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<document id="9FB93B0A5F83BF397CF2C016F05BBE10" ID-CLB-Dataset="8626" ID-DOI="10.5281/zenodo.4476396" ID-GBIF-Dataset="f638455f-2539-4bb0-816f-bafbc0b7c1ac" ID-ISSN="ISSN 0374-1036" ID-Zenodo-Dep="4476396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1611663472831" checkinUser="felipe" docAuthor="Kment, Petr" docDate="2005" docId="039887C4A312F47E7CCC7E5D32AD500D" docLanguage="en" docName="ActaEntMusNatPra.45.21-50.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Mahea durrelli, sp. nov." docType="treatment" docVersion="6" lastPageNumber="36" masterDocId="FFA1FFBCA31EF4717D327C42312D533A" masterDocTitle="Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles" masterLastPageNumber="50" masterPageNumber="21" pageNumber="33" updateTime="1698925359088" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="765AA24BE3BD14109519FB08833ED96C">Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles</mods:title>
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<mods:namePart id="A2DDF42BAA5D56464D03B5AC926F7B6E">Kment, Petr</mods:namePart>
<mods:affiliation id="80E44F00D5EBF1F8782D1CDFD1EA444A">) &amp;) Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: sigara @ post. cz &amp;) &amp;) Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:date id="F6BED4C5BB3260AA44617D4026527B7D">2005</mods:date>
<mods:detail id="D6E24A328CD1D4729DF66E091E390041" type="series">
<mods:title id="90541043CB0A2C0E3321F46F294F7A9F">suppl.</mods:title>
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<mods:number id="2B642F0F1DDC97DD0D0515FC21AD17A5">2005-12-31</mods:number>
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<mods:classification id="D5B1FD0FE47E34D9757E20D73AA8AA2A">journal article</mods:classification>
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<treatment id="039887C4A312F47E7CCC7E5D32AD500D" ID-DOI="http://doi.org/10.5281/zenodo.4476432" ID-GBIF-Taxon="177135177" ID-Zenodo-Dep="4476432" LSID="urn:lsid:plazi:treatment:039887C4A312F47E7CCC7E5D32AD500D" httpUri="http://treatment.plazi.org/id/039887C4A312F47E7CCC7E5D32AD500D" lastPageId="15" lastPageNumber="36" pageId="12" pageNumber="33">
<subSubSection id="C32B6559A312F47D7CCC7E5D322D510D" box="[510,768,543,567]" pageId="12" pageNumber="33" type="nomenclature">
<paragraph id="8B8E36D2A312F47D7CCC7E5D322D510D" blockId="12.[510,768,543,567]" box="[510,768,543,567]" pageId="12" pageNumber="33">
<heading id="D0C681BEA312F47D7CCC7E5D322D510D" bold="true" box="[510,768,543,567]" centered="true" fontSize="8" level="2" pageId="12" pageNumber="33" reason="3">
<emphasis id="B945EAC0A312F47D7CCC7E5D322D510D" bold="true" box="[510,768,543,567]" pageId="12" pageNumber="33">
<taxonomicName id="4C314D51A312F47D7CCC7E5D338E510D" ID-CoL="7VRC5" authority="Kment, 2005" box="[510,675,543,567]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="durrelli" status="sp. nov.">
<emphasis id="B945EAC0A312F47D7CCC7E5D338E510D" bold="true" box="[510,675,543,567]" italics="true" pageId="12" pageNumber="33">Mahea durrelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657BBA312F47D7F997E5D322D510D" box="[683,768,543,567]" pageId="12" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B6559A312F47D7F047E7D33EA5168" box="[566,711,575,594]" pageId="12" pageNumber="33" type="description">
<paragraph id="8B8E36D2A312F47D7F047E7D33EA5168" blockId="12.[566,711,575,594]" box="[566,711,575,594]" pageId="12" pageNumber="33">
(
<figureCitation id="130A2A57A312F47D7F0F7E7D338C5168" box="[573,673,575,594]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="13.[113,152,1072,1091]" captionStartId-1="14.[151,190,1263,1282]" captionTargetBox-0="[115,1088,220,1043]" captionTargetBox-1="[151,1126,220,1229]" captionTargetId-0="figure-165@13.[113,1088,220,1043]" captionTargetId-1="figure-112@14.[151,1126,220,1229]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." captionText-1="Figs. 24-27. Mahea durrelli sp. nov. 24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4476418" figureDoi-1="http://doi.org/10.5281/zenodo.4476422" httpUri-0="https://zenodo.org/record/4476418/files/figure.png" httpUri-1="https://zenodo.org/record/4476422/files/figure.png" pageId="12" pageNumber="33">Figs. 20-27</figureCitation>
, 44)
</paragraph>
</subSubSection>
<subSubSection id="C32B6559A312F47D7DA57E3735335737" pageId="12" pageNumber="33" type="materials_examined">
<paragraph id="8B8E36D2A312F47D7DA57E3733B051B7" blockId="12.[151,1127,628,1037]" box="[151,669,628,653]" pageId="12" pageNumber="33">
<emphasis id="B945EAC0A312F47D7DA57E37300351B6" bold="true" box="[151,302,628,653]" pageId="12" pageNumber="33">
<typeStatus id="548A8870A312F47D7DA57E3731E351B7" box="[151,206,629,653]" pageId="12" pageNumber="33">Type</typeStatus>
locality.
</emphasis>
North-east
<collectingCountry id="F3267642A312F47D7C9E7E37330351B7" box="[428,558,629,653]" name="Madagascar" pageId="12" pageNumber="33">Madagascar</collectingCountry>
, Vohémar.
</paragraph>
<paragraph id="8B8E36D2A312F47D7DA57ED7338750F7" blockId="12.[151,1127,628,1037]" pageId="12" pageNumber="33">
<emphasis id="B945EAC0A312F47D7DA57ED730175196" bold="true" box="[151,314,660,685]" pageId="12" pageNumber="33">Type material.</emphasis>
<materialsCitation id="3B593C8FA312F47D7C0D7ED730FA51F7" ID-GBIF-Occurrence="3026650302" collectorName="J. L. Stehlik" country="Madagascar" location="Madagascar" pageId="12" pageNumber="33" specimenCount="1" typeStatus="holotype">
<typeStatus id="548A8870A312F47D7C0D7ED7308B5191" box="[319,422,661,684]" pageId="12" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
: ♁, Vohémar /
<collectingCountry id="F3267642A312F47D7F787ED733E75197" box="[586,714,661,685]" name="Madagascar" pageId="12" pageNumber="33">Madagascar</collectingCountry>
[p] // Collectio /
<collectorName id="26C45304A312F47D7E4B7ED732C15197" box="[889,1004,661,685]" pageId="12" pageNumber="33">J.L.Stehlík</collectorName>
/ Moravské museum, Brno [p] // ♁ [p] //
</materialsCitation>
<materialsCitation id="3B593C8FA312F47D7CD27EF7308E5037" ID-GBIF-Occurrence="3026650306" collectionCode="MMBC" country="Madagascar" pageId="12" pageNumber="33" specimenCount="2" typeStatus="holotype">
<typeStatus id="548A8870A312F47D7CD27EF7335351F7" box="[480,638,693,717]" pageId="12" pageNumber="33" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="4C314D51A312F47D7FAB7EF733D451F7" authorityName=": KUMAR" authorityYear="1974" box="[665,761,693,717]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="47" phylum="Arthropoda" rank="genus">MAHEA</taxonomicName>
/ DURRELLI / sp. nov. / det. P. KMENT 2005 [p, red label] (
<collectionCode id="ED20AE17A312F47D7CEB7E97331C51D7" box="[473,561,725,749]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="12" pageNumber="33">MMBC</collectionCode>
). The
<typeStatus id="548A8870A312F47D7F417E9733FD51D7" box="[627,720,725,749]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
is mounted with the pygophore glued on the same piece of card
</materialsCitation>
; I have dissected the aedeagus and left paramere and placed them in a plastic microvial with glycerol attached to the same pin.
<typeStatus id="548A8870A312F47D7FC57F5732745011" box="[759,857,789,812]" pageId="12" pageNumber="33" type="paratype">PARATYPE</typeStatus>
: ♁, Vohémar /
<collectingCountry id="F3267642A312F47D7ECC7F57319A5076" name="Madagascar" pageId="12" pageNumber="33">Madagascar</collectingCountry>
[p] // COLLECTIO / NATIONAL MUSEUM /
<collectingRegion id="49F5F830A312F47D7FFD7F7732235077" box="[719,782,821,845]" country="Czech Republic" name="Praha" pageId="12" pageNumber="33">Praha</collectingRegion>
,
<collectingCountry id="F3267642A312F47D7E2E7F7732E55077" box="[796,968,821,845]" name="Czech Republic" pageId="12" pageNumber="33">Czech Republic</collectingCountry>
[p] // ♁ [p] //
<materialsCitation id="3B593C8FA312F47D7DA57F17326D50B7" ID-GBIF-Occurrence="3026650303" collectionCode="NMPC" country="Madagascar" pageId="12" pageNumber="33" specimenCount="2" typeStatus="paratype">
<typeStatus id="548A8870A312F47D7DA57F1730065057" box="[151,299,853,877]" pageId="12" pageNumber="33" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="4C314D51A312F47D7C097F1730B75057" authorityName=": KUMAR" authorityYear="1974" box="[315,410,853,877]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="47" phylum="Arthropoda" rank="genus">MAHEA</taxonomicName>
/ DURRELLI / sp. nov. / det. P. KMENT 2005 [p, red label] (
<collectionCode id="ED20AE17A312F47D793E7F1735715057" box="[1036,1116,853,877]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="12" pageNumber="33">NMPC</collectionCode>
). The
<typeStatus id="548A8870A312F47D7DF57F37300F50B7" box="[199,290,885,909]" pageId="12" pageNumber="33" type="paratype">paratype</typeStatus>
is mounted on the tip of a pentagonal piece of card
</materialsCitation>
; left antennomere 2 broken, left antennomeres 3-4 missing, and right spine on sternum 6 broken; I have removed the pygophore and glued it on the same piece of card.
</paragraph>
<paragraph id="8B8E36D2A312F47D7D847F9735335737" blockId="12.[151,1127,628,1037]" pageId="12" pageNumber="33">
Note: The locality label Vohémar is identical with those found in material collected in
<collectingCountry id="F3267642A312F47D7DA57FB7303A5737" box="[151,279,1013,1037]" name="Madagascar" pageId="12" pageNumber="33">Madagascar</collectingCountry>
by C. Lamberton in 1937 and housed in NMPC (see
<bibRefCitation id="EFA04B23A312F47D7E627FB7353E5737" author="HOBERLANDT L." box="[848,1043,1013,1037]" pageId="12" pageNumber="33" pagination="135 - 150" refId="ref13474" refString="HOBERLANDT L. 1942: Heteroptera Madagascariensia in Museo Pragensi. III. Ad Ploiariinarum cognitionem. Sbornik Entomologickeho Oddeleni pri Zoologickych Sbirkach Zemskeho Musea v Praze 20: 135 - 150 (in Czech and Latin)." type="journal article" year="1942">HOBERLANDT 1942</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B6559A312F47E7DA57861326652CD" lastPageId="15" lastPageNumber="36" pageId="12" pageNumber="33" type="description">
<paragraph id="8B8E36D2A312F47D7DA57861351F5706" blockId="12.[151,1127,1059,1596]" box="[151,1074,1059,1084]" pageId="12" pageNumber="33">
<emphasis id="B945EAC0A312F47D7DA57861300C5701" bold="true" box="[151,289,1059,1083]" pageId="12" pageNumber="33">Description.</emphasis>
Body narrow, ochraceous with reddish-brown punctures, slightly shining.
</paragraph>
<paragraph id="8B8E36D2A312F47D7D847806354A5761" blockId="12.[151,1127,1059,1596]" box="[182,1127,1092,1116]" pageId="12" pageNumber="33">
Male (
<typeStatus id="548A8870A312F47D7DC47806307E5766" box="[246,339,1092,1116]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
) (Fig. 44). Length
<quantity id="4CC99B37A312F47D7F22780633605761" box="[528,589,1092,1116]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="33" unit="mm" value="6.0">6 mm</quantity>
, width of pronotum between humeral spines 3.1 mm.
</paragraph>
<paragraph id="8B8E36D2A312F47D7D84782632F156E1" blockId="12.[151,1127,1059,1596]" pageId="12" pageNumber="33">
Head ochraceous with irregular rows and groups of coarse black or brown punctures (two rows on mandibular plates less distinct than in other species, vertex with U-shaped figure). Head shorter than wide across eyes (1: 1.42), ratio of width of head: width of the pronotum between humeral angles equal to 1: 1.97. Mandibular plates only slightly narrowing basally, apically arcuately curved inwards, not meeting in front of clypeus; apex of clypeus free, clypeus slightly shorter than mandibular plates. Eyes ochraceous with brownish spots; ocelli reddish, situated near anterior pronotal margin. Antenniferous tubercles each with black longitudinal spot laterally; antennae castaneous, antennomeres 1-2 basally paler; antennomere 2 with erect pubescence, some hairs longer than its diameter; lengths of antennomeres: 1
<quantity id="4CC99B37A312F47D7EF4792635355641" box="[966,1048,1380,1404]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="33" unit="mm" value="0.3">0.3 mm</quantity>
, 2
<quantity id="4CC99B37A312F47D7977792631EE56A1" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="33" unit="mm" value="1.7">1.7 mm</quantity>
, 3 1.0 mm, 4
<quantity id="4CC99B37A312F47D7C4479C630EB56A1" box="[374,454,1412,1436]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="33" unit="mm" value="0.9">0.9 mm</quantity>
(ratio 1: 5.67: 3.33: 3). Head ventrally ochraceous with sparse reddish punctures and one pair of larger, blackish L-shaped spots postero-medially next to bucculae; rostrum ochraceous, apex blackened, reaching middle of metacoxae.
</paragraph>
<paragraph id="8B8E36D2A312F47C7D8479A6304F56B8" blockId="12.[151,1127,1059,1596]" lastBlockId="13.[113,1089,1162,1602]" lastPageId="13" lastPageNumber="34" pageId="12" pageNumber="33">
Pronotum ochraceous with thick, coarse, uniformly distributed reddish-brown punctures; two rows of black punctures in area of cicatrices; humeral angles reddish. Anterior margin of pronotum only slightly concave, almost straight medially; antero-lateral margins rounded, proximally almost parallel, medially abruptly curved laterad; humeral angles prominent, conically produced laterad and upwards, apically constricted, with long narrow spine (
<figureCitation id="130A2A57A313F47C7ED378E831A157D8" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="13" pageNumber="34">Figs. 20- 21</figureCitation>
); posterior margin of pronotum medially more concave than in other species. Scutellum narrowly triangular, slightly sinuated medially, almost flat, only anterior part slightly sloping caudad; ochraceous, with reddish-brown punctures, colouration of neighbouring punctures locally coalescent in anterior part; apex narrowly triangular, with small black spot. Hemelytra narrow, suture brownish, membrane slightly brownish infuscate, not surpassing postero-lateral angles of sternum 7.
</paragraph>
<caption id="DF4E665AA313F47C7D43787230D45764" ID-DOI="http://doi.org/10.5281/zenodo.4476418" ID-Zenodo-Dep="4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="13" pageNumber="34" startId="13.[113,152,1072,1091]" targetBox="[115,1088,220,1043]" targetPageId="13">
<paragraph id="8B8E36D2A313F47C7D43787230D45764" blockId="13.[113,1089,1072,1118]" pageId="13" pageNumber="34">
Figs. 20-23.
<taxonomicName id="4C314D51A313F47C7DD37872304E5779" box="[225,355,1072,1091]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="durrelli" status="sp. nov.">
<emphasis id="B945EAC0A313F47C7DD37872304E5779" box="[225,355,1072,1091]" italics="true" pageId="13" pageNumber="34">Mahea durrelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657BBA313F47C7C5878733082577E" box="[362,431,1073,1092]" pageId="13" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view.
</paragraph>
</caption>
<paragraph id="8B8E36D2A313F47C7DA279C8301C5578" blockId="13.[113,1089,1162,1602]" pageId="13" pageNumber="34">
Thorax ventrally ochraceous with reddish-brown punctures; colouration of punctures especially on meso- and metapleuron coalescent, forming brown spots alternating with ochraceous unpunctured spots; acetabula, peritreme, and elevated unpunctured spots on pleura ochraceous. Metapleura not swollen, its margin only slightly visible from above. Legs ochraceous, tarsomeres 2 brownish. Hind femora and tibiae straight, lacking granulation on inner margins (
<figureCitation id="130A2A57A313F47C7DE77A6830095578" box="[213,292,1578,1602]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="13" pageNumber="34">Fig. 22</figureCitation>
).
</paragraph>
<caption id="DF4E665AA310F47F7DA578AD32F75627" ID-DOI="http://doi.org/10.5281/zenodo.4476422" ID-Zenodo-Dep="4476422" httpUri="https://zenodo.org/record/4476422/files/figure.png" pageId="14" pageNumber="35" startId="14.[151,190,1263,1282]" targetBox="[151,1126,220,1229]" targetPageId="14">
<paragraph id="8B8E36D2A310F47F7DA578AD32F75627" blockId="14.[151,1127,1263,1309]" pageId="14" pageNumber="35">
Figs. 24-27.
<taxonomicName id="4C314D51A310F47F7C3B78AD30A15638" box="[265,396,1263,1282]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="durrelli" status="sp. nov.">
<emphasis id="B945EAC0A310F47F7C3B78AD30A15638" box="[265,396,1263,1282]" italics="true" pageId="14" pageNumber="35">Mahea durrelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657BBA310F47F7CA778AD30F65638" box="[405,475,1263,1282]" pageId="14" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B8E36D2A310F47F7D84790833AB5578" blockId="14.[151,1127,1354,1602]" pageId="14" pageNumber="35">
Abdomen ventrally ochraceous with small and shallow red punctures, sterna 3-6 with three pairs of small black spots: one oval spot on anterior margin next to median carina, one round spot medially on each side, one small, round, impressed spot laterally on suture between sterna; sternum 7 with pair of oval blackish spots on anterior margin, postero-lateral angles of sternum 7 black. Median carina well developed on sterna 3-6, only basally on sternum 7, disappearing caudad (
<figureCitation id="130A2A57A310F47F7CB279A830FD5538" box="[384,464,1514,1538]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="14" pageNumber="35">Fig. 23</figureCitation>
). Connexiva ochraceous, postero-lateral angles without spines except sternum 6 with long spine curved postero-laterad; postero-lateral angles of sternum 7 lanceolate, produced postero-laterad (
<figureCitation id="130A2A57A310F47F7F1B7A6833545578" box="[553,633,1578,1602]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="14" pageNumber="35">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8B8E36D2A311F47E7DA27C9D3300526D" blockId="15.[113,1089,223,824]" pageId="15" pageNumber="36">
Male genitalia. Pygophore brownish; less dorso-ventrally flattened (
<figureCitation id="130A2A57A311F47E7E627C9D32B053CD" box="[848,925,223,247]" captionStart="Figs" captionStartId="14.[151,190,1263,1282]" captionTargetBox="[151,1126,220,1229]" captionTargetId="figure-112@14.[151,1126,220,1229]" captionTargetPageId="14" captionText="Figs. 24-27. Mahea durrelli sp. nov. 24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4476422" httpUri="https://zenodo.org/record/4476422/files/figure.png" pageId="15" pageNumber="36">Fig. 24</figureCitation>
); exterior opening with small triangular tubercle dorso-medially (
<figureCitation id="130A2A57A311F47E7F4D7CBD33D7522D" box="[639,762,255,279]" captionStart="Figs" captionStartId="14.[151,190,1263,1282]" captionTargetBox="[151,1126,220,1229]" captionTargetId="figure-112@14.[151,1126,220,1229]" captionTargetPageId="14" captionText="Figs. 24-27. Mahea durrelli sp. nov. 24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4476422" httpUri="https://zenodo.org/record/4476422/files/figure.png" pageId="15" pageNumber="36">Figs. 24-25</figureCitation>
); hypophysis of paramere oval, apically broadly rounded, not pointed, inner margin only slightly incised (
<figureCitation id="130A2A57A311F47E7E4D7D5D32E2520D" box="[895,975,287,311]" captionStart="Figs" captionStartId="14.[151,190,1263,1282]" captionTargetBox="[151,1126,220,1229]" captionTargetId="figure-112@14.[151,1126,220,1229]" captionTargetPageId="14" captionText="Figs. 24-27. Mahea durrelli sp. nov. 24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4476422" httpUri="https://zenodo.org/record/4476422/files/figure.png" pageId="15" pageNumber="36">Fig. 26</figureCitation>
); vesica of aedeagus long, looped medially (
<figureCitation id="130A2A57A311F47E7CE27D7D330D526D" box="[464,544,319,343]" captionStart="Figs" captionStartId="14.[151,190,1263,1282]" captionTargetBox="[151,1126,220,1229]" captionTargetId="figure-112@14.[151,1126,220,1229]" captionTargetPageId="14" captionText="Figs. 24-27. Mahea durrelli sp. nov. 24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4476422" httpUri="https://zenodo.org/record/4476422/files/figure.png" pageId="15" pageNumber="36">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8B8E36D2A311F47E7DA27D1D3077524D" blockId="15.[113,1089,223,824]" box="[144,346,351,375]" pageId="15" pageNumber="36">Female. Unknown.</paragraph>
<paragraph id="8B8E36D2A311F47E7D437D3D326652CD" blockId="15.[113,1089,223,824]" pageId="15" pageNumber="36">
<emphasis id="B945EAC0A311F47E7D437D3D31C252AD" bold="true" box="[113,239,383,407]" pageId="15" pageNumber="36">Variability.</emphasis>
<typeStatus id="548A8870A311F47E7DCB7DC2307B52A2" box="[249,342,384,408]" pageId="15" pageNumber="36" type="paratype">Paratype</typeStatus>
male (length
<quantity id="4CC99B37A311F47E7CDD7D3D336852AD" box="[495,581,383,407]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="15" pageNumber="36" unit="mm" value="5.9">5.9 mm</quantity>
, width of pronotum between humeral spines
<quantity id="4CC99B37A311F47E79017D3D31B0528D" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="36" unit="mm" value="3.0">3 mm</quantity>
, length of antennomeres: 1
<quantity id="4CC99B37A311F47E7CE67DDD3305528D" box="[468,552,415,439]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="36" unit="mm" value="0.3">0.3 mm</quantity>
, 2
<quantity id="4CC99B37A311F47E7F6B7DDD3381528D" box="[601,684,415,439]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="36" unit="mm" value="1.5">1.5 mm</quantity>
, 3
<quantity id="4CC99B37A311F47E7FEC7DDD321C528D" box="[734,817,415,439]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="36" unit="mm" value="0.9">0.9 mm</quantity>
, 4
<quantity id="4CC99B37A311F47E7E517DDD3298528D" box="[867,949,415,439]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="36" unit="mm" value="0.9">0.9 mm</quantity>
) differs from the
<typeStatus id="548A8870A311F47E7DA57DFD31D952ED" box="[151,244,447,471]" pageId="15" pageNumber="36" type="holotype">holotype</typeStatus>
by having the mandibular plates meeting in front of the clypeus, punctures on the head darker, and the pronotum with a vague ochraceous median line.
</paragraph>
</subSubSection>
<subSubSection id="C32B6559A311F47E7D437DBD323651AD" pageId="15" pageNumber="36" type="diagnosis">
<paragraph id="8B8E36D2A311F47E7D437DBD323651AD" blockId="15.[113,1089,223,824]" pageId="15" pageNumber="36">
<emphasis id="B945EAC0A311F47E7D437DBD3044512D" bold="true" box="[113,361,511,535]" pageId="15" pageNumber="36">Differential diagnosis.</emphasis>
<taxonomicName id="4C314D51A311F47E7C5D7DBD3323512D" box="[367,526,511,535]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="durrelli">
<emphasis id="B945EAC0A311F47E7C5D7DBD3323512D" box="[367,526,511,535]" italics="true" pageId="15" pageNumber="36">Mahea durrelli</emphasis>
</taxonomicName>
differes from
<taxonomicName id="4C314D51A311F47E7F917E423239512D" baseAuthorityName="Cachan" baseAuthorityYear="1952" box="[675,788,511,535]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="andriai">
<emphasis id="B945EAC0A311F47E7F917E423239512D" box="[675,788,511,535]" italics="true" pageId="15" pageNumber="36">M. andriai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D51A311F47E7E137E4232BE512D" box="[801,915,511,535]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="distanti">
<emphasis id="B945EAC0A311F47E7E137E4232BE512D" box="[801,915,511,535]" italics="true" pageId="15" pageNumber="36">M. distanti</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C314D51A311F47E7EFE7E42356D512D" box="[972,1088,511,535]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B945EAC0A311F47E7EFE7E42356D512D" box="[972,1088,511,535]" italics="true" pageId="15" pageNumber="36">M. parvula</emphasis>
</taxonomicName>
in having each humeral angle abruptly produced into a long spine. From
<taxonomicName id="4C314D51A311F47E7E577E6232F1510D" authorityName="Distant" authorityYear="1909" box="[869,988,543,567]" class="Insecta" family="Acanthosomatidae" genus="Mahea" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="sexualis">
<emphasis id="B945EAC0A311F47E7E577E6232F1510D" box="[869,988,543,567]" italics="true" pageId="15" pageNumber="36">M. sexualis</emphasis>
</taxonomicName>
, it differs by having the lateral pronotal margins deeply sinuated (
<figureCitation id="130A2A57A311F47E7FF97E7D3236516D" box="[715,795,575,599]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="15" pageNumber="36">Fig. 20</figureCitation>
); the humeral spines raised (
<figureCitation id="130A2A57A311F47E7D4B7E1D31DE514D" box="[121,243,607,631]" captionStart="Figs" captionStartId="13.[113,152,1072,1091]" captionTargetBox="[115,1088,220,1043]" captionTargetId="figure-165@13.[113,1088,220,1043]" captionTargetPageId="13" captionText="Figs. 20-23. Mahea durrelli sp. nov. 20 pronotum, dorsal view; 21 pronotum, posterior view; 22 hind leg (claws missing); 23 abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4476418" httpUri="https://zenodo.org/record/4476418/files/figure.png" pageId="15" pageNumber="36">Figs. 20-21</figureCitation>
); the metapleura not swollen; the shape of connexival spines; and the apices of parameres rounded, not pointed (
<figureCitation id="130A2A57A311F47E7CE37E3D336151AD" box="[465,588,639,663]" captionStart="Figs" captionStartId="14.[151,190,1263,1282]" captionTargetBox="[151,1126,220,1229]" captionTargetId="figure-112@14.[151,1126,220,1229]" captionTargetPageId="14" captionText="Figs. 24-27. Mahea durrelli sp. nov. 24 pygophore, posterior view; 25 pygophore, ventral view; 26 left paramere, ventral view; 27 aedeagus (basal articulatory apparatus omitted). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4476422" httpUri="https://zenodo.org/record/4476422/files/figure.png" pageId="15" pageNumber="36">Figs. 24-26</figureCitation>
) (see also the key).
</paragraph>
</subSubSection>
<subSubSection id="C32B6559A311F47E7D437EDD307F502D" pageId="15" pageNumber="36" type="etymology">
<paragraph id="8B8E36D2A311F47E7D437EDD331151CD" blockId="15.[113,1089,223,824]" pageId="15" pageNumber="36">
<emphasis id="B945EAC0A311F47E7D437EDD31DC518D" bold="true" box="[113,241,671,695]" pageId="15" pageNumber="36">Etymology.</emphasis>
The species is named in honour of Gerald Durrell (1925-1995), the famous English amateur naturalist, nature conservator and writer who paid a lot of attention to the nature of
<collectingCountry id="F3267642A311F47E7DEA7E9D307551CD" box="[216,344,735,759]" name="Madagascar" pageId="15" pageNumber="36">Madagascar</collectingCountry>
and adjacent islands.
</paragraph>
<paragraph id="8B8E36D2A311F47E7D437EBD307F502D" blockId="15.[113,1089,223,824]" box="[113,338,767,791]" pageId="15" pageNumber="36">
<emphasis id="B945EAC0A311F47E7D437EBD31F1502D" bold="true" box="[113,220,767,791]" pageId="15" pageNumber="36">Bionomy.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32B6559A311F47E7D437F5D32AD500D" box="[113,896,799,824]" pageId="15" pageNumber="36" type="distribution">
<paragraph id="8B8E36D2A311F47E7D437F5D32AD500D" blockId="15.[113,1089,223,824]" box="[113,896,799,824]" pageId="15" pageNumber="36">
<emphasis id="B945EAC0A311F47E7D437F5D302F500D" bold="true" box="[113,258,799,823]" pageId="15" pageNumber="36">Distribution.</emphasis>
North-east
<collectingCountry id="F3267642A311F47E7C4D7F5D332C500D" box="[383,513,799,823]" name="Madagascar" pageId="15" pageNumber="36">Madagascar</collectingCountry>
. Known only from the
<typeStatus id="548A8870A311F47E7FC57F6232085002" box="[759,805,800,824]" pageId="15" pageNumber="36">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>