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<document id="3B494AFBFE82FA93AF1BD0C18ABAD779" ID="10.11646/zootaxa.4242.2.7SLASH10440" ID-CLB-Dataset="36303" ID-DOI="10.11646/zootaxa.4242.2.7" ID-GBIF-Dataset="8632c8ac-22fd-42bc-b95c-820c0c029c20" ID-ISSN="1175-5326" ID-Zenodo-Dep="376427" ID-ZooBank="8511A4B8-D82F-49EE-9132-E941B4C5D2C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489137585520" checkinUser="plazi" docAuthor="Puchalski, Mateusz &amp; Giłka, Wojciech" docDate="2017" docId="03988791FFBCE772FF33266D42D4FC5D" docLanguage="en" docName="zootaxa.4242.2.7SLASH10440.pdf" docOrigin="Zootaxa 4242 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladotanytarsus bilyji Gilka et Puchalski, sp. nov." docType="treatment" docVersion="9" lastPageNumber="350" masterDocId="FFA1FFE9FFBFE774FFA4207E474CFF82" masterDocTitle="Cladotanytarsus Kieffer (Diptera: Chironomidae): several distinctive species reviewed on the basis of records from Canada and USA" masterLastPageNumber="358" masterPageNumber="344" pageNumber="347" updateTime="1698437824612" updateUser="ExternalLinkService">
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<mods:title id="7AE96358AEB634E1FCC955939FB31869">Cladotanytarsus Kieffer (Diptera: Chironomidae): several distinctive species reviewed on the basis of records from Canada and USA</mods:title>
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<mods:namePart id="3DD478B9B09491BA70AF620C2BC1A34D">Puchalski, Mateusz</mods:namePart>
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<mods:namePart id="DB9EBBB3FC8147BD2CA5263EB86920B4">Giłka, Wojciech</mods:namePart>
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<mods:date id="63E91E26B41B5FA810B55C52BCC5DB71">2017</mods:date>
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<treatment id="03988791FFBCE772FF33266D42D4FC5D" ID-DOI="http://doi.org/10.5281/zenodo.5613457" ID-GBIF-Taxon="127843608" ID-Zenodo-Dep="5613457" LSID="urn:lsid:plazi:treatment:03988791FFBCE772FF33266D42D4FC5D" httpUri="http://treatment.plazi.org/id/03988791FFBCE772FF33266D42D4FC5D" lastPageId="6" lastPageNumber="350" pageId="3" pageNumber="347">
<subSubSection id="C32B650CFFBCE777FF33266D4646F9CD" pageId="3" pageNumber="347" type="nomenclature">
<paragraph id="8B8E3687FFBCE777FF33266D45BAF9AC" blockId="3.[151,758,1555,1615]" box="[151,758,1555,1582]" pageId="3" pageNumber="347">
<heading id="D0C681EBFFBCE777FF33266D45BAF9AC" bold="true" box="[151,758,1555,1582]" fontSize="11" level="1" pageId="3" pageNumber="347" reason="1">
<emphasis id="B945EA95FFBCE777FF33266D45BAF9AC" bold="true" box="[151,758,1555,1582]" pageId="3" pageNumber="347">
<taxonomicName id="4C314D04FFBCE777FF33266D45C0F9AC" authority="Gilka et Puchalski" authorityName="Gilka et Puchalski" box="[151,652,1555,1582]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="bilyji" status="sp. nov.">
<emphasis id="B945EA95FFBCE777FF33266D46D2F9AF" bold="true" box="[151,414,1555,1581]" italics="true" pageId="3" pageNumber="347">Cladotanytarsus bilyji</emphasis>
Giłka
<emphasis id="B945EA95FFBCE777FE57266D4544F9AF" bold="true" box="[499,520,1555,1581]" italics="true" pageId="3" pageNumber="347">et</emphasis>
Puchalski
</taxonomicName>
,
<taxonomicNameLabel id="A27657EEFFBCE777FD3C266A45BAF9AC" box="[664,758,1556,1582]" pageId="3" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E3687FFBCE777FF3326484646F9CD" blockId="3.[151,758,1555,1615]" box="[151,266,1590,1615]" pageId="3" pageNumber="347">
(
<figureCitation id="130A2A02FFBCE777FF3B264847ABF9CD" box="[159,231,1590,1615]" captionStart="FIGURE 2" captionStartId="4.[151,250,599,621]" captionTargetBox="[151,1436,193,578]" captionTargetId="figure@4.[151,1436,193,578]" captionTargetPageId="4" captionText="FIGURE 2. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. Wing." httpUri="https://zenodo.org/record/376429/files/figure.png" pageId="3" pageNumber="347">Figs 2</figureCitation>
,
<figureCitation id="130A2A02FFBCE777FF572649464DF9CD" box="[243,257,1591,1615]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="3" pageNumber="347">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B650CFFBCE777FF3326004442F880" pageId="3" pageNumber="347" type="materials_examined">
<paragraph id="8B8E3687FFBCE777FF3326004442F880" blockId="3.[151,1437,1662,2011]" pageId="3" pageNumber="347">
<materialsCitation id="3B593CDAFFBCE777FF332600435FF939" ID-GBIF-Occurrence="1438637184" collectingDate="1971-06-16" collectorName="E. Johnson &amp; M. Roberts &amp; O. A. Saether" country="Canada" latitude="51.55" location="Old Fishing Dock" longLatPrecision="1089" longitude="-96.71667" municipality="Lake Winnipeg" pageId="3" pageNumber="347" specimenCount="2" stateProvince="MANITOBA" typeStatus="holotype">
<emphasis id="B945EA95FFBCE777FF332600460BF915" bold="true" box="[151,327,1662,1687]" pageId="3" pageNumber="347">
<typeStatus id="548A8825FFBCE777FF332600479EF915" box="[151,210,1662,1687]" pageId="3" pageNumber="347">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548A8825FFBCE777FEF4260046F7F915" box="[336,443,1662,1687]" pageId="3" pageNumber="347" type="holotype">Holotype</typeStatus>
, adult male:
<collectingCountry id="F3267617FFBCE777FDF626014581F915" box="[594,717,1663,1687]" name="Canada" pageId="3" pageNumber="347">CANADA</collectingCountry>
,
<collectingRegion id="49F5F865FFBCE777FD7E2601443FF915" box="[730,883,1663,1687]" country="Canada" name="Manitoba" pageId="3" pageNumber="347">MANITOBA</collectingRegion>
,
<collectingMunicipality id="6BEAACFDFFBCE777FC252600437FF915" box="[897,1075,1662,1687]" pageId="3" pageNumber="347">Lake Winnipeg</collectingMunicipality>
,
<location id="8EEE605CFFBCE777FBE426004242F915" LSID="urn:lsid:plazi:treatment:03988791FFBCE772FF33266D42D4FC5D:8EEE605CFFBCE777FBE426004242F915" box="[1088,1294,1662,1687]" country="Canada" latitude="51.55" longLatPrecision="1089" longitude="-96.71667" municipality="Lake Winnipeg" name="Old Fishing Dock" pageId="3" pageNumber="347" stateProvince="MANITOBA">Old Fishing Dock</location>
(
<geoCoordinate id="EE055040FFBCE777FA84260042C6F914" box="[1312,1418,1662,1687]" direction="north" orientation="latitude" pageId="3" pageNumber="347" precision="925" value="51.55">51°33 N</geoCoordinate>
/
<geoCoordinate id="EE055040FFBCE777FF3326DC4647F939" box="[151,267,1698,1723]" direction="west" orientation="longitude" pageId="3" pageNumber="347" precision="925" value="-96.71667">96°43 W</geoCoordinate>
),
<date id="FF8F1047FFBCE777FE8526DC46F1F939" box="[289,445,1698,1723]" pageId="3" pageNumber="347" value="1971-06-16">
<collectingDate id="EFCBE9AFFFBCE777FE8526DC46F1F939" box="[289,445,1698,1723]" pageId="3" pageNumber="347" value="1971-06-16">16 June 1971</collectingDate>
</date>
, leg.
<collectorName id="26C45351FFBCE777FE5A26DD4537F939" box="[510,635,1698,1723]" pageId="3" pageNumber="347">E. Johnson</collectorName>
&amp;
<collectorName id="26C45351FFBCE777FD0026DD4468F939" box="[676,804,1698,1723]" pageId="3" pageNumber="347">M. Roberts</collectorName>
(prep.
<collectorName id="26C45351FFBCE777FCDD26DD4341F939" box="[889,1037,1698,1723]" pageId="3" pageNumber="347">O.A. Saether</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B593CDAFFBCE777FB8426DD4563F881" ID-GBIF-Occurrence="1438637182" collectingDate="1999-09-20" collectorName="J. Trybula. Ex &amp; M. &amp; J. E. Sublette" country="United States of America" county="Clermont County" latitude="39.118332" location="Baldwin Road" longLatPrecision="19" longitude="-84.216385" municipality="Shayler Run" pageId="3" pageNumber="347" specimenCount="1" specimenCount-adult="1" stateProvince="OHIO" typeStatus="paratype">
<typeStatus id="548A8825FFBCE777FB8426DD43CAF939" box="[1056,1158,1699,1723]" pageId="3" pageNumber="347" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F3267617FFBCE777FB3026DD4383F939" box="[1172,1231,1699,1723]" name="United States of America" pageId="3" pageNumber="347">USA</collectingCountry>
.
<collectingRegion id="49F5F865FFBCE777FB7826DD4269F939" box="[1244,1317,1699,1723]" country="United States of America" name="Ohio" pageId="3" pageNumber="347">OHIO</collectingRegion>
.
<collectingCounty id="62EF4E0BFFBCE777FA9626DC47A7F95D" pageId="3" pageNumber="347">Clermont County</collectingCounty>
,
<collectingMunicipality id="6BEAACFDFFBCE777FF5C26B846CAF95C" box="[248,390,1734,1759]" pageId="3" pageNumber="347">Shayler Run</collectingMunicipality>
at
<location id="8EEE605CFFBCE777FE0A26B8451EF95D" LSID="urn:lsid:plazi:treatment:03988791FFBCE772FF33266D42D4FC5D:8EEE605CFFBCE777FE0A26B8451EF95D" box="[430,594,1734,1759]" country="United States of America" county="Clermont County" latitude="39.118332" longLatPrecision="19" longitude="-84.216385" municipality="Shayler Run" name="Baldwin Road" pageId="3" pageNumber="347" stateProvince="OHIO">Baldwin Road</location>
(
<geoCoordinate id="EE055040FFBCE777FDC026B845B4F95C" box="[612,760,1734,1759]" direction="north" orientation="latitude" pageId="3" pageNumber="347" precision="15" value="39.118332">39°0706” N</geoCoordinate>
/
<geoCoordinate id="EE055040FFBCE777FCB726B844FCF95D" box="[787,944,1734,1759]" direction="west" orientation="longitude" pageId="3" pageNumber="347" precision="15" value="-84.216385">84°1259” W</geoCoordinate>
),
<date id="FF8F1047FFBCE777FC6226B943E5F95D" box="[966,1193,1734,1759]" pageId="3" pageNumber="347" value="1999-09-20">
<collectingDate id="EFCBE9AFFFBCE777FC6226B943E5F95D" box="[966,1193,1734,1759]" pageId="3" pageNumber="347" value="1999-09-20">20 September 1999</collectingDate>
</date>
,
<specimenCount id="9D37FD0EFFBCE777FB1326B9424AF95D" box="[1207,1286,1734,1759]" pageId="3" pageNumber="347" type="adult">1 adult</specimenCount>
male, leg.
<collectorName id="26C45351FFBCE777FA2E26B94657F880" pageId="3" pageNumber="347">J. Trybula. Ex</collectorName>
coll.
<collectorName id="26C45351FFBCE777FEFF26954563F881" box="[347,559,1770,1795]" pageId="3" pageNumber="347">M. &amp; J.E. Sublette</collectorName>
</materialsCitation>
. Deposit in DEUM.
</paragraph>
</subSubSection>
<subSubSection id="C32B650CFFBCE777FF63277045B8F8C8" pageId="3" pageNumber="347" type="etymology">
<paragraph id="8B8E3687FFBCE777FF63277045B8F8C8" blockId="3.[151,1437,1662,2011]" pageId="3" pageNumber="347">
<emphasis id="B945EA95FFBCE777FF63277046E4F8A5" bold="true" box="[199,424,1806,1831]" pageId="3" pageNumber="347">Derivatio nominis.</emphasis>
The specific epithet honours Bohdan Bilyj (Etobicoke,
<collectingCountry id="F3267617FFBCE777FB9A277043DAF8A5" box="[1086,1174,1806,1831]" name="Canada" pageId="3" pageNumber="347">Canada</collectingCountry>
), to commemorate his contribution to the study of Nearctic
<taxonomicName id="4C314D04FFBCE777FD90274C45A1F8C9" box="[564,749,1842,1867]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA95FFBCE777FD90274C45A1F8C9" box="[564,749,1842,1867]" italics="true" pageId="3" pageNumber="347">Cladotanytarsus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B650CFFBCE777FF632728430DF859" pageId="3" pageNumber="347" type="diagnosis">
<paragraph id="8B8E3687FFBCE777FF632728430DF859" blockId="3.[151,1437,1662,2011]" pageId="3" pageNumber="347">
<emphasis id="B945EA95FFBCE777FF632728460DF8ED" bold="true" box="[199,321,1878,1903]" pageId="3" pageNumber="347">Diagnosis.</emphasis>
Anal point strongly elongate, evenly tapering toward apex. Superior volsella slender, narrowed at mid length, with field of microtrichia on basal part dorsally. Stem of median volsella
<emphasis id="B945EA95FFBCE777FBEC27054326F810" box="[1096,1130,1915,1938]" italics="true" pageId="3" pageNumber="347">ca.</emphasis>
40 µm long, longer than its setiform and 78 furcate lamellae, all arranged evenly on median and apical part of stem. Inferior volsella with lateral knee-like extension at base, with well-developed angular dorsomedian ridge.
</paragraph>
</subSubSection>
<caption id="DF4E660FFFBBE770FF33222944EDFDEF" box="[151,929,599,621]" httpUri="https://zenodo.org/record/376429/files/figure.png" pageId="4" pageNumber="348" targetBox="[151,1436,193,578]" targetPageId="4">
<paragraph id="8B8E3687FFBBE770FF33222944EDFDEF" blockId="4.[151,929,599,621]" box="[151,929,599,621]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF332229465EFDEE" bold="true" box="[151,274,599,621]" pageId="4" pageNumber="348">FIGURE 2.</emphasis>
<taxonomicName id="4C314D04FFBBE770FEBD222945F1FDEF" authority="Gilka et Puchalski" authorityName="Gilka et Puchalski" box="[281,701,599,621]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="bilyji" status="sp. nov.">
<emphasis id="B945EA95FFBBE770FEBD222946B6FDEF" box="[281,506,599,621]" italics="true" pageId="4" pageNumber="348">Cladotanytarsus bilyji</emphasis>
Giłka
<emphasis id="B945EA95FFBBE770FD9B2226451DFDEF" box="[575,593,600,621]" italics="true" pageId="4" pageNumber="348">et</emphasis>
Puchalski
</taxonomicName>
,
<emphasis id="B945EA95FFBBE770FD6322294454FDEE" bold="true" box="[711,792,599,620]" pageId="4" pageNumber="348">
<taxonomicNameLabel id="A27657EEFFBBE770FD6322294454FDEE" box="[711,792,599,620]" pageId="4" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. Wing.
</paragraph>
</caption>
<subSubSection id="C32B650CFFBBE770FF6322E44427F8EF" pageId="4" pageNumber="348" type="description">
<paragraph id="8B8E3687FFBBE770FF6322E44573FD30" blockId="4.[151,1436,666,1161]" box="[199,575,666,691]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF6322E4461DFD31" bold="true" box="[199,337,666,691]" pageId="4" pageNumber="348">Description</emphasis>
.
<emphasis id="B945EA95FFBBE770FEFB22E446A9FD31" bold="true" box="[351,485,666,691]" pageId="4" pageNumber="348">Adult male</emphasis>
(n = 2).
</paragraph>
<paragraph id="8B8E3687FFBBE770FF6322C3464BFCC1" blockId="4.[151,1436,666,1161]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF6322C3461CFD54" box="[199,336,701,726]" italics="true" pageId="4" pageNumber="348">Colouration</emphasis>
(slide-mounted specimens). Eyes black. Antennal pedicel, tentorium, scutal stripes, scutellum, postnotum and sternum dark brown. Head capsule, antennal flagellum, mouthparts and abdomen greenish to light brown; hypopygium and legs slightly darker. Wing membrane transparent with greenish undertone, veins brownish.
</paragraph>
<paragraph id="8B8E3687FFBBE770FF6323334456FC2C" blockId="4.[151,1436,666,1161]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF6323334648FCE4" box="[199,260,845,870]" italics="true" pageId="4" pageNumber="348">Head</emphasis>
. Eyes reniform, broadly separated. Antenna with 13 flagellomeres, AR 0.920.94; plume fullydeveloped. Frontal tubercles cylindrical, 1224 µm long. Lengths of palpomeres 25 (in µm): 3236, 84104, 100 108, 164 (n = 1). Clypeus semicircular, with 1415 setae.
</paragraph>
<paragraph id="8B8E3687FFBBE770FF6323C445A2FC51" blockId="4.[151,1436,666,1161]" box="[199,750,954,979]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF6323C446DAFC51" box="[199,406,954,979]" italics="true" pageId="4" pageNumber="348">Thorax chaetotaxy</emphasis>
. Ac 78, Dc 8, Pa 1, Scts 46.
</paragraph>
<paragraph id="8B8E3687FFBBE770FF6323A14343FB98" blockId="4.[151,1436,666,1161]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF6323A147B3FC74" box="[199,255,991,1014]" italics="true" pageId="4" pageNumber="348">Wing</emphasis>
(
<figureCitation id="130A2A02FFBBE770FEAB23A3461FFC74" box="[271,339,989,1014]" captionStart="FIGURE 2" captionStartId="4.[151,250,599,621]" captionTargetBox="[151,1436,193,578]" captionTargetId="figure@4.[151,1436,193,578]" captionTargetPageId="4" captionText="FIGURE 2. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. Wing." httpUri="https://zenodo.org/record/376429/files/figure.png" pageId="4" pageNumber="348">Fig. 2</figureCitation>
). Length 15201720 µm. Shape, venation pattern and chaetotaxy typical of the genus, as shown in
<figureCitation id="130A2A02FFBBE770FF33247C4797FB98" box="[151,219,1026,1051]" captionStart="FIGURE 2" captionStartId="4.[151,250,599,621]" captionTargetBox="[151,1436,193,578]" captionTargetId="figure@4.[151,1436,193,578]" captionTargetPageId="4" captionText="FIGURE 2. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. Wing." httpUri="https://zenodo.org/record/376429/files/figure.png" pageId="4" pageNumber="348">Fig. 2</figureCitation>
; Cu1, R1 and m3+4 bare or with few macrotrichia at most; VR Cu 1.181.28.
</paragraph>
<paragraph id="8B8E3687FFBBE770FF632454438DFB0B" blockId="4.[151,1436,666,1161]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF63245447B0FBC3" box="[199,252,1066,1089]" italics="true" pageId="4" pageNumber="348">Legs</emphasis>
. Fore leg tibia with straight spur
<emphasis id="B945EA95FFBBE770FD28245445E2FBC3" box="[652,686,1066,1089]" italics="true" pageId="4" pageNumber="348">ca.</emphasis>
20 µm long. Combs of mid and hind leg tibiae separated, each bearing straight spur:
<emphasis id="B945EA95FFBBE770FE34243046FDFBE7" box="[400,433,1102,1125]" italics="true" pageId="4" pageNumber="348">ca.</emphasis>
1520 µm long on mid leg and up to
<emphasis id="B945EA95FFBBE770FCC1243044CBFBE7" box="[869,903,1102,1125]" italics="true" pageId="4" pageNumber="348">ca.</emphasis>
30 µm long on hind leg. Basitarsus of mid leg with 24 hook-shaped sensilla chaetica. For lengths of leg segments and leg ratios, see
<tableCitation id="C6B3033CFFBBE770FBCC240E43F1FB0B" box="[1128,1213,1136,1161]" captionStart="TABLE 2" captionStartId="4.[151,239,1208,1231]" captionTargetBox="[159,1406,1291,1448]" captionTargetPageId="4" captionText="TABLE 2. Leg segment lengths (µm) and leg ratios of male Cladotanytarsus bilyji sp. nov. (n = 1 when segments missing or deformed)." httpUri="http://table.plazi.org/id/DF4E660FFFBBE770FF3324C646C8FB6D" pageId="4" pageNumber="348" tableUuid="DF4E660FFFBBE770FF3324C646C8FB6D">Table 2</tableCitation>
.
</paragraph>
<caption id="DF4E660FFFBBE770FF3324C646C8FB6D" ID-Table-UUID="DF4E660FFFBBE770FF3324C646C8FB6D" httpUri="http://table.plazi.org/id/DF4E660FFFBBE770FF3324C646C8FB6D" pageId="4" pageNumber="348" targetBox="[159,1406,1291,1448]" targetIsTable="true" targetPageId="4">
<paragraph id="8B8E3687FFBBE770FF3324C646C8FB6D" blockId="4.[151,1436,1207,1263]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF3324C64640FB4D" bold="true" box="[151,268,1208,1231]" pageId="4" pageNumber="348">TABLE 2.</emphasis>
Leg segment lengths (µm) and leg ratios of male
<taxonomicName id="4C314D04FFBBE770FCE024C9437AFB4D" box="[836,1078,1207,1231]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="bilyji" status="sp. nov.">
<emphasis id="B945EA95FFBBE770FCE024C9437AFB4D" box="[836,1078,1207,1231]" italics="true" pageId="4" pageNumber="348">Cladotanytarsus bilyji</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657EEFFBBE770FBE424C643D5FB4D" box="[1088,1177,1208,1231]" pageId="4" pageNumber="348" rank="species">
<emphasis id="B945EA95FFBBE770FBE424C643D5FB4D" bold="true" box="[1088,1177,1208,1231]" pageId="4" pageNumber="348">sp. nov.</emphasis>
</taxonomicNameLabel>
(n = 1 when segments missing or deformed).
</paragraph>
</caption>
<paragraph id="8B8E3687FFBBE770FE8A25754208FA27" pageId="4" pageNumber="348">
<table id="F931C427FFBB188BFF3B25754232FA2A" box="[159,1406,1291,1448]" gridcols="9" gridrows="4" pageId="4" pageNumber="348">
<tr id="350134C5FFBB188BFF3B25754232FAA7" box="[159,1406,1291,1317]" gridrow="0" pageId="4" pageNumber="348" rowspan-0="1">
<th id="76D05DB9FFBB188BFE8A257546CBFAA7" box="[302,391,1291,1317]" gridcol="1" gridrow="0" pageId="4" pageNumber="348">fe</th>
<th id="76D05DB9FFBB188BFE192575455AFAA7" box="[445,534,1291,1317]" gridcol="2" gridrow="0" pageId="4" pageNumber="348">Ti</th>
<th id="76D05DB9FFBB188BFDE8257545E9FAA7" box="[588,677,1291,1317]" gridcol="3" gridrow="0" pageId="4" pageNumber="348">ta1</th>
<th id="76D05DB9FFBB188BFD7F2575447FFAA7" box="[731,819,1291,1317]" gridcol="4" gridrow="0" pageId="4" pageNumber="348">ta2</th>
<th id="76D05DB9FFBB188BFCCE2575448EFAA7" box="[874,962,1291,1317]" gridcol="5" gridrow="0" pageId="4" pageNumber="348">ta3</th>
<th id="76D05DB9FFBB188BFC5D25754375FAA7" box="[1017,1081,1291,1317]" gridcol="6" gridrow="0" pageId="4" pageNumber="348">ta4</th>
<th id="76D05DB9FFBB188BFB2D25754384FAA7" box="[1161,1224,1291,1317]" gridcol="7" gridrow="0" pageId="4" pageNumber="348">ta5</th>
<th id="76D05DB9FFBB188BFABC25754232FAA7" box="[1304,1406,1291,1317]" gridcol="8" gridrow="0" pageId="4" pageNumber="348">LR</th>
</tr>
<tr id="350134C5FFBB188BFF3B25494232FAD2" box="[159,1406,1335,1360]" gridrow="1" pageId="4" pageNumber="348">
<th id="76D05DB9FFBB188BFF3B254947FFFAD2" box="[159,179,1335,1360]" gridcol="0" gridrow="1" pageId="4" pageNumber="348">p1</th>
<td id="76D05DB9FFBB188BFE8A254946CBFAD2" box="[302,391,1335,1360]" gridcol="1" gridrow="1" pageId="4" pageNumber="348">627715</td>
<td id="76D05DB9FFBB188BFE192549455AFAD2" box="[445,534,1335,1360]" gridcol="2" gridrow="1" pageId="4" pageNumber="348">369</td>
<td id="76D05DB9FFBB188BFDE8254945E9FAD2" box="[588,677,1335,1360]" gridcol="3" gridrow="1" pageId="4" pageNumber="348">731</td>
<td id="76D05DB9FFBB188BFD7F2549447FFAD2" box="[731,819,1335,1360]" gridcol="4" gridrow="1" pageId="4" pageNumber="348">398</td>
<td id="76D05DB9FFBB188BFCCE2549448EFAD2" box="[874,962,1335,1360]" gridcol="5" gridrow="1" pageId="4" pageNumber="348">399</td>
<td id="76D05DB9FFBB188BFC5D25494375FAD2" box="[1017,1081,1335,1360]" gridcol="6" gridrow="1" pageId="4" pageNumber="348">221</td>
<td id="76D05DB9FFBB188BFB2D25494384FAD2" box="[1161,1224,1335,1360]" gridcol="7" gridrow="1" pageId="4" pageNumber="348">111</td>
<td id="76D05DB9FFBB188BFABC25494232FAD2" box="[1304,1406,1335,1360]" gridcol="8" gridrow="1" pageId="4" pageNumber="348">1.98</td>
</tr>
<tr id="350134C5FFBB188BFF3B251D4232FAFF" box="[159,1406,1379,1405]" gridrow="2" pageId="4" pageNumber="348">
<th id="76D05DB9FFBB188BFF3B251D47FFFAFF" box="[159,179,1379,1405]" gridcol="0" gridrow="2" pageId="4" pageNumber="348">p2</th>
<td id="76D05DB9FFBB188BFE8A251D46CBFAFF" box="[302,391,1379,1405]" gridcol="1" gridrow="2" pageId="4" pageNumber="348">649900</td>
<td id="76D05DB9FFBB188BFE19251D455AFAFF" box="[445,534,1379,1405]" gridcol="2" gridrow="2" pageId="4" pageNumber="348">472642</td>
<td id="76D05DB9FFBB188BFDE8251D45E9FAFF" box="[588,677,1379,1405]" gridcol="3" gridrow="2" pageId="4" pageNumber="348">288295</td>
<td id="76D05DB9FFBB188BFD7F251D447FFAFF" box="[731,819,1379,1405]" gridcol="4" gridrow="2" pageId="4" pageNumber="348">162192</td>
<td id="76D05DB9FFBB188BFCCE251D448EFAFF" box="[874,962,1379,1405]" gridcol="5" gridrow="2" pageId="4" pageNumber="348">118140</td>
<td id="76D05DB9FFBB188BFC5D251D4375FAFF" box="[1017,1081,1379,1405]" gridcol="6" gridrow="2" pageId="4" pageNumber="348">6673</td>
<td id="76D05DB9FFBB188BFB2D251D4384FAFF" box="[1161,1224,1379,1405]" gridcol="7" gridrow="2" pageId="4" pageNumber="348">5270</td>
<td id="76D05DB9FFBB188BFABC251D4232FAFF" box="[1304,1406,1379,1405]" gridcol="8" gridrow="2" pageId="4" pageNumber="348">0.460.61</td>
</tr>
<tr id="350134C5FFBB188BFF3B25F14232FA2A" box="[159,1406,1423,1448]" gridrow="3" pageId="4" pageNumber="348">
<th id="76D05DB9FFBB188BFF3B25F147FFFA2A" box="[159,179,1423,1448]" gridcol="0" gridrow="3" pageId="4" pageNumber="348">p3</th>
<td id="76D05DB9FFBB188BFE8A25F146CBFA2A" box="[302,391,1423,1448]" gridcol="1" gridrow="3" pageId="4" pageNumber="348">704826</td>
<td id="76D05DB9FFBB188BFE1925F1455AFA2A" box="[445,534,1423,1448]" gridcol="2" gridrow="3" pageId="4" pageNumber="348">915964</td>
<td id="76D05DB9FFBB188BFDE825F145E9FA2A" box="[588,677,1423,1448]" gridcol="3" gridrow="3" pageId="4" pageNumber="348">450</td>
<td id="76D05DB9FFBB188BFD7F25F1447FFA2A" box="[731,819,1423,1448]" gridcol="4" gridrow="3" pageId="4" pageNumber="348">288</td>
<td id="76D05DB9FFBB188BFCCE25F1448EFA2A" box="[874,962,1423,1448]" gridcol="5" gridrow="3" pageId="4" pageNumber="348">244</td>
<td id="76D05DB9FFBB188BFC5D25F14375FA2A" box="[1017,1081,1423,1448]" gridcol="6" gridrow="3" pageId="4" pageNumber="348">155</td>
<td id="76D05DB9FFBB188BFB2D25F14384FA2A" box="[1161,1224,1423,1448]" gridcol="7" gridrow="3" pageId="4" pageNumber="348">103</td>
<td id="76D05DB9FFBB188BFABC25F14232FA2A" box="[1304,1406,1423,1448]" gridcol="8" gridrow="3" pageId="4" pageNumber="348">0.65</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8E3687FFBBE770FF6325934427F8EF" blockId="4.[151,1437,1516,1901]" pageId="4" pageNumber="348">
<emphasis id="B945EA95FFBBE770FF632593461AF986" box="[199,342,1517,1540]" italics="true" pageId="4" pageNumber="348">Hypopygium</emphasis>
(
<figureCitation id="130A2A02FFBBE770FEC2259246E0F987" box="[358,428,1516,1541]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="4" pageNumber="348">Fig. 3</figureCitation>
). Gonostylus shorter than gonocoxite,
<emphasis id="B945EA95FFBBE770FCC7259344C8F986" box="[867,900,1517,1540]" italics="true" pageId="4" pageNumber="348">ca.</emphasis>
70 µm long, with blunt apex. Anal tergite with V-type separated bands and 78 median setae placed in two rows. Anal point elongate, evenly tapering towards slender apex (slightly deformed in
<typeStatus id="548A8825FFBBE770FD95264B45DDF9CF" box="[561,657,1589,1613]" pageId="4" pageNumber="348" type="paratype">paratype</typeStatus>
specimen,
<figureCitation id="130A2A02FFBBE770FCBC264A4412F9CF" box="[792,862,1588,1613]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="4" pageNumber="348">Fig. 3</figureCitation>
C), bearing 78 spinulae between well-developed crests, entire area surrounding base of anal point covered with microtrichia (
<figureCitation id="130A2A02FFBBE770FC542626437EF9F2" box="[1008,1074,1624,1649]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="4" pageNumber="348">Fig. 3</figureCitation>
A, C). Superior volsella slender, elongate, more or less narrowed at mid length, bearing field of microtrichia dorsally only on basal part or proximal 1/3 at most, 611 dorsal setae, 12 setae on anteromedian margin and 3 long setae placed on conical tubercles at base. Digitus curved, long, extending far beyond superior volsella, with finger-like tip (
<figureCitation id="130A2A02FFBBE770FB0626BA43A4F95F" box="[1186,1256,1732,1757]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="4" pageNumber="348">Fig. 3</figureCitation>
A, D). Stem of median volsella slightly curved and directed posteriorly, long (
<emphasis id="B945EA95FFBBE770FCF726974438F882" box="[851,884,1769,1792]" italics="true" pageId="4" pageNumber="348">ca.</emphasis>
40 µm), bearing several setiform and 78 furcate lamellae: 4 strong and 34 weaker, all arranged evenly on median and apical part of stem (
<figureCitation id="130A2A02FFBBE770FB10277243B5F8A7" box="[1204,1273,1804,1829]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="4" pageNumber="348">Fig. 3</figureCitation>
B, E). Inferior volsella with lateral knee-like extension at base, evenly curved and directed posteromedially, darkly pigmented dorsomedian ridge distinctly protruding, angular (
<figureCitation id="130A2A02FFBBE770FD62272A444BF8EF" box="[710,775,1876,1901]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="4" pageNumber="348">Fig. 3</figureCitation>
A, B, F).
</paragraph>
</subSubSection>
<caption id="DF4E660FFFBAE771FF3327514266F801" httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="5" pageNumber="349" targetBox="[151,1436,193,1818]" targetPageId="5">
<paragraph id="8B8E3687FFBAE771FF3327514266F801" blockId="5.[151,1436,1839,1923]" pageId="5" pageNumber="349">
<emphasis id="B945EA95FFBAE771FF332751465FF8C6" bold="true" box="[151,275,1839,1861]" pageId="5" pageNumber="349">FIGURE 3.</emphasis>
<taxonomicName id="4C314D04FFBAE771FEBD275145F2F8C7" authority="Gilka et Puchalski" authorityName="Gilka et Puchalski" box="[281,702,1839,1862]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="bilyji" status="sp. nov.">
<emphasis id="B945EA95FFBAE771FEBD275146B7F8C7" box="[281,507,1839,1861]" italics="true" pageId="5" pageNumber="349">Cladotanytarsus bilyji</emphasis>
Giłka
<emphasis id="B945EA95FFBAE771FDE4274F451EF8C4" box="[576,594,1841,1862]" italics="true" pageId="5" pageNumber="349">et</emphasis>
Puchalski
</taxonomicName>
,
<emphasis id="B945EA95FFBAE771FD6C274E4455F8C7" bold="true" box="[712,793,1840,1861]" pageId="5" pageNumber="349">
<taxonomicNameLabel id="A27657EEFFBAE771FD6C274E4455F8C7" box="[712,793,1840,1861]" pageId="5" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B945EA95FFBAE771FCC02751443AF8C6" bold="true" box="[868,886,1839,1860]" pageId="5" pageNumber="349">A</emphasis>
,
<emphasis id="B945EA95FFBAE771FC20274E44D9F8C7" bold="true" box="[900,917,1840,1861]" pageId="5" pageNumber="349">B</emphasis>
—hypopygium in dorsal (A) and ventral aspect (B) (holotype);
<emphasis id="B945EA95FFBAE771FEAB2730466DF8E6" bold="true" box="[271,289,1870,1892]" pageId="5" pageNumber="349">C</emphasis>
—anal point (variation, paratype);
<emphasis id="B945EA95FFBAE771FD2C273145D6F8E6" bold="true" box="[648,666,1871,1892]" pageId="5" pageNumber="349">D</emphasis>
—superior volsella and digitus (variation);
<emphasis id="B945EA95FFBAE771FBFC27314325F8E6" bold="true" box="[1112,1129,1871,1892]" pageId="5" pageNumber="349">E</emphasis>
—median volsella (holotype);
<emphasis id="B945EA95FFBAE771FF33271347EAF800" bold="true" box="[151,166,1901,1922]" pageId="5" pageNumber="349">F</emphasis>
—inferior volsella (paratype) (D—magnified
<emphasis id="B945EA95FFBAE771FDD2271045D9F801" box="[630,661,1902,1923]" italics="true" pageId="5" pageNumber="349">ca.</emphasis>
2 times relative to A; E, F—magnified
<emphasis id="B945EA95FFBAE771FB832710430AF801" box="[1063,1094,1902,1923]" italics="true" pageId="5" pageNumber="349">ca.</emphasis>
3 times relative to A).
</paragraph>
</caption>
<subSubSection id="C32B650CFFB9E772FF6320E942D4FC5D" pageId="6" pageNumber="350" type="discussion">
<paragraph id="8B8E3687FFB9E772FF6320E94687FEB4" blockId="6.[151,1437,151,991]" pageId="6" pageNumber="350">
<emphasis id="B945EA95FFB9E772FF6320E94677FF32" bold="true" box="[199,315,151,176]" pageId="6" pageNumber="350">Remarks.</emphasis>
Apart from the strongly elongate anal tergite point, this new species can be distinguished from other
<taxonomicName id="4C314D04FFB9E772FF3320C7461CFF50" box="[151,336,185,210]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA95FFB9E772FF3320C7461CFF50" box="[151,336,185,210]" italics="true" pageId="6" pageNumber="350">Cladotanytarsus</emphasis>
</taxonomicName>
on the following combination of characters: the superior volsella slender and narrowed at mid length, with field of microtrichia only on basal part dorsally, the median volsella composed of a long stem and 78 furcate lamellae and the inferior volsella with a lateral knee-like extension at the base, bearing a well-developed angular dorsomedian ridge.
</paragraph>
<paragraph id="8B8E3687FFB9E772FF63214042D4FC5D" blockId="6.[151,1437,151,991]" pageId="6" pageNumber="350">
According to preliminary determinations (Sublette, Saether; pers. comm.), there are at least 8 or 9 additional
<taxonomicName id="4C314D04FFB9E772FF33211C461CFEF9" box="[151,336,354,379]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA95FFB9E772FF33211C461CFEF9" box="[151,336,354,379]" italics="true" pageId="6" pageNumber="350">Cladotanytarsus</emphasis>
</taxonomicName>
species recorded from the Lake Winnipeg region (
<emphasis id="B945EA95FFB9E772FC00211C4375FEF9" box="[932,1081,354,379]" italics="true" pageId="6" pageNumber="350">locus typicus</emphasis>
of
<taxonomicName id="4C314D04FFB9E772FBC1211D438FFEF9" box="[1125,1219,354,379]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="bilyji">
<emphasis id="B945EA95FFB9E772FBC1211D438FFEF9" box="[1125,1219,354,379]" italics="true" pageId="6" pageNumber="350">C. bilyji</emphasis>
</taxonomicName>
), which should be compared with those described by
<bibRefCitation id="EFA04B76FFB9E772FD9221F8441FFE1D" author="Bilyj" box="[566,851,390,415]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj and Davies (1989)</bibRefCitation>
from northwestern
<collectingRegion id="49F5F865FFB9E772FBE421F843D5FE1D" box="[1088,1177,390,415]" country="Canada" name="Ontario" pageId="6" pageNumber="350">Ontario</collectingRegion>
. We examined all the
<taxonomicName id="4C314D04FFB9E772FF3321D4461CFE41" box="[151,336,426,451]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B945EA95FFB9E772FF3321D4461CFE41" box="[151,336,426,451]" italics="true" pageId="6" pageNumber="350">Cladotanytarsus</emphasis>
</taxonomicName>
species by Bilyj (
<emphasis id="B945EA95FFB9E772FDB921D24526FE41" box="[541,618,428,451]" italics="true" pageId="6" pageNumber="350">op. cit.</emphasis>
) known from adult males (excl.
<taxonomicName id="4C314D04FFB9E772FC7921D54257FE41" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" box="[989,1307,426,451]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="aeiparthenus">
<emphasis id="B945EA95FFB9E772FC7921D543DCFE41" box="[989,1168,426,451]" italics="true" pageId="6" pageNumber="350">C. aeiparthenus</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FB3D21D44257FE41" author="Bilyj" box="[1177,1307,426,451]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
). However, even when atypical variations are included, most of the males examined (except for
<taxonomicName id="4C314D04FFB9E772FBF521B14220FE65" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" box="[1105,1388,462,487]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="muricatus">
<emphasis id="B945EA95FFB9E772FBF521B143AEFE64" box="[1105,1250,463,486]" italics="true" pageId="6" pageNumber="350">C. muricatus</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FB4E21B04220FE65" author="Bilyj" box="[1258,1388,462,487]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
, see remarks below) do not fully fit the present concept of the strongly elongate and narrow hypopygial anal point. Males of those species have relatively short anal points (
<taxonomicName id="4C314D04FFB9E772FC972269437AFDAD" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" box="[819,1078,534,559]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="daviesi">
<emphasis id="B945EA95FFB9E772FC97226944EAFDAD" box="[819,934,534,559]" italics="true" pageId="6" pageNumber="350">C. daviesi</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FC162268437AFDAD" author="Bilyj" box="[946,1078,534,559]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C314D04FFB9E772FBE1226942C3FDAD" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" box="[1093,1423,534,559]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="pinnaticornis">
<emphasis id="B945EA95FFB9E772FBE1226943B3FDAC" box="[1093,1279,535,558]" italics="true" pageId="6" pageNumber="350">C. pinnaticornis</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FAAD226842C3FDAD" author="Bilyj" box="[1289,1423,534,559]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
), without a distinct distal elongation and/or at least distinctly broadened at the base. Males of
<taxonomicName id="4C314D04FFB9E772FB38224542D0FDD1" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" box="[1180,1436,570,595]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="elaensis">
<emphasis id="B945EA95FFB9E772FB382245425BFDD1" box="[1180,1303,570,595]" italics="true" pageId="6" pageNumber="350">C. elaensis</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FABA224442D0FDD1" author="Bilyj" box="[1310,1436,570,595]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
have acute anal points, but are relatively broad and surrounded with a microtrichia-free area at the base; in
<taxonomicName id="4C314D04FFB9E772FA2622214638FD19" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="tribelus">
<emphasis id="B945EA95FFB9E772FA26222147A1FD19" italics="true" pageId="6" pageNumber="350">C. tribelus</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FF5022FC4638FD19" author="Bilyj" box="[244,372,642,667]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
the anal point is narrow and parallel-sided but moderately long (
<bibRefCitation id="EFA04B76FFB9E772FBF322FC47EBFD3D" author="Bilyj" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj &amp; Davies 1989; figs 6, 8</bibRefCitation>
). As far as the general shape of the anal point is concerned, there is a slight resemblance between the males of
<taxonomicName id="4C314D04FFB9E772FA2622D946C7FD61" authority="Bilyj, 1989" authorityName="Bilyj" authorityYear="1989" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="fusiformis">
<emphasis id="B945EA95FFB9E772FA2622D9464BFD61" italics="true" pageId="6" pageNumber="350">C. fusiformis</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FEA922B446C7FD61" author="Bilyj" box="[269,395,714,739]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj, 1989</bibRefCitation>
</taxonomicName>
and those of
<taxonomicName id="4C314D04FFB9E772FD8422B54535FD61" box="[544,633,714,739]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="bilyji">
<emphasis id="B945EA95FFB9E772FD8422B54535FD61" box="[544,633,714,739]" italics="true" pageId="6" pageNumber="350">C. bilyji</emphasis>
</taxonomicName>
; although these two species can be easily separated also by the shape of the superior volsella, which in the former is nearly half as long as the digitus, and the median volsella has a smaller number of lamellae (
<emphasis id="B945EA95FFB9E772FEDB236D46D7FCA9" box="[383,411,787,811]" italics="true" pageId="6" pageNumber="350">cf.</emphasis>
<figureCitation id="130A2A02FFB9E772FE04236C46ADFCA9" box="[416,481,786,811]" captionStart="FIGURE 3" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[151,1436,193,1818]" captionTargetId="figure@5.[151,1436,193,1818]" captionTargetPageId="5" captionText="FIGURE 3. Cladotanytarsus bilyji Giłka et Puchalski, sp. nov., male. A, B — hypopygium in dorsal (A) and ventral aspect (B) (holotype); C — anal point (variation, paratype); D — superior volsella and digitus (variation); E — median volsella (holotype); F — inferior volsella (paratype) (D—magnified ca. 2 times relative to A; E, F—magnified ca. 3 times relative to A)." httpUri="https://zenodo.org/record/376430/files/figure.png" pageId="6" pageNumber="350">Fig. 3</figureCitation>
and
<bibRefCitation id="EFA04B76FFB9E772FDB2236C45B3FCA9" author="Bilyj" box="[534,767,786,811]" pageId="6" pageNumber="350" refString="Bilyj, B. &amp; Davies, I. J. (1989) Descriptions and ecological notes on seven new species of Cladotanytarsus (Chironomidae: Diptera) collected from an experimentally acidified lake. Canadian Journal of Zoology, 67, 948 - 962. https: // doi. org / 10.1139 / z 89 - 138" type="journal article" year="1989">Bilyj &amp; Davies 1989</bibRefCitation>
, fig. 4). Originally,
<taxonomicName id="4C314D04FFB9E772FC7E236D4326FCA9" box="[986,1130,787,811]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="fusiformis">
<emphasis id="B945EA95FFB9E772FC7E236D4326FCA9" box="[986,1130,787,811]" italics="true" pageId="6" pageNumber="350">C. fusiformis</emphasis>
</taxonomicName>
was also compared with
<taxonomicName id="4C314D04FFB9E772FA26236D4554FCCD" authority="Goetghebuer, 1922" authorityName="Goetghebuer" authorityYear="1922" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="nigrovittatus">
<emphasis id="B945EA95FFB9E772FA26236D4669FCCC" italics="true" pageId="6" pageNumber="350">C. nigrovittatus</emphasis>
(
<bibRefCitation id="EFA04B76FFB9E772FE902348455CFCCD" author="Goetghebuer" box="[308,528,822,847]" pageId="6" pageNumber="350" refString="Goetghebuer, M. (1922) Nouveaux materiaux pour l'etude de la fauna de Chironomides de Belgique. Annales de Biologie Lacustre, 11, 38 - 62." type="journal article" year="1922">Goetghebuer, 1922</bibRefCitation>
)
</taxonomicName>
. The latter species is presumed to be the closest relative of
<taxonomicName id="4C314D04FFB9E772FB1023494242FCCD" box="[1204,1294,822,847]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="bilyji">
<emphasis id="B945EA95FFB9E772FB1023494242FCCD" box="[1204,1294,822,847]" italics="true" pageId="6" pageNumber="350">C. bilyji</emphasis>
</taxonomicName>
(see the key below). Finally, the males of
<taxonomicName id="4C314D04FFB9E772FE5323254467FCF1" authority="Walker, 1856" authorityName="Walker" authorityYear="1856" box="[503,811,858,883]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="mancus">
<emphasis id="B945EA95FFB9E772FE532325453EFCF1" box="[503,626,859,883]" italics="true" pageId="6" pageNumber="350">C. mancus</emphasis>
(Walker, 1856)
</taxonomicName>
[or the
<taxonomicName id="4C314D04FFB9E772FC3623254341FCF1" box="[914,1037,859,883]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="mancus">
<emphasis id="B945EA95FFB9E772FC3623254341FCF1" box="[914,1037,859,883]" italics="true" pageId="6" pageNumber="350">C. mancus</emphasis>
</taxonomicName>
group
<emphasis id="B945EA95FFB9E772FBC3232243E9FCF1" box="[1127,1189,860,883]" italics="true" pageId="6" pageNumber="350">sensu</emphasis>
<bibRefCitation id="EFA04B76FFB9E772FB1523244207FCF1" author="Gilka" box="[1201,1355,858,883]" pageId="6" pageNumber="350" refString="Gilka, W. (2001) A review of Polish Cladotanytarsus Kieffer (Diptera: Chironomidae) with description of three new species. Polish Journal of Entomology, 70, 307 - 328." type="journal article" year="2001">Giłka (2001)</bibRefCitation>
], often misidentified with
<taxonomicName id="4C314D04FFB9E772FEDE2301457DFC14" box="[378,561,895,918]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="nigrovittatus">
<emphasis id="B945EA95FFB9E772FEDE2301457DFC14" box="[378,561,895,918]" italics="true" pageId="6" pageNumber="350">C. nigrovittatus</emphasis>
</taxonomicName>
, should also be included in this comparison. They differ in the length proportions of the median volsella stem-lamellae (the stem longer relative to its lamellae in
<taxonomicName id="4C314D04FFB9E772FB0D23DD4248FC39" box="[1193,1284,930,955]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="bilyji">
<emphasis id="B945EA95FFB9E772FB0D23DD4248FC39" box="[1193,1284,930,955]" italics="true" pageId="6" pageNumber="350">C. bilyji</emphasis>
</taxonomicName>
), the number of lamellae (greater in
<taxonomicName id="4C314D04FFB9E772FE3223B946BCFC5D" box="[406,496,966,991]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="bilyji">
<emphasis id="B945EA95FFB9E772FE3223B946BCFC5D" box="[406,496,966,991]" italics="true" pageId="6" pageNumber="350">C. bilyji</emphasis>
</taxonomicName>
), and the arrangement of median setae on the anal tergite (dispersed in
<taxonomicName id="4C314D04FFB9E772FAB523B942C0FC5D" box="[1297,1420,967,991]" class="Insecta" family="Chironomidae" genus="Cladotanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="mancus">C. mancus</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>