="4C314D53FFD89E3FFC4C432EFB520E47" box="[1002,1199,1290,1315]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="50" pageNumber="534" phylum="Annelida" rank="species" species="lornensis">
<emphasis id="B945EAC2FFD89E3FFC4C432EFB520E47" box="[1002,1199,1290,1315]" italics="true" pageId="50" pageNumber="534">Pistella lornensis</emphasis>
</taxonomicName>
, ZB 1968: 16. Nontype of
<taxonomicName id="4C314D53FFD89E3FFF4A430AFE520E23" box="[236,431,1326,1351]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="50" pageNumber="534" phylum="Annelida" rank="species" species="lornensis">
<emphasis id="B945EAC2FFD89E3FFF4A430AFE520E23" box="[236,431,1326,1351]" italics="true" pageId="50" pageNumber="534">Pistella lornensis</emphasis>
</taxonomicName>
,
<collectionCode id="ED20AE15FFD89E3FFE1B430BFDF30E23" box="[445,526,1327,1351]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="50" pageNumber="534">ZMUC</collectionCode>
Pol 1744.
</paragraph>
</subSubSection>
<subSubSection id="C32B655BFFD89E38FF614376FDAD0AB4" lastPageId="53" lastPageNumber="537" pageId="50" pageNumber="534" type="description">
<paragraph id="8B8E36D0FFD89E38FF614376FDAD0AB4" blockId="50.[151,1437,930,2035]" lastBlockId="53.[151,1436,151,464]" lastPageId="53" lastPageNumber="537" pageId="50" pageNumber="534">
<emphasis id="B945EAC2FFD89E3FFF614376FEA40E0F" bold="true" box="[199,345,1362,1387]" pageId="50" pageNumber="534">Description.</emphasis>
In life, body pink to greenish, with colourless buccal tentacles and mid-ventral shields with bright red posterior section (
<figureCitation id="130A2A55FFD89E3FFE2D4352FE300EEB" box="[395,461,1398,1423]" captionStart="FIGURE 2" captionStartId="3.[151,250,1615,1637]" captionTargetBox="[173,1421,183,1594]" captionTargetId="figure@3.[166,1426,179,1607]" captionTargetPageId="3" captionText="FIGURE 2. Live specimens. A B. Pista kristiani n. sp. AM W. 45451, MI QLD 2445; C D. Lanicides rubra n. sp. AM W. 44272, MI QLD 2344; E G. Eupolymnia chlorobranchiata n. sp. AM W. 45455, MI QLD 2446; H I. Pistella franciscana n. sp. H. Discarded specimen; I. AM W. 44593, MI QLD 2394; J L. Loimia tuberculata n. sp; J K. Spec. AM W. 44280, MI QLD 2359; L. Discarded specimen. Photos: A K—Alexander Semenov, L—Gary Cranitch." httpUri="https://zenodo.org/record/289529/files/figure.png" pageId="50" pageNumber="534">Fig. 2</figureCitation>
HI). Transverse prostomium attached to dorsal surface of upper lip; basal part without eyespots; distal part shelf-like (
<figureCitation id="130A2A55FFD89E3FFE5243BEFDB50ED7" box="[500,584,1434,1459]" captionStart="FIGURE 32" captionStartId="51.[151,250,1919,1941]" captionTargetBox="[155,1432,198,1888]" captionTargetId="figure@51.[152,1435,193,1898]" captionTargetPageId="51" captionText="FIGURE 32. Pistella franciscana n. sp. AM W. 45445: A B. Entire worm, ventro-lateral and dorsal views, respectively; C F. Anterior ends, dorsal, right lateral, ventral and left lateral views, respectively; G I. Posterior end, in progressively lower magnifications, ventral views; J K. Close ups of the anterior end, ventral and right lateral views, respectively. Numbers refer to segments. Abbreviations: ul = upper lip. Scale bars: A B = 0.5 mm, C, E, G, K = 0.2 mm, D, F, H I = 0.3 mm, J = 0.15 mm." httpUri="https://zenodo.org/record/289559/files/figure.png" pageId="50" pageNumber="534">Figs 32</figureCitation>
EF, JK; 33A, D, F, J). Buccal tentacles, longer than region with notopodia in life, but shorter after preservation (
<figureCitation id="130A2A55FFD89E3FFD9C439AFD7C0EB3" box="[570,641,1470,1495]" captionStart="FIGURE 2" captionStartId="3.[151,250,1615,1637]" captionTargetBox="[173,1421,183,1594]" captionTargetId="figure@3.[166,1426,179,1607]" captionTargetPageId="3" captionText="FIGURE 2. Live specimens. A B. Pista kristiani n. sp. AM W. 45451, MI QLD 2445; C D. Lanicides rubra n. sp. AM W. 44272, MI QLD 2344; E G. Eupolymnia chlorobranchiata n. sp. AM W. 45455, MI QLD 2446; H I. Pistella franciscana n. sp. H. Discarded specimen; I. AM W. 44593, MI QLD 2394; J L. Loimia tuberculata n. sp; J K. Spec. AM W. 44280, MI QLD 2359; L. Discarded specimen. Photos: A K—Alexander Semenov, L—Gary Cranitch." httpUri="https://zenodo.org/record/289529/files/figure.png" pageId="50" pageNumber="534">Figs 2</figureCitation>
HI; 32AF, JK; 33AK). Peristomium forming lips; hood-like upper lip, short, as wide as long; small, swollen lower lip, restricted to oral area (
<figureCitation id="130A2A55FFD89E3FFC7443C6FBDA0E9F" box="[978,1063,1506,1531]" captionStart="FIGURE 32" captionStartId="51.[151,250,1919,1941]" captionTargetBox="[155,1432,198,1888]" captionTargetId="figure@51.[152,1435,193,1898]" captionTargetPageId="51" captionText="FIGURE 32. Pistella franciscana n. sp. AM W. 45445: A B. Entire worm, ventro-lateral and dorsal views, respectively; C F. Anterior ends, dorsal, right lateral, ventral and left lateral views, respectively; G I. Posterior end, in progressively lower magnifications, ventral views; J K. Close ups of the anterior end, ventral and right lateral views, respectively. Numbers refer to segments. Abbreviations: ul = upper lip. Scale bars: A B = 0.5 mm, C, E, G, K = 0.2 mm, D, F, H I = 0.3 mm, J = 0.15 mm." httpUri="https://zenodo.org/record/289559/files/figure.png" pageId="50" pageNumber="534">Figs 32</figureCitation>
E, J; 33GH). Segment 1 reduced dorsally, expanded ventrally with paired ventral lobes lateral to mouth, connected to each other by low lobe marginal to mouth; segment 2 with paired ventro-lateral lobes connected to each other by protruding mid-ventral crest, lobes almost semi-circular, covering posterior half of segment 1; segment 3 with pair of triangular, distally rounded lateral lobes, lobes short, only covering posterior part of segment 2, dorsal margins aligned with line of notopodia, ventral margins terminating in line with the mid-length of subsequent neuropodial tori (
<figureCitation id="130A2A55FFD89E3FFB5640B2FAB80DCB" box="[1264,1349,1686,1711]" captionStart="FIGURE 32" captionStartId="51.[151,250,1919,1941]" captionTargetBox="[155,1432,198,1888]" captionTargetId="figure@51.[152,1435,193,1898]" captionTargetPageId="51" captionText="FIGURE 32. Pistella franciscana n. sp. AM W. 45445: A B. Entire worm, ventro-lateral and dorsal views, respectively; C F. Anterior ends, dorsal, right lateral, ventral and left lateral views, respectively; G I. Posterior end, in progressively lower magnifications, ventral views; J K. Close ups of the anterior end, ventral and right lateral views, respectively. Numbers refer to segments. Abbreviations: ul = upper lip. Scale bars: A B = 0.5 mm, C, E, G, K = 0.2 mm, D, F, H I = 0.3 mm, J = 0.15 mm." httpUri="https://zenodo.org/record/289559/files/figure.png" pageId="50" pageNumber="534">Figs 32</figureCitation>
AF, J K; 33AL). Anterior segments not markedly inflated dorsally. Paired dorso-lateral plumous branchiae present on segment 2, single pair, frequently one branchia missing, presumably lost by predation, supported by the fact that some individuals have branchiae of remarkably different sizes (
<figureCitation id="130A2A55FFD89E3FFCCD4126FC3C0C7F" box="[875,961,1794,1819]" captionStart="FIGURE 32" captionStartId="51.[151,250,1919,1941]" captionTargetBox="[155,1432,198,1888]" captionTargetId="figure@51.[152,1435,193,1898]" captionTargetPageId="51" captionText="FIGURE 32. Pistella franciscana n. sp. AM W. 45445: A B. Entire worm, ventro-lateral and dorsal views, respectively; C F. Anterior ends, dorsal, right lateral, ventral and left lateral views, respectively; G I. Posterior end, in progressively lower magnifications, ventral views; J K. Close ups of the anterior end, ventral and right lateral views, respectively. Numbers refer to segments. Abbreviations: ul = upper lip. Scale bars: A B = 0.5 mm, C, E, G, K = 0.2 mm, D, F, H I = 0.3 mm, J = 0.15 mm." httpUri="https://zenodo.org/record/289559/files/figure.png" pageId="50" pageNumber="534">Figs 32</figureCitation>
AF, K; 33AF, IK); each branchia with conspicuous, crenulate basal stem, and branchial filaments originating in a spiral at tip; branchial filaments dichotomously branching for 12 levels, with relatively long tips, bearing ciliary tracks laterally and around bald tips (
<figureCitation id="130A2A55FFD89E3FFF75414AFEDB0CE3" box="[211,294,1902,1927]" captionStart="FIGURE 33" captionStartId="52.[151,250,1820,1842]" captionTargetBox="[205,1359,194,1785]" captionTargetId="figure@52.[204,1360,193,1796]" captionTargetPageId="52" captionText="FIGURE 33. Pistella franciscana n. sp. AM W. 44593: A E. Anterior end, left ventro-lateral, right lateral, right ventro-lateral, left lateral and right dorso-lateral views, respectively; F K. Close ups of the anterior end, dorsal, left and right (2) ventro-lateral views, left dorso-lateral and left lateral views, respectively; L. Close up of segments 2 8, left lateral view, white and black arrows point to the two sets of papillae; M N. Progressively closer views of the branchiae; O. Posterior end, left lateral view. Numbers refer to segments. Abbreviations: ll = lower lip, P = basal part of prostomium, ul = upper lip, * = distal part of prostomium. Scale bars: A, C E = 200 µm, B = 300 µm, F, I = 150 µm, G, L = 70 µm, H, J K, O = 100 µm, M = 40 µm, N = 10 µm." httpUri="https://zenodo.org/record/289560/files/figure.png" pageId="50" pageNumber="534">Fig. 33</figureCitation>
AF, IK, MN). Anterior segments with glandular, rectangular to trapezoidal, smooth to slightly corrugated mid-ventral shields; shields typically divided in two parts, anterior part white, posterior part bright red, not always visible after preservation (
<figureCitation id="130A2A55FFD89E3FFDEF4192FD6F0CAB" box="[585,658,1974,1999]" captionStart="FIGURE 2" captionStartId="3.[151,250,1615,1637]" captionTargetBox="[173,1421,183,1594]" captionTargetId="figure@3.[166,1426,179,1607]" captionTargetPageId="3" captionText="FIGURE 2. Live specimens. A B. Pista kristiani n. sp. AM W. 45451, MI QLD 2445; C D. Lanicides rubra n. sp. AM W. 44272, MI QLD 2344; E G. Eupolymnia chlorobranchiata n. sp. AM W. 45455, MI QLD 2446; H I. Pistella franciscana n. sp. H. Discarded specimen; I. AM W. 44593, MI QLD 2394; J L. Loimia tuberculata n. sp; J K. Spec. AM W. 44280, MI QLD 2359; L. Discarded specimen. Photos: A K—Alexander Semenov, L—Gary Cranitch." httpUri="https://zenodo.org/record/289529/files/figure.png" pageId="50" pageNumber="534">Figs 2</figureCitation>
HI; 32A, DE, JK; 33A, C, GI, K). Short and conical notopodia beginning from segment 4, typically extending for 17 segments, until segment 20. Winged notochaetae in both rows, broadly-winged, wings as broad as width of shaft or broader (
<figureCitation id="130A2A55FFDF9E38FCFC46B3FC4F0BD4" box="[858,946,151,176]" captionStart="FIGURE 34" captionStartId="53.[151,250,1838,1860]" captionTargetBox="[151,1434,519,1808]" captionTargetId="figure@53.[151,1435,517,1817]" captionTargetPageId="53" captionText="FIGURE 34. Pistella franciscana n. sp. AM W. 44593: A. Pygidium, frontal view; B. Neuropodium, segment 16; C D. Notochaetae, segment 8, general and chaetae from anterior row, respectively; E G. Notochaetae of segment 15, general, chaetae from anterior and posterior rows, respectively; H. Uncini, segment 10; I J. Uncini, segment 15 under higher magnifications; K. Uncini, segment 24; L M. Posterior uncini, under higher magnifications. Scale bars: A = 20 µm, B, D, L = 7 µm, C = 15 µm, E G = 10 µm, H, M = 2 µm, I, K = 5 µm, J = 3 µm." httpUri="https://zenodo.org/record/289561/files/figure.png" pageId="53" pageNumber="537">Figs 34</figureCitation>
CG; 35AB, F). Neuropodia present from segment 5, as low ridges until termination of notopodia, as low pinnules thereafter. Neurochaetae throughout as short-handled avicular uncini, arranged in partially to completely intercalated double rows from segment 11 until termination of notopodia; uncini with dorsal button at mid-length of base, conspicuous at least until termination of notopodia, and crest with 34 transverse rows of secondary teeth (
<figureCitation id="130A2A55FFDF9E38FC204703FC230A24" box="[902,990,295,320]" captionStart="FIGURE 34" captionStartId="53.[151,250,1838,1860]" captionTargetBox="[151,1434,519,1808]" captionTargetId="figure@53.[151,1435,517,1817]" captionTargetPageId="53" captionText="FIGURE 34. Pistella franciscana n. sp. AM W. 44593: A. Pygidium, frontal view; B. Neuropodium, segment 16; C D. Notochaetae, segment 8, general and chaetae from anterior row, respectively; E G. Notochaetae of segment 15, general, chaetae from anterior and posterior rows, respectively; H. Uncini, segment 10; I J. Uncini, segment 15 under higher magnifications; K. Uncini, segment 24; L M. Posterior uncini, under higher magnifications. Scale bars: A = 20 µm, B, D, L = 7 µm, C = 15 µm, E G = 10 µm, H, M = 2 µm, I, K = 5 µm, J = 3 µm." httpUri="https://zenodo.org/record/289561/files/figure.png" pageId="53" pageNumber="537">Figs 34</figureCitation>
B, HM; 35CE, GK). Nephridial and genital papillae usually present on segments 3 and 67, respectively, the latter posterior and dorsal to notopodia (
<figureCitation id="130A2A55FFDF9E38FF39474BFF0C0AEC" box="[159,241,367,392]" captionStart="FIGURE 33" captionStartId="52.[151,250,1820,1842]" captionTargetBox="[205,1359,194,1785]" captionTargetId="figure@52.[204,1360,193,1796]" captionTargetPageId="52" captionText="FIGURE 33. Pistella franciscana n. sp. AM W. 44593: A E. Anterior end, left ventro-lateral, right lateral, right ventro-lateral, left lateral and right dorso-lateral views, respectively; F K. Close ups of the anterior end, dorsal, left and right (2) ventro-lateral views, left dorso-lateral and left lateral views, respectively; L. Close up of segments 2 8, left lateral view, white and black arrows point to the two sets of papillae; M N. Progressively closer views of the branchiae; O. Posterior end, left lateral view. Numbers refer to segments. Abbreviations: ll = lower lip, P = basal part of prostomium, ul = upper lip, * = distal part of prostomium. Scale bars: A, C E = 200 µm, B = 300 µm, F, I = 150 µm, G, L = 70 µm, H, J K, O = 100 µm, M = 40 µm, N = 10 µm." httpUri="https://zenodo.org/record/289560/files/figure.png" pageId="53" pageNumber="537">Fig. 33</figureCitation>
D, K, L). In addition there are additional minute papillae on segments 26 which are labelled in
<figureCitation id="130A2A55FFDF9E38FA92474BFA780AEC" box="[1332,1413,367,392]" captionStart="FIGURE 33" captionStartId="52.[151,250,1820,1842]" captionTargetBox="[205,1359,194,1785]" captionTargetId="figure@52.[204,1360,193,1796]" captionTargetPageId="52" captionText="FIGURE 33. Pistella franciscana n. sp. AM W. 44593: A E. Anterior end, left ventro-lateral, right lateral, right ventro-lateral, left lateral and right dorso-lateral views, respectively; F K. Close ups of the anterior end, dorsal, left and right (2) ventro-lateral views, left dorso-lateral and left lateral views, respectively; L. Close up of segments 2 8, left lateral view, white and black arrows point to the two sets of papillae; M N. Progressively closer views of the branchiae; O. Posterior end, left lateral view. Numbers refer to segments. Abbreviations: ll = lower lip, P = basal part of prostomium, ul = upper lip, * = distal part of prostomium. Scale bars: A, C E = 200 µm, B = 300 µm, F, I = 150 µm, G, L = 70 µm, H, J K, O = 100 µm, M = 40 µm, N = 10 µm." httpUri="https://zenodo.org/record/289560/files/figure.png" pageId="53" pageNumber="537">Fig. 33</figureCitation>
L, the function of which is unknown, perhaps they are sensory. Pygidium with larger ventral papillae (
<figureCitation id="130A2A55FFDF9E38FAB947B0FA940AC8" box="[1311,1385,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1615,1637]" captionTargetBox="[173,1421,183,1594]" captionTargetId="figure@3.[166,1426,179,1607]" captionTargetPageId="3" captionText="FIGURE 2. Live specimens. A B. Pista kristiani n. sp. AM W. 45451, MI QLD 2445; C D. Lanicides rubra n. sp. AM W. 44272, MI QLD 2344; E G. Eupolymnia chlorobranchiata n. sp. AM W. 45455, MI QLD 2446; H I. Pistella franciscana n. sp. H. Discarded specimen; I. AM W. 44593, MI QLD 2394; J L. Loimia tuberculata n. sp; J K. Spec. AM W. 44280, MI QLD 2359; L. Discarded specimen. Photos: A K—Alexander Semenov, L—Gary Cranitch." httpUri="https://zenodo.org/record/289529/files/figure.png" pageId="53" pageNumber="537">Figs 2</figureCitation>
HI; 32AB, GI; 33
<collectingCountry id="F3267640FFDF9E38FEE8479CFE990AB4" box="[334,356,440,464]" name="Iceland" pageId="53" pageNumber="537">O</collectingCountry>
; 36A). Mucous tube.
</paragraph>
</subSubSection>
<caption id="DF4E6658FFD99E3EFF31415BFA790C95" httpUri="https://zenodo.org/record/289559/files/figure.png" pageId="51" pageNumber="535" targetBox="[155,1432,198,1888]" targetPageId="51">
<paragraph id="8B8E36D0FFD99E3EFF31415BFA790C95" blockId="51.[151,1437,1919,2033]" pageId="51" pageNumber="535">
<emphasis id="B945EAC2FFD99E3EFF31415BFEDD0CF1" bold="true" box="[151,288,1919,1941]" pageId="51" pageNumber="535">FIGURE 32.</emphasis>
<taxonomicName id="4C314D53FFD99E3EFE81415BFE0E0CF1" box="[295,499,1919,1941]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="51" pageNumber="535" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFD99E3EFE81415BFE0E0CF1" box="[295,499,1919,1941]" italics="true" pageId="51" pageNumber="535">Pistella franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFD99E3EFE5C41A4FDCC0CF1" box="[506,561,1920,1941]" pageId="51" pageNumber="535" rank="species">n. sp.</taxonomicNameLabel>
AM W.45445: AB. Entire worm, ventro-lateral and dorsal views, respectively; CF. Anterior ends, dorsal, right lateral, ventral and left lateral views, respectively; GI. Posterior end, in progressively lower magnifications, ventral views; JK. Close ups of the anterior end, ventral and right lateral views, respectively. Numbers refer to segments. Abbreviations: ul = upper lip. Scale bars: AB = 0.5 mm, C, E, G, K = 0.2 mm, D, F, HI = 0.3 mm, J = 0.15 mm.
</paragraph>
</caption>
<caption id="DF4E6658FFDE9E39FF314138FF3D0C8E" httpUri="https://zenodo.org/record/289560/files/figure.png" pageId="52" pageNumber="536" targetBox="[205,1359,194,1785]" targetPageId="52">
<paragraph id="8B8E36D0FFDE9E39FF314138FF3D0C8E" blockId="52.[151,1436,1820,2026]" pageId="52" pageNumber="536">
<emphasis id="B945EAC2FFDE9E39FF314138FEDD0C56" bold="true" box="[151,288,1820,1842]" pageId="52" pageNumber="536">FIGURE 33.</emphasis>
<taxonomicName id="4C314D53FFDE9E39FE804138FE0F0C56" box="[294,498,1820,1842]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="52" pageNumber="536" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFDE9E39FE804138FE0F0C56" box="[294,498,1820,1842]" italics="true" pageId="52" pageNumber="536">Pistella franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFDE9E39FE5E4139FDD20C56" box="[504,559,1821,1842]" pageId="52" pageNumber="536" rank="species">n. sp.</taxonomicNameLabel>
AM W.44593: AE. Anterior end, left ventro-lateral, right lateral, right ventro-lateral, left lateral and right dorso-lateral views, respectively; FK. Close ups of the anterior end, dorsal, left and right (2) ventro-lateral views, left dorso-lateral and left lateral views, respectively; L. Close up of segments 28, left lateral view, white and black arrows point to the two sets of papillae; MN. Progressively closer views of the branchiae; O. Posterior end, left lateral view. Numbers refer to segments. Abbreviations: ll = lower lip, P = basal part of prostomium, ul = upper lip, * = distal part of prostomium. Scale bars: A, CE = 200 µm, B = 300 µm, F, I = 150 µm, G, L = 70 µm, H, JK, O = 100 µm, M = 40 µm, N = 10 µm.
</paragraph>
</caption>
<caption id="DF4E6658FFDF9E38FF31410AFC9F0CDB" httpUri="https://zenodo.org/record/289561/files/figure.png" pageId="53" pageNumber="537" targetBox="[151,1434,519,1808]" targetPageId="53">
<paragraph id="8B8E36D0FFDF9E38FF31410AFC9F0CDB" blockId="53.[151,1436,1838,1983]" pageId="53" pageNumber="537">
<emphasis id="B945EAC2FFDF9E38FF31410AFED90C20" bold="true" box="[151,292,1838,1860]" pageId="53" pageNumber="537">FIGURE 34.</emphasis>
<taxonomicName id="4C314D53FFDF9E38FE89410AFE020C20" box="[303,511,1838,1860]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="53" pageNumber="537" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFDF9E38FE89410AFE020C20" box="[303,511,1838,1860]" italics="true" pageId="53" pageNumber="537">Pistella franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFDF9E38FDAC410BFDB80C20" box="[522,581,1839,1860]" pageId="53" pageNumber="537" rank="species">n. sp.</taxonomicNameLabel>
AM W.44593: A. Pygidium, frontal view; B. Neuropodium, segment 16; CD. Notochaetae, segment 8, general and chaetae from anterior row, respectively; EG. Notochaetae of segment 15, general, chaetae from anterior and posterior rows, respectively; H. Uncini, segment 10; IJ. Uncini, segment 15 under higher magnifications; K. Uncini, segment 24; LM. Posterior uncini, under higher magnifications. Scale bars: A = 20 µm, B, D, L = 7 µm, C = 15 µm, EG = 10 µm, H, M = 2 µm, I, K = 5 µm, J = 3 µm.
</paragraph>
</caption>
<caption id="DF4E6658FFDC9E3BFF31412FFDD40C3A" httpUri="https://zenodo.org/record/289562/files/figure.png" pageId="54" pageNumber="538" targetBox="[153,1434,194,1773]" targetPageId="54">
<paragraph id="8B8E36D0FFDC9E3BFF31412FFDD40C3A" blockId="54.[151,1436,1803,1886]" pageId="54" pageNumber="538">
<emphasis id="B945EAC2FFDC9E3BFF31412FFEDF0C45" bold="true" box="[151,290,1803,1825]" pageId="54" pageNumber="538">FIGURE 35.</emphasis>
<taxonomicName id="4C314D53FFDC9E3BFE8C412FFE0A0C45" box="[298,503,1803,1825]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="54" pageNumber="538" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFDC9E3BFE8C412FFE0A0C45" box="[298,503,1803,1825]" italics="true" pageId="54" pageNumber="538">Pistella franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFDC9E3BFDA64128FDC40C45" box="[512,569,1804,1825]" pageId="54" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
AM W.44628: AB, F. Notochaetae, segments 8 and 19, general and chaetae from anterior row, respectively; CE, GK. Uncini, segments 5, 6 (2), 7 (2), 19, 24 and posterior, respectively. Scale bars: AB = 30 µm, CE, GI, K = 7 µm, F, J = 10 µm.
</paragraph>
</caption>
<subSubSection id="C32B655BFFDC9E3AFF61418AFAB30AB4" lastPageId="55" lastPageNumber="539" pageId="54" pageNumber="538" type="discussion">
<paragraph id="8B8E36D0FFDC9E3AFF61418AFF050A79" blockId="54.[151,1436,1966,2028]" lastBlockId="55.[151,1437,151,608]" lastPageId="55" lastPageNumber="539" pageId="54" pageNumber="538">
<emphasis id="B945EAC2FFDC9E3BFF61418AFEC60CA3" bold="true" box="[199,315,1966,1991]" pageId="54" pageNumber="538">Remarks.</emphasis>
<taxonomicName id="4C314D53FFDC9E3BFEE0418AFDD60CA3" box="[326,555,1966,1991]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="54" pageNumber="538" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFDC9E3BFEE0418AFDD60CA3" box="[326,555,1966,1991]" italics="true" pageId="54" pageNumber="538">Pistella franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFDC9E3BFD914194FD8A0CA3" box="[567,631,1968,1991]" pageId="54" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
differs from the only other known Australian species of
<taxonomicName id="4C314D53FFDC9E3BFAB9418AFA890CA3" box="[1311,1396,1966,1991]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="54" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B945EAC2FFDC9E3BFAB9418AFA890CA3" box="[1311,1396,1966,1991]" italics="true" pageId="54" pageNumber="538">Pistella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D53FFDC9E3BFA214194FDC90C8F" authority="Hutchings &amp; Glasby, 1990" authorityName="Hutchings &amp; Glasby" authorityYear="1990" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="54" pageNumber="538" phylum="Annelida" rank="species" species="anthela">
<emphasis id="B945EAC2FFDC9E3BFA214194FF160C88" italics="true" pageId="54" pageNumber="538">P. anthela</emphasis>
(
<bibRefCitation id="EFA04B21FFDC9E3BFF5D41F7FDD10C8F" author="Hutchings" box="[251,556,2003,2028]" pageId="54" pageNumber="538" refString="Hutchings, P. A. &amp; Glasby, C. J. (1990) Additional new species of the family Terebellidae (Polychaeta) from Western Australia, with a key to all described species of the region. In: Wells, F. E., Walker, D. I., Kirkman, H. &amp; Lethbridge, R. (Eds.), Proceedings of the Third International Marine Biological Workshop: the marine flora and fauna of Albany, Western Australia. Western Australian Museum, Perth, pp. 251 - 289." type="book chapter" year="1990">Hutchings &amp; Glasby, 1990</bibRefCitation>
)
</taxonomicName>
described from the Dampier Archipelago in Western
<collectingCountry id="F3267640FFDC9E3BFB3E41F7FAFF0C88" box="[1176,1282,2003,2028]" name="Australia" pageId="54" pageNumber="538">Australia</collectingCountry>
, by the shape and arrangement of lateral lobes. In the new species no lobes continue across the dorsum as glandular ridges as they do in
<emphasis id="B945EAC2FFDD9E3AFEA84699FE870BB0" box="[270,378,188,213]" italics="true" pageId="55" pageNumber="539">P. an t he l a</emphasis>
. The tips of the branchial filaments are completely bald surrounded by well developed cilated sides (
<figureCitation id="130A2A55FFDD9E3AFF4246FBFEC70B9C" box="[228,314,223,248]" captionStart="FIGURE 33" captionStartId="52.[151,250,1820,1842]" captionTargetBox="[205,1359,194,1785]" captionTargetId="figure@52.[204,1360,193,1796]" captionTargetPageId="52" captionText="FIGURE 33. Pistella franciscana n. sp. AM W. 44593: A E. Anterior end, left ventro-lateral, right lateral, right ventro-lateral, left lateral and right dorso-lateral views, respectively; F K. Close ups of the anterior end, dorsal, left and right (2) ventro-lateral views, left dorso-lateral and left lateral views, respectively; L. Close up of segments 2 8, left lateral view, white and black arrows point to the two sets of papillae; M N. Progressively closer views of the branchiae; O. Posterior end, left lateral view. Numbers refer to segments. Abbreviations: ll = lower lip, P = basal part of prostomium, ul = upper lip, * = distal part of prostomium. Scale bars: A, C E = 200 µm, B = 300 µm, F, I = 150 µm, G, L = 70 µm, H, J K, O = 100 µm, M = 40 µm, N = 10 µm." httpUri="https://zenodo.org/record/289560/files/figure.png" pageId="55" pageNumber="539">Fig. 33</figureCitation>
M, N), whereas those of
<emphasis id="B945EAC2FFDD9E3AFDCA46C5FD230B9C" box="[620,734,223,248]" italics="true" pageId="55" pageNumber="539">P. a nt he l a</emphasis>
appear ciliated throughout including the tips (JMMN pers observ.).
</paragraph>
<paragraph id="8B8E36D0FFDD9E3AFF614703FAB30AB4" blockId="55.[151,1437,151,608]" pageId="55" pageNumber="539">
The
<typeStatus id="548A8872FFDD9E3AFF5D470CFED60A24" box="[251,299,296,320]" pageId="55" pageNumber="539">type</typeStatus>
species of the genus,
<taxonomicName id="4C314D53FFDD9E3AFD80470DFC9B0A24" authority="Pearson, 1969" authorityName="Pearson" authorityYear="1969" box="[550,870,295,320]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="55" pageNumber="539" phylum="Annelida" rank="species" species="lornensis">
<emphasis id="B945EAC2FFDD9E3AFD80470DFD570A24" box="[550,682,295,320]" italics="true" pageId="55" pageNumber="539">P. lornensis</emphasis>
(
<bibRefCitation id="EFA04B21FFDD9E3AFD1C470CFCA20A24" author="Pearson" box="[698,863,295,320]" pageId="55" pageNumber="539" refString="Pearson, T. H. (1969) Scionella lornensis sp. nov., a new terebellid (Polychaeta: Annelida) from the west coast of Scotland, with notes on the genus Scionella Moore, and a key to the genera of the Terebellidae recorded from European waters. Journal of Natural History, 3, 509 - 516. http: // dx. doi. org / 10.1080 / 00222936900770441" type="journal article" year="1969">Pearson, 1969</bibRefCitation>
)
</taxonomicName>
, has much larger lobes on segments 13, those of segment 1 extending ventrally from one side to another of body, lobes of segment 2 much longer, almost reaching level of prostomium and with straight distal margins, and those of segment 3 extending to the corners of the midventral shield, while in
<taxonomicName id="4C314D53FFDD9E3AFE3A47B1FDC60AC8" box="[412,571,404,428]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="55" pageNumber="539" phylum="Annelida" rank="species" species="franciscana" status="sp. nov.">
<emphasis id="B945EAC2FFDD9E3AFE3A47B1FDC60AC8" box="[412,571,404,428]" italics="true" pageId="55" pageNumber="539">P. franciscana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B9FFDD9E3AFDE747B1FD810AC8" box="[577,636,405,428]" pageId="55" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
the lobe of segment 1 is only marginal to mouth, lobes of segment 2 are almost semi-circular and only cover the posterior part of segment 1, and those of segment 3 are only lateral.
</paragraph>
</subSubSection>
<subSubSection id="C32B655BFFDD9E3AFF6147F8FAFA097C" pageId="55" pageNumber="539" type="etymology">
<paragraph id="8B8E36D0FFDD9E3AFF6147F8FAFA097C" blockId="55.[151,1437,151,608]" pageId="55" pageNumber="539">
<emphasis id="B945EAC2FFDD9E3AFF6147F8FEB30A91" bold="true" box="[199,334,476,501]" pageId="55" pageNumber="539">Etymology.</emphasis>
We name this species as “
<taxonomicName id="4C314D53FFDD9E3AFD3A47F8FCDD0A90" box="[668,800,476,500]" class="Polychaeta" family="Terebellidae" genus="Pistella" kingdom="Animalia" order="Terebellida" pageId="55" pageNumber="539" phylum="Annelida" rank="species" species="franciscana">
<emphasis id="B945EAC2FFDD9E3AFD3A47F8FCDD0A90" box="[668,800,476,500]" italics="true" pageId="55" pageNumber="539">franciscana</emphasis>
</taxonomicName>
” due to the unusual pattern of ciliation of branchial filaments, which encircle a bald tip, resembling the head of Franciscan monks from the Middle Ages.
</paragraph>
</subSubSection>
<subSubSection id="C32B655BFFDD9E3AFF614400FB1A0959" box="[199,1255,548,573]" pageId="55" pageNumber="539" type="materials_examined">
<paragraph id="8B8E36D0FFDD9E3AFF614400FB1A0959" blockId="55.[151,1437,151,608]" box="[199,1255,548,573]" pageId="55" pageNumber="539">
<emphasis id="B945EAC2FFDD9E3AFF614400FE980959" bold="true" box="[199,357,548,573]" pageId="55" pageNumber="539">
<typeStatus id="548A8872FFDD9E3AFF614400FEFF0959" box="[199,258,548,573]" pageId="55" pageNumber="539">Type</typeStatus>
locality.
</emphasis>
North of North Point,
<geoCoordinate id="EE055017FFDD9E3AFDC24400FD180959" box="[612,741,548,573]" direction="south" orientation="latitude" pageId="55" pageNumber="539" precision="15" value="-14.643333">14°38'36&quot;S</geoCoordinate>
,
<geoCoordinate id="EE055017FFDD9E3AFD574400FC7F0959" box="[753,898,548,573]" direction="east" orientation="longitude" pageId="55" pageNumber="539" precision="15" value="145.4525">145°27'09&quot;E</geoCoordinate>
, Lizard
<collectingCountry id="F3267640FFDD9E3AFC784400FBD80959" box="[990,1061,548,573]" name="Iceland" pageId="55" pageNumber="539">Island</collectingCountry>
, GBR,
<collectingCountry id="F3267640FFDD9E3AFBDE4400FB1E0959" box="[1144,1251,548,573]" name="Australia" pageId="55" pageNumber="539">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B655BFFDD9E3AFF614463FD190904" box="[199,740,583,608]" pageId="55" pageNumber="539" type="distribution">
<paragraph id="8B8E36D0FFDD9E3AFF614463FD190904" blockId="55.[151,1437,151,608]" box="[199,740,583,608]" pageId="55" pageNumber="539">
<emphasis id="B945EAC2FFDD9E3AFF614463FE9D0904" bold="true" box="[199,352,583,608]" pageId="55" pageNumber="539">Distribution.</emphasis>
Widespread around Lizard
<collectingCountry id="F3267640FFDD9E3AFD314463FD220904" box="[663,735,583,608]" name="Iceland" pageId="55" pageNumber="539">Island</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>