391 lines
60 KiB
XML
391 lines
60 KiB
XML
<document id="00CEA2CCEAFD6FE9A7A4BEBF191264D1" ID-CLB-Dataset="5696" ID-DOI="10.11646/zootaxa.4985.3.6" ID-GBIF-Dataset="ac091d94-86ff-45b9-9eb3-a00c8b173096" ID-ISSN="1175-5326" ID-Zenodo-Dep="4956267" ID-ZooBank="D0116147-5733-4276-9DC8-95FBE3762BD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623759304789" checkinUser="plazi" docAuthor="Techakijvej, Chotiwut, Sareein, Nattawut, Hwang, Jeong Mi, Bae, Yeon Jae & Phalaraksh, Chitchol" docDate="2021" docId="03966810F20CFF8D95D4D4245570F9B2" docLanguage="en" docName="zootaxa.4985.3.6.pdf" docOrigin="Zootaxa 4985 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ephoron ookaewae Techakijvej & Sareein & Hwang & Bae & Phalaraksh 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="398" masterDocId="FFAF1068F20EFF8B9543D606540AFFDE" masterDocTitle="A new species of Ephoron Williamson, 1802 (Ephemeroptera: Polymitarcyidae) from Thailand" masterLastPageNumber="402" masterPageNumber="392" pageNumber="394" updateTime="1699016653363" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="F70287FD768FDB16A6261DD05BDEA0EB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="23AC1D7F238F69DFC7D73EFEC46A363C">
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<mods:title id="C8BA5EB56AA5E0C381C9A06B6976F4F8">A new species of Ephoron Williamson, 1802 (Ephemeroptera: Polymitarcyidae) from Thailand</mods:title>
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<mods:name id="B7E012C52088C5AA3C1788130DEDB4BD" type="personal">
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<mods:roleTerm id="AB36EF4B6F294626FFE56C8CCAE07EBF">Author</mods:roleTerm>
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<mods:namePart id="C5F732D1470EF9B2C256363E3C16A6BE">Techakijvej, Chotiwut</mods:namePart>
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<mods:affiliation id="F40A3F7496A22BE1694180AD48028F06">PhD Degree Program in Environmental Science, Environmental Science Research Center, Faculty of Science, Chiang Mai University,</mods:affiliation>
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<mods:roleTerm id="F431F440C9DD4BBAFEDB0B4C713BE706">Author</mods:roleTerm>
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<mods:namePart id="0443F7EEC82CD0A454FE9BBC32181E96">Sareein, Nattawut</mods:namePart>
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<mods:nameIdentifier id="7092A591A8E609FB40E9B89BFF702680" type="ORCID">0000-0003-4167-8902</mods:nameIdentifier>
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<mods:affiliation id="BBD6F4FA52A42C65E4E29AB54A7C11FC">Environmental Science Research Center, Faculty of Science, Chiang Mai University, Chiang Mai, 50200, Thailand. & https: // orcid. org / 0000 - 0003 - 4167 - 8902</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="1007CEE612AC8EF7E58460703595E44D">Author</mods:roleTerm>
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<mods:namePart id="038428E4BA2E5B1C4A1E7F16E8EEC4C0">Hwang, Jeong Mi</mods:namePart>
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<mods:nameIdentifier id="0E9E3413128FB86B18E276CE3AC746A1" type="ORCID">0000-0003-4743-4474</mods:nameIdentifier>
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<mods:affiliation id="4892B05ADB4883828838CBE5478C4ED0">Korean Entomological Institute, College of Life Sciences and Biotechnology, Korea University, Seoul 02841, Korea. & https: // orcid. org / 0000 - 0003 - 4743 - 4474</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="A2ADBCA3BA7C5943A92E09AC5CF11E05">Author</mods:roleTerm>
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<mods:namePart id="BCC150463D5F4B9F224401B5020844C0">Bae, Yeon Jae</mods:namePart>
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<mods:nameIdentifier id="1BA46FE25A05F536932F09A6E9465351" type="ORCID">0000-0001-7810-5409</mods:nameIdentifier>
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<mods:affiliation id="E755F6FAFB68103941EB82B548836269">Korean Entomological Institute, College of Life Sciences and Biotechnology, Korea University, Seoul 02841, Korea. & Department of Environmental Science and Ecological Engineering, Graduate School, Korea University, Seoul 02841, Korea. https: // orcid. org / 0000 - 0001 - 7810 - 5409</mods:affiliation>
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<mods:namePart id="AEE00B5A6D8DA7D1DAECF575A5FEDFD2">Phalaraksh, Chitchol</mods:namePart>
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<mods:affiliation id="EB4D26FA979CBEF795BD722132FE4EE1">Environmental Science Research Center, Faculty of Science, Chiang Mai University, Chiang Mai, 50200, Thailand. & Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, 50200, Thailand.</mods:affiliation>
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<mods:title id="3326B71B7337260CE0381ECD2D72BCED">Zootaxa</mods:title>
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<mods:part id="AC4D8FE03F6D7FD1658ED99B9DC67B66">
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<mods:date id="D5E5BB095108637793E997654AFCB68A">2021</mods:date>
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<mods:number id="4A89D6310860F06F4449FCB6F067EC83">2021-06-15</mods:number>
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<mods:number id="6FC2DAD232CFB6B0980C393AEDCC0D58">4985</mods:number>
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<treatment id="03966810F20CFF8D95D4D4245570F9B2" ID-DOI="http://doi.org/10.5281/zenodo.5074203" ID-GBIF-Taxon="183336377" ID-Zenodo-Dep="5074203" LSID="urn:lsid:plazi:treatment:03966810F20CFF8D95D4D4245570F9B2" httpUri="http://treatment.plazi.org/id/03966810F20CFF8D95D4D4245570F9B2" lastPageId="6" lastPageNumber="398" pageId="2" pageNumber="394">
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<subSubSection id="C3258A8DF20CFF8995D4D4245736FDE3" box="[151,828,546,573]" pageId="2" pageNumber="394" type="nomenclature">
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<paragraph id="8B80D906F20CFF8995D4D4245736FDE3" blockId="2.[151,828,546,609]" box="[151,828,546,573]" pageId="2" pageNumber="394">
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<heading id="D0C86E6AF20CFF8995D4D4245736FDE3" bold="true" box="[151,828,546,573]" fontSize="11" level="1" pageId="2" pageNumber="394" reason="1">
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<emphasis id="B94B0514F20CFF8995D4D4245736FDE3" bold="true" box="[151,828,546,573]" pageId="2" pageNumber="394">
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<taxonomicName id="4C3FA285F20CFF8995D4D42456D8FDE3" authority="Techakijvej & Phalaraksh" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[151,722,546,573]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
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<emphasis id="B94B0514F20CFF8995D4D424558BFDE3" bold="true" box="[151,385,546,573]" italics="true" pageId="2" pageNumber="394">Ephoron ookaewae</emphasis>
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Techakijvej & Phalaraksh
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</taxonomicName>
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,
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<taxonomicNameLabel id="A278B86FF20CFF89979CD4255736FDE3" box="[735,828,547,573]" pageId="2" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3258A8DF20CFF8995D4D4415518FDBF" box="[151,274,583,609]" pageId="2" pageNumber="394" type="description">
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<paragraph id="8B80D906F20CFF8995D4D4415518FDBF" blockId="2.[151,828,546,609]" box="[151,274,583,609]" pageId="2" pageNumber="394">
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(
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<figureCitation id="1304C583F20CFF8995DCD4415500FDBF" box="[159,266,583,609]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1616,1640]" captionStartId-1="4.[151,250,1941,1965]" captionStartId-2="5.[151,250,1954,1978]" captionStartId-3="6.[151,250,1152,1176]" captionTargetBox-0="[151,1426,273,1593]" captionTargetBox-1="[158,1429,181,1918]" captionTargetBox-2="[224,1362,181,1930]" captionTargetBox-3="[151,1436,489,1128]" captionTargetId-0="figure-74@3.[151,1436,273,1593]" captionTargetId-1="figure-18@4.[158,1429,181,1918]" captionTargetId-2="figure-30@5.[224,1362,181,1930]" captionTargetId-3="figure-183@6.[151,1436,489,1128]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." captionText-1="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." captionText-2="FIGURE 3. Ephoron ookaewae sp. nov., mature larva (SEM): a, apex of right mandible; b, same, detail of small triangular spine (arrow)." captionText-3="FIGURE 4. Ephoron ookaewae sp. nov., eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide)." figureDoi-0="http://doi.org/10.5281/zenodo.4956271" figureDoi-1="http://doi.org/10.5281/zenodo.4956273" figureDoi-2="http://doi.org/10.5281/zenodo.4956275" figureDoi-3="http://doi.org/10.5281/zenodo.4956277" httpUri-0="https://zenodo.org/record/4956271/files/figure.png" httpUri-1="https://zenodo.org/record/4956273/files/figure.png" httpUri-2="https://zenodo.org/record/4956275/files/figure.png" httpUri-3="https://zenodo.org/record/4956277/files/figure.png" pageId="2" pageNumber="394">Figs. 1–4</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3258A8DF20CFF8995D4D4885193FC83" pageId="2" pageNumber="394" type="diagnosis">
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<paragraph id="8B80D906F20CFF8995D4D4885193FC83" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="394">
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<emphasis id="B94B0514F20CFF8995D4D4885518FD76" bold="true" box="[151,274,654,680]" pageId="2" pageNumber="394">Diagnosis.</emphasis>
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<taxonomicName id="4C3FA285F20CFF899459D48955F9FD77" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[282,499,655,681]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
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<emphasis id="B94B0514F20CFF899459D48955F9FD77" box="[282,499,655,681]" italics="true" pageId="2" pageNumber="394">Ephoron ookaewae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A278B86FF20CFF8994B8D4895659FD77" box="[507,595,655,681]" pageId="2" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from the other species of the genus by the following characters, in the larvae: 5–7 spike-like tubercles on dorsal side of mandibular tusk, mandibular tusk not curved inwards as much as in other species, mandible with small triangular shaped spine between outer and inner incisors, head with three to four-pointed median frontal process. In the adult: male and female forewing with
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<specimenCount id="9D39128FF20CFF8991BDD4FD5196FCCB" box="[1278,1436,763,789]" pageId="2" pageNumber="394" type="intercalary">4 intercalaries</specimenCount>
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between CuA and CuP, male hindwings with MA forked near middle of wing. And in the eggs: acorn shaped with concave indentation at posterior area, chorion sculptured in hexagonal netted pattern, convex area inside each cell.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3258A8DF20CFF899584D560566DFB87" pageId="2" pageNumber="394" type="materials_examined">
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<paragraph id="8B80D906F20CFF899584D560566DFB87" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="394">
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<emphasis id="B94B0514F20CFF899584D560553CFC5E" bold="true" box="[199,310,870,896]" pageId="2" pageNumber="394">Material.</emphasis>
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<materialsCitation id="3B57D35BF20CFF899402D5615069FC17" ID-GBIF-Occurrence="3314062303" collectingDate="2018-03-04" collectionCode="FBRL" collectorName="C. Techakijvej" country="Thailand" county="Ping River" elevation="59" latitude="15.752944" location="Freshwater Biomonitor Research Laboratory" longLatPrecision="1" longitude="100.08614" municipality="Nakonsawan Province" pageId="2" pageNumber="394" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
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<emphasis id="B94B0514F20CFF899402D56155A4FC5F" bold="true" box="[321,430,871,897]" pageId="2" pageNumber="394">
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<typeStatus id="548467A4F20CFF899402D56155A4FC5F" box="[321,430,871,897]" pageId="2" pageNumber="394" type="holotype">Holotype</typeStatus>
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</emphasis>
|
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(
|
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<location id="8EE08FDDF20CFF899483D56157CCFC5F" LSID="urn:lsid:plazi:treatment:03966810F20CFF8D95D4D4245570F9B2:8EE08FDDF20CFF899483D56157CCFC5F" box="[448,966,871,897]" country="Thailand" county="Ping River" latitude="15.752944" longLatPrecision="1" longitude="100.08614" municipality="Nakonsawan Province" name="Freshwater Biomonitor Research Laboratory" pageId="2" pageNumber="394" stateProvince="Chiang Mai">Freshwater Biomonitor Research Laboratory</location>
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:
|
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<collectionCode id="ED2E41C3F20CFF899697D5615017FC5F" box="[980,1053,871,897]" pageId="2" pageNumber="394">FBRL</collectionCode>
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,
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<location id="8EE08FDDF20CFF899168D561510FFC5F" LSID="urn:lsid:plazi:treatment:03966810F20CFF8D95D4D4245570F9B2:8EE08FDDF20CFF899168D561510FFC5F" box="[1067,1285,871,897]" country="Thailand" county="Ping River" latitude="15.752944" longLatPrecision="1" longitude="100.08614" municipality="Nakonsawan Province" name="Faculty of Science" pageId="2" pageNumber="394" stateProvince="Chiang Mai">Faculty of Science</location>
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,
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<collectingRegion id="49FB17E4F20CFF899057D5615196FC5F" box="[1300,1436,871,897]" country="Thailand" name="Chiang Mai" pageId="2" pageNumber="394">Chiang Mai</collectingRegion>
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<location id="8EE08FDDF20CFF8995D4D58D5504FC7B" LSID="urn:lsid:plazi:treatment:03966810F20CFF8D95D4D4245570F9B2:8EE08FDDF20CFF8995D4D58D5504FC7B" box="[151,270,907,933]" country="Thailand" county="Ping River" latitude="15.752944" longLatPrecision="1" longitude="100.08614" municipality="Nakonsawan Province" name="University" pageId="2" pageNumber="394" stateProvince="Chiang Mai">University</location>
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,
|
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<collectingRegion id="49FB17E4F20CFF89945AD58D5595FC7B" box="[281,415,907,933]" country="Thailand" name="Chiang Mai" pageId="2" pageNumber="394">Chiang Mai</collectingRegion>
|
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,
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<location id="8EE08FDDF20CFF8994EAD58D5604FC78" LSID="urn:lsid:plazi:treatment:03966810F20CFF8D95D4D4245570F9B2:8EE08FDDF20CFF8994EAD58D5604FC78" box="[425,526,907,934]" country="Thailand" county="Ping River" latitude="15.752944" longLatPrecision="1" longitude="100.08614" municipality="Nakonsawan Province" name="Thailand" pageId="2" pageNumber="394" stateProvince="Chiang Mai">Thailand</location>
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||
) male imago (in alcohol) from
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<emphasis id="B94B0514F20CFF899624D58C57DEFC7A" bold="true" box="[871,980,906,932]" pageId="2" pageNumber="394">
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<collectingCountry id="F3289996F20CFF899624D58C57DEFC7A" box="[871,980,906,932]" name="Thailand" pageId="2" pageNumber="394">Thailand</collectingCountry>
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</emphasis>
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:
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<collectingMunicipality id="6BE4437CF20CFF8996A1D58D50D4FC7B" box="[994,1246,907,933]" pageId="2" pageNumber="394">Nakonsawan Province</collectingMunicipality>
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,
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<collectingCounty id="62E1A18AF20CFF8991AAD58D516AFC7B" box="[1257,1376,907,933]" pageId="2" pageNumber="394">Ping River</collectingCounty>
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(
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<figureCitation id="1304C583F20CFF89902CD58D54B8FC17" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1a</figureCitation>
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||
),
|
||
<elevation id="00123E35F20CFF89958AD5A85535FC17" box="[201,319,942,969]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="394" unit="m" value="59.0">
|
||
<quantity id="4CC774E3F20CFF89958AD5A85508FC17" box="[201,258,942,969]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="394" unit="m" value="59.0">59 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="EE0BBFC1F20CFF89940FD5A855E0FC17" box="[332,490,942,969]" degrees="15" direction="north" minutes="45" orientation="latitude" pageId="2" pageNumber="394" precision="1" seconds="10.6" value="15.752944">15°45’10.6”N</geoCoordinate>
|
||
<geoCoordinate id="EE0BBFC1F20CFF8994B2D5A85696FC17" box="[497,668,942,969]" degrees="100" direction="east" minutes="05" orientation="longitude" pageId="2" pageNumber="394" precision="1" seconds="10.1" value="100.08614">100°05’10.1”E</geoCoordinate>
|
||
,
|
||
<date id="FF81FFC6F20CFF8997EBD5A9571CFC17" box="[680,790,943,969]" pageId="2" pageNumber="394" value="2018-03-04">
|
||
<collectingDate id="EFC5062EF20CFF8997EBD5A9571CFC17" box="[680,790,943,969]" pageId="2" pageNumber="394" value="2018-03-04">4.iii.2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="527EA111F20CFF899661D5A9578EFC17" box="[802,900,943,969]" pageId="2" pageNumber="394">hand net</collectingMethod>
|
||
,
|
||
<collectorName id="26CABCD0F20CFF8996D3D5A95038FC17" box="[912,1074,943,969]" pageId="2" pageNumber="394">C. Techakijvej</collectorName>
|
||
col.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B57D35BF20CFF899129D5A9566FFC33" ID-GBIF-Occurrence="3314062301" collectingDate="2018-03-04" collectionCode="FBRL" collectorName="C. Techakijvej" country="Thailand" county="Ping River" elevation="59" latitude="15.752944" location="Freshwater Biomonitor Research Laboratory" longLatPrecision="1" longitude="100.08614" municipality="Nakonsawan Province" pageId="2" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
|
||
<emphasis id="B94B0514F20CFF899129D5A950E8FC17" bold="true" box="[1130,1250,943,969]" pageId="2" pageNumber="394">
|
||
<typeStatus id="548467A4F20CFF899129D5A950E8FC17" box="[1130,1250,943,969]" pageId="2" pageNumber="394" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED2E41C3F20CFF8991B2D5A95134FC17" box="[1265,1342,943,969]" pageId="2" pageNumber="394">FBRL</collectionCode>
|
||
):
|
||
<specimenCount id="9D39128FF20CFF899012D5A95196FC17" box="[1361,1436,943,969]" pageId="2" pageNumber="394" type="male">1 male</specimenCount>
|
||
mature larva (same data as holotype) and
|
||
</materialsCitation>
|
||
<specimenCount id="9D39128FF20CFF89972ED5D556CCFC33" box="[621,710,979,1005]" pageId="2" pageNumber="394" type="male">10 male</specimenCount>
|
||
imagos, and
|
||
<materialsCitation id="3B57D35BF20CFF89961ED5D554CBFBEB" ID-GBIF-Occurrence="3314062302" collectingDate="2019-11-06" collectorName="C. Techakijvej" country="Thailand" elevation="59" latitude="15.728722" location="Nakonsawan Province" longLatPrecision="1" longitude="100.11684" municipality="Ping River" pageId="2" pageNumber="394" specimenCount="10" specimenCount-female="10" stateProvince="Chiang Mai" typeStatus="paratype">
|
||
<specimenCount id="9D39128FF20CFF89961ED5D557C6FC33" box="[861,972,979,1005]" pageId="2" pageNumber="394" type="female">10 female</specimenCount>
|
||
subimagos with eggs, from
|
||
<location id="8EE08FDDF20CFF89904FD5D554F7FBCF" LSID="urn:lsid:plazi:treatment:03966810F20CFF8D95D4D4245570F9B2:8EE08FDDF20CFF89904FD5D554F7FBCF" country="Thailand" latitude="15.728722" longLatPrecision="1" longitude="100.11684" municipality="Ping River" name="Nakonsawan Province" pageId="2" pageNumber="394" stateProvince="Chiang Mai">Nakonsawan Province</location>
|
||
,
|
||
<collectingMunicipality id="6BE4437CF20CFF899448D5F1558DFBCF" box="[267,391,1015,1041]" pageId="2" pageNumber="394">Ping River</collectingMunicipality>
|
||
,
|
||
<elevation id="00123E35F20CFF8994D6D5F05605FBCF" box="[405,527,1014,1041]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="394" unit="m" value="59.0">
|
||
<quantity id="4CC774E3F20CFF8994D6D5F055DAFBCF" box="[405,464,1014,1041]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="394" unit="m" value="59.0">59 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="EE0BBFC1F20CFF89975ED5F056B1FBCF" box="[541,699,1014,1041]" degrees="15" direction="north" minutes="43" orientation="latitude" pageId="2" pageNumber="394" precision="1" seconds="43.4" value="15.728722">15°43’43.4”N</geoCoordinate>
|
||
<geoCoordinate id="EE0BBFC1F20CFF899786D5F1577AFBCF" box="[709,880,1015,1041]" degrees="100" direction="east" minutes="07" orientation="longitude" pageId="2" pageNumber="394" precision="1" seconds="00.6" value="100.11684">100°07’00.6”E</geoCoordinate>
|
||
,
|
||
<date id="FF81FFC6F20CFF89963DD5F157E0FBCF" box="[894,1002,1015,1041]" pageId="2" pageNumber="394" value="2019-11-06">
|
||
<collectingDate id="EFC5062EF20CFF89963DD5F157E0FBCF" box="[894,1002,1015,1041]" pageId="2" pageNumber="394" value="2019-11-06">6.xi.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="527EA111F20CFF8996BBD5F1506AFBCF" box="[1016,1120,1015,1041]" pageId="2" pageNumber="394">light trap</collectingMethod>
|
||
(
|
||
<figureCitation id="1304C583F20CFF899132D5F150E8FBCF" box="[1137,1250,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1714,1738]" captionTargetBox="[151,1436,741,1691]" captionTargetId="figure-313@7.[151,1436,741,1691]" captionTargetPageId="7" captionText="FIGURE 5. Habitat of Ephoron ookaewae sp. nov.: a, Ping River, Nakonsawan Province, Thailand; b, light trap near a Ping River; c, larva inside a broken U-shaped cavity; d) broken hard clay substrate." figureDoi="http://doi.org/10.5281/zenodo.4956281" httpUri="https://zenodo.org/record/4956281/files/figure.png" pageId="2" pageNumber="394">Fig. 5a–b</figureCitation>
|
||
),
|
||
<collectorName id="26CABCD0F20CFF8991BBD5F15196FBCF" box="[1272,1436,1015,1041]" pageId="2" pageNumber="394">C. Techakijvej</collectorName>
|
||
col.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B57D35BF20CFF899585D21D566EFB87" ID-GBIF-Occurrence="3314062304" collectingDate="2019-11-06" collectorName="C. Techakijvej" country="Thailand" elevation="59" latitude="15.728722" location="Nakonsawan Province" longLatPrecision="1" longitude="100.11684" municipality="Ping River" pageId="2" pageNumber="394" specimenCount="10" specimenCount-adult="4" specimenCount-female="2" specimenCount-larva="2" specimenCount-male="2" stateProvince="Chiang Mai" typeStatus="paratype">
|
||
Additional material, 1 mature larva and eggs for SEM,
|
||
<specimenCount id="9D39128FF20CFF899677D21D5787FBEB" box="[820,909,1051,1077]" pageId="2" pageNumber="394" type="larva">2 larvae</specimenCount>
|
||
and
|
||
<specimenCount id="9D39128FF20CFF899686D21D5017FBEB" box="[965,1053,1051,1077]" pageId="2" pageNumber="394" type="adult">4 adults</specimenCount>
|
||
(
|
||
<specimenCount id="9D39128FF20CFF89916ED21D508EFBEB" box="[1069,1156,1051,1077]" pageId="2" pageNumber="394" type="male">2 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D39128FF20CFF8991FFD21D5120FBE8" box="[1212,1322,1051,1078]" pageId="2" pageNumber="394" type="female">2 females</specimenCount>
|
||
) for DNA extraction (same data as adult
|
||
<typeStatus id="548467A4F20CFF8994A8D2395656FB87" box="[491,604,1087,1113]" pageId="2" pageNumber="394" type="paratype">paratypes</typeStatus>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3258A8DF20CFF8D9584D264515BFE0F" lastPageId="6" lastPageNumber="398" pageId="2" pageNumber="394" type="description">
|
||
<paragraph id="8B80D906F20CFF899584D264566DF963" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="394">
|
||
<emphasis id="B94B0514F20CFF899584D2645553FBA2" bold="true" box="[199,345,1122,1148]" pageId="2" pageNumber="394">Description.</emphasis>
|
||
<emphasis id="B94B0514F20CFF89941CD26555E7FBA2" box="[351,493,1123,1149]" italics="true" pageId="2" pageNumber="394">Male imago.</emphasis>
|
||
(
|
||
<figureCitation id="1304C583F20CFF8994BFD265567BFBA0" box="[508,625,1123,1150]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Figs. 1a–d</figureCitation>
|
||
) Length (mm) (n=10): body (excluding caudal filament and cerci) 10.4– 11.1; forewings 7.4–8.7; hindwings 4.1–4.4; caudal filament 0.4–0.6; cerci 21.4–24.4. Head. Head as wide as pronotum, whitish to very pale yellowish, antennae whitish (
|
||
<figureCitation id="1304C583F20CFF899665D2AD577DFB1B" box="[806,887,1195,1221]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1a</figureCitation>
|
||
). Ecdysal line outlined with black, from margin of lateral ocelli to mid-line.
|
||
<emphasis id="B94B0514F20CFF899499D2C95639FB37" box="[474,563,1231,1257]" italics="true" pageId="2" pageNumber="394">Thorax.</emphasis>
|
||
Pronotum whitish with brown pattern (
|
||
<figureCitation id="1304C583F20CFF8996BBD2C95043FB37" box="[1016,1097,1231,1257]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1a</figureCitation>
|
||
), meso- and metanotum, pale yellowish. Legs (
|
||
<figureCitation id="1304C583F20CFF899414D2F555A2FAD0" box="[343,424,1267,1294]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1d</figureCitation>
|
||
). Forelegs (
|
||
<quantity id="4CC774E3F20CFF89976FD2F4569FFAD3" box="[556,661,1266,1293]" metricMagnitude="-3" metricUnit="m" metricValue="5.92" pageId="2" pageNumber="394" unit="mm" value="5.92">5.92 mm</quantity>
|
||
), forefemur with dark brown line on dorsal and ventral surfaces, foretibiae with brown line along its length, length ratio of forefemur to foretibiae = 1: 2.9, length ratio of five foretarsal segments = 0.2: 2.2: 1.4: 1.3: 1, tarsi with whitish forked claws. Mid- and hindlegs reduced, whitish. Mid-leg (
|
||
<quantity id="4CC774E3F20CFF899029D33D54CDFAA7" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="2" pageNumber="394" unit="mm" value="1.78">1.78 mm</quantity>
|
||
), length ratio of midfemur and midtibia = 1: 0.8, length ratio of three midtarsal segments = 1: 0.9: 1. Hindleg (
|
||
<quantity id="4CC774E3F20CFF8995DCD3855500FA43" box="[159,266,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="2" pageNumber="394" unit="mm" value="2.36">2.36 mm</quantity>
|
||
), length ratio of hindfemur to hindtibia = 1: 0.7, length ratio of three hindtarsal segments = 1: 0.7: 0.8. Wings (
|
||
<figureCitation id="1304C583F20CFF8995B3D3A15549FA1C" box="[240,323,1447,1474]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1b</figureCitation>
|
||
), forewing hyaline, numerous cross-veins. Main vein of costal margin brownish, particularly near the wing base. Membrane on costal area especially pterostigma whitish, AA to wing base whitish and posterior margin pale whitish, fork of MA distal to fork of RS,
|
||
<specimenCount id="9D39128FF20CFF89978DD3E95767F9D7" box="[718,877,1519,1545]" pageId="2" pageNumber="394" type="intercalary">4 intercalaries</specimenCount>
|
||
between CuA and CuP. Hindwing hyaline, costal projection well developed,
|
||
<specimenCount id="9D39128FF20CFF89948DD0155657F9F3" box="[462,605,1555,1581]" pageId="2" pageNumber="394" type="intercalary">1 intercalary</specimenCount>
|
||
between RA and RS, MA forked near middle of hindwing,
|
||
<specimenCount id="9D39128FF20CFF89904DD0155197F9F3" box="[1294,1437,1555,1581]" pageId="2" pageNumber="394" type="intercalary">1 intercalary</specimenCount>
|
||
between CuA and CuP. Abdomen. Abdominal terga whitish, with dark brownish markings becoming darker and wider posteriorly, separated laterally and meeting at the posterior end. Genitalia: forceps 4-segmented, penes whitish (width:
|
||
<quantity id="4CC774E3F20CFF899459D07855B7F947" box="[282,445,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.25" metricValueMin="1.23" pageId="2" pageNumber="394" unit="mm" value="1.24" valueMax="1.25" valueMin="1.23">1.23-1.25 mm</quantity>
|
||
), penis lobes straight and diverging laterally, apex of each lobe hooked (
|
||
<figureCitation id="1304C583F20CFF8991B4D0795142F947" box="[1271,1352,1663,1689]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1c</figureCitation>
|
||
). Cerci well-developed, caudal filament reduced.
|
||
</paragraph>
|
||
<paragraph id="8B80D906F20CFF889584D0C15033FF27" blockId="2.[151,1437,654,2013]" lastBlockId="3.[151,1436,151,249]" lastPageId="3" lastPageNumber="395" pageId="2" pageNumber="394">
|
||
<emphasis id="B94B0514F20CFF899584D0C155A9F93F" box="[199,419,1735,1761]" italics="true" pageId="2" pageNumber="394">Female sub-imago.</emphasis>
|
||
Length (mm) (n=10): body (excluding caudal filament and cerci) 13.7–13.8; forewings 10.9–11.4; hindwings, 4.5–4.9; caudal filament, 5.7–6.1; cerci, 7.7–8.5. Head. Markings and color similar to male. Thorax. Color and pattern same as male. All legs reduced (
|
||
<figureCitation id="1304C583F20CFF899663D1095765F8F7" box="[800,879,1807,1833]" captionStart="FIGURE 1" captionStartId="3.[151,250,1616,1640]" captionTargetBox="[151,1426,273,1593]" captionTargetId="figure-74@3.[151,1436,273,1593]" captionTargetPageId="3" captionText="FIGURE 1. Ephoron ookaewae sp. nov., adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="2" pageNumber="394">Fig. 1e</figureCitation>
|
||
). Foreleg (
|
||
<quantity id="4CC774E3F20CFF8996ABD108505BF8F7" box="[1000,1105,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" pageId="2" pageNumber="394" unit="mm" value="2.53">2.53 mm</quantity>
|
||
), forefemora with dark brown line on dorsal and ventral surfaces, foretibia dark brown, length ratio of forefemur to foretibia = 1: 1.1, foretarsus whitish, length ratio of five foretarsal segments = 1: 1.6: 1.4: 0.9: 1.2. Mid- and hindlegs whitish. Midleg (
|
||
<quantity id="4CC774E3F20CFF899029D15154CDF84B" metricMagnitude="-3" metricUnit="m" metricValue="2.41" pageId="2" pageNumber="394" unit="mm" value="2.41">2.41 mm</quantity>
|
||
), length ratio of midfemur and midtibia = 1: 1.3, length ratio of three midtarsal segments = 1: 0.6: 0.7. Hindleg (
|
||
<quantity id="4CC774E3F20CFF8995DCD1995500F867" box="[159,266,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="394" unit="mm" value="3.2">3.20 mm</quantity>
|
||
), length ratio of three hindtarsal segments = 1: 1.2: 1. Forewing whitish hyaline, numerous cross veins. Main vein of costal margin brownish especially near the wing base. Membrane of wing in costal area especially pterostigma whitish. Four intercalaries between CuA and CuP. Hindwing, costal projection well developed, no intercalary vein between RA and RS. Abdomen. Abdominal terga whitish, with dark brownish markings, becoming darker and wider at posterior end. Caudal filament and cerci developed, with setae.
|
||
</paragraph>
|
||
<caption id="DF40898EF20DFF8895D4D056565FF96E" ID-DOI="http://doi.org/10.5281/zenodo.4956271" ID-Zenodo-Dep="4956271" httpUri="https://zenodo.org/record/4956271/files/figure.png" pageId="3" pageNumber="395" startId="3.[151,250,1616,1640]" targetBox="[151,1426,273,1593]" targetPageId="3">
|
||
<paragraph id="8B80D906F20DFF8895D4D056565FF96E" blockId="3.[151,1436,1616,1713]" pageId="3" pageNumber="395">
|
||
<emphasis id="B94B0514F20DFF8895D4D0565519F9B6" bold="true" box="[151,275,1616,1640]" pageId="3" pageNumber="395">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C3FA285F20DFF88945AD05655D6F9B6" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[281,476,1616,1640]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
|
||
<emphasis id="B94B0514F20DFF88945AD05655D6F9B6" box="[281,476,1616,1640]" italics="true" pageId="3" pageNumber="395">Ephoron ookaewae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94B0514F20DFF8894A0D0575639F9B7" bold="true" box="[483,563,1617,1641]" pageId="3" pageNumber="395">
|
||
<taxonomicNameLabel id="A278B86FF20DFF8894A0D0575639F9B7" box="[483,563,1617,1641]" pageId="3" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult: a, male imago, dorsal view; b, wings of male imago, (1) RS fork, (2) MA fork; c, penes, ventral view; d, male legs; e, female subimago legs. Cx = coxa, Tr = trochanter, Fe = femur, Ti = tibia, T = tarsal segments 1–5 and C = claw; Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B80D906F20DFF8D9584D0D95064FEC3" blockId="3.[151,1437,1759,2037]" lastBlockId="6.[151,1437,151,465]" lastPageId="6" lastPageNumber="398" pageId="3" pageNumber="395">
|
||
<emphasis id="B94B0514F20DFF889584D0D95563F927" box="[199,361,1759,1785]" italics="true" pageId="3" pageNumber="395">Mature larvae</emphasis>
|
||
(
|
||
<figureCitation id="1304C583F20DFF889439D0D955D7F927" box="[378,477,1759,1785]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,250,1941,1965]" captionStartId-1="5.[151,250,1954,1978]" captionTargetBox-0="[158,1429,181,1918]" captionTargetBox-1="[224,1362,181,1930]" captionTargetId-0="figure-18@4.[158,1429,181,1918]" captionTargetId-1="figure-30@5.[224,1362,181,1930]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." captionText-1="FIGURE 3. Ephoron ookaewae sp. nov., mature larva (SEM): a, apex of right mandible; b, same, detail of small triangular spine (arrow)." figureDoi-0="http://doi.org/10.5281/zenodo.4956273" figureDoi-1="http://doi.org/10.5281/zenodo.4956275" httpUri-0="https://zenodo.org/record/4956273/files/figure.png" httpUri-1="https://zenodo.org/record/4956275/files/figure.png" pageId="3" pageNumber="395">Fig. 2–3</figureCitation>
|
||
). Length (mm) (n=2): body (excluding mandibular tusks, caudal filament and cerci) 10.0–10.3; mandibular tusks 0.95–0.98; caudal filament 3.9–4.0; cerci 6.7–6.7. Head. Head as wide as pronotum, whitish. Ecdysal line outlined as in male imago. Head with a three to four-pointed median frontal process (
|
||
<figureCitation id="1304C583F20DFF88902CD12154BFF8BB" captionStart="FIGURE 2" captionStartId="4.[151,250,1941,1965]" captionTargetBox="[158,1429,181,1918]" captionTargetId="figure-18@4.[158,1429,181,1918]" captionTargetPageId="4" captionText="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956273" httpUri="https://zenodo.org/record/4956273/files/figure.png" pageId="3" pageNumber="395">Fig. 2c</figureCitation>
|
||
). Mandibular tusks shorter than maximal width of pronotum, straight with apex curved upwards, in lateral view (
|
||
<figureCitation id="1304C583F20DFF8895E3D16954F9F854" box="[160,243,1903,1930]" captionStart="FIGURE 2" captionStartId="4.[151,250,1941,1965]" captionTargetBox="[158,1429,181,1918]" captionTargetId="figure-18@4.[158,1429,181,1918]" captionTargetPageId="4" captionText="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956273" httpUri="https://zenodo.org/record/4956273/files/figure.png" pageId="3" pageNumber="395">Fig. 2b</figureCitation>
|
||
). Tusks with 5–7 tubercles, tubercles smaller at base, with dense group of short setae on upper surface near bases of tusk (
|
||
<figureCitation id="1304C583F20DFF889476D195558FF873" box="[309,389,1939,1965]" captionStart="FIGURE 2" captionStartId="4.[151,250,1941,1965]" captionTargetBox="[158,1429,181,1918]" captionTargetId="figure-18@4.[158,1429,181,1918]" captionTargetPageId="4" captionText="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956273" httpUri="https://zenodo.org/record/4956273/files/figure.png" pageId="3" pageNumber="395">Fig. 2c</figureCitation>
|
||
) and long setae on lateral surface. Right mandible (
|
||
<figureCitation id="1304C583F20DFF889682D195501AF873" box="[961,1040,1939,1965]" captionStart="FIGURE 3" captionStartId="5.[151,250,1954,1978]" captionTargetBox="[224,1362,181,1930]" captionTargetId="figure-30@5.[224,1362,181,1930]" captionTargetPageId="5" captionText="FIGURE 3. Ephoron ookaewae sp. nov., mature larva (SEM): a, apex of right mandible; b, same, detail of small triangular spine (arrow)." figureDoi="http://doi.org/10.5281/zenodo.4956275" httpUri="https://zenodo.org/record/4956275/files/figure.png" pageId="3" pageNumber="395">Fig. 3a</figureCitation>
|
||
) with small triangular shaped spine between outer and inner incisors (
|
||
<figureCitation id="1304C583F20DFF889759D1B15665F80C" box="[538,623,1975,2002]" captionStart="FIGURE 3" captionStartId="5.[151,250,1954,1978]" captionTargetBox="[224,1362,181,1930]" captionTargetId="figure-30@5.[224,1362,181,1930]" captionTargetPageId="5" captionText="FIGURE 3. Ephoron ookaewae sp. nov., mature larva (SEM): a, apex of right mandible; b, same, detail of small triangular spine (arrow)." figureDoi="http://doi.org/10.5281/zenodo.4956275" httpUri="https://zenodo.org/record/4956275/files/figure.png" pageId="3" pageNumber="395">Fig. 3b</figureCitation>
|
||
). Maxillary and labial palp, two segmented. Thorax. Pronotum whitish, brown pattern as in male imago. Meso and metanotum pale yellowish, wing pad dark brown (
|
||
<figureCitation id="1304C583F20DFF889192D1DD512FF82B" box="[1233,1317,2011,2037]" captionStart="FIGURE 2" captionStartId="4.[151,250,1941,1965]" captionTargetBox="[158,1429,181,1918]" captionTargetId="figure-18@4.[158,1429,181,1918]" captionTargetPageId="4" captionText="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956273" httpUri="https://zenodo.org/record/4956273/files/figure.png" pageId="3" pageNumber="395">Fig. 2a</figureCitation>
|
||
). Forelegs stout and fossorial with rows of long setae. Inner side of forefemur has a U-shaped row of setae. Foretibiae flattened, with two rows of ventral tubercles along margins. Middle legs small. Abdomen. Same color and pattern as male. Gills whitish, gills present on abdominal segments 1–7, black dot on gills base, gill 1 single, gills 2–7 forked and each lamella with fringed lateral margin (
|
||
<figureCitation id="1304C583F208FF8D9724D70556B0FEC0" box="[615,698,259,286]" captionStart="FIGURE 2" captionStartId="4.[151,250,1941,1965]" captionTargetBox="[158,1429,181,1918]" captionTargetId="figure-18@4.[158,1429,181,1918]" captionTargetPageId="4" captionText="FIGURE 2. Ephoron ookaewae sp. nov., mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4956273" httpUri="https://zenodo.org/record/4956273/files/figure.png" pageId="6" pageNumber="398">Fig. 2d</figureCitation>
|
||
). Caudal filament and cerci with setae.
|
||
</paragraph>
|
||
<caption id="DF40898EF20AFF8F95D4D19350A5F80F" ID-DOI="http://doi.org/10.5281/zenodo.4956273" ID-Zenodo-Dep="4956273" httpUri="https://zenodo.org/record/4956273/files/figure.png" pageId="4" pageNumber="396" startId="4.[151,250,1941,1965]" targetBox="[158,1429,181,1918]" targetPageId="4">
|
||
<paragraph id="8B80D906F20AFF8F95D4D19350A5F80F" blockId="4.[151,1437,1941,2002]" pageId="4" pageNumber="396">
|
||
<emphasis id="B94B0514F20AFF8F95D4D1935518F873" bold="true" box="[151,274,1941,1965]" pageId="4" pageNumber="396">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3FA285F20AFF8F945BD19355D0F873" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[280,474,1941,1965]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
|
||
<emphasis id="B94B0514F20AFF8F945BD19355D0F873" box="[280,474,1941,1965]" italics="true" pageId="4" pageNumber="396">Ephoron ookaewae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94B0514F20AFF8F94A3D190563AF870" bold="true" box="[480,560,1942,1966]" pageId="4" pageNumber="396">
|
||
<taxonomicNameLabel id="A278B86FF20AFF8F94A3D190563AF870" box="[480,560,1942,1966]" pageId="4" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, mature larva: a, male larva, dorsal view; b, mandibular tusks, lateral view; c, head and pronotum, dorsal view (arrow indicates median frontal process); d, abdominal gill 2. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF40898EF20BFF8E95D4D1A4552CF800" ID-DOI="http://doi.org/10.5281/zenodo.4956275" ID-Zenodo-Dep="4956275" httpUri="https://zenodo.org/record/4956275/files/figure.png" pageId="5" pageNumber="397" startId="5.[151,250,1954,1978]" targetBox="[224,1362,181,1930]" targetPageId="5">
|
||
<paragraph id="8B80D906F20BFF8E95D4D1A4552CF800" blockId="5.[151,1436,1954,2014]" pageId="5" pageNumber="397">
|
||
<emphasis id="B94B0514F20BFF8E95D4D1A4551FF864" bold="true" box="[151,277,1954,1978]" pageId="5" pageNumber="397">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3FA285F20BFF8E945ED1A455E8F864" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[285,482,1954,1978]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
|
||
<emphasis id="B94B0514F20BFF8E945ED1A455E8F864" box="[285,482,1954,1978]" italics="true" pageId="5" pageNumber="397">Ephoron ookaewae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94B0514F20BFF8E94A8D1A45637F864" bold="true" box="[491,573,1954,1978]" pageId="5" pageNumber="397">
|
||
<taxonomicNameLabel id="A278B86FF20BFF8E94A8D1A45637F864" box="[491,573,1954,1978]" pageId="5" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, mature larva (SEM): a, apex of right mandible; b, same, detail of small triangular spine (arrow).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B80D906F208FF8D9584D721515BFE0F" blockId="6.[151,1437,151,465]" pageId="6" pageNumber="398">
|
||
<emphasis id="B94B0514F208FF8D9584D72154F6FE9E" box="[199,252,295,320]" italics="true" pageId="6" pageNumber="398">Egg.</emphasis>
|
||
(
|
||
<figureCitation id="1304C583F208FF8D9449D7215574FE9F" box="[266,382,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1152,1176]" captionTargetBox="[151,1436,489,1128]" captionTargetId="figure-183@6.[151,1436,489,1128]" captionTargetPageId="6" captionText="FIGURE 4. Ephoron ookaewae sp. nov., eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide)." figureDoi="http://doi.org/10.5281/zenodo.4956277" httpUri="https://zenodo.org/record/4956277/files/figure.png" pageId="6" pageNumber="398">Figs. 4a–e</figureCitation>
|
||
) Length (µm) (n=10) cap width 230–266, width 161–209. Eggs acorn shaped (
|
||
<figureCitation id="1304C583F208FF8D91A5D7215156FE9C" box="[1254,1372,295,322]" captionStart="FIGURE 4" captionStartId="6.[151,250,1152,1176]" captionTargetBox="[151,1436,489,1128]" captionTargetId="figure-183@6.[151,1436,489,1128]" captionTargetPageId="6" captionText="FIGURE 4. Ephoron ookaewae sp. nov., eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide)." figureDoi="http://doi.org/10.5281/zenodo.4956277" httpUri="https://zenodo.org/record/4956277/files/figure.png" pageId="6" pageNumber="398">Figs. 4a–b</figureCitation>
|
||
) with a concave indentation at anterior area (
|
||
<figureCitation id="1304C583F208FF8D9716D74D56ACFEB8" box="[597,678,331,358]" captionStart="FIGURE 4" captionStartId="6.[151,250,1152,1176]" captionTargetBox="[151,1436,489,1128]" captionTargetId="figure-183@6.[151,1436,489,1128]" captionTargetPageId="6" captionText="FIGURE 4. Ephoron ookaewae sp. nov., eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide)." figureDoi="http://doi.org/10.5281/zenodo.4956277" httpUri="https://zenodo.org/record/4956277/files/figure.png" pageId="6" pageNumber="398">Figs. 4</figureCitation>
|
||
b-c). Eggs with one polar cap (multi-unit cap;
|
||
<bibRefCitation id="EFAEA4F7F208FF8D91F8D74D54D8FE57" author="Gaino, E. & Flannagan, J." pageId="6" pageNumber="398" pagination="527 - 533" refId="ref5088" refString="Gaino, E. & Flannagan, J. (1995) Fine external morphology of the eggs of Ephoron album (Say) and Ephoron shigae (Takahashi) (Ephemeroptera, Polymitarcyidae). The Canadian Entomologist, 127 (4), 527 - 533. https: // doi. org / 10.4039 / Ent 127527 - 4" type="journal article" year="1995">Gaino & Flannagan 1995</bibRefCitation>
|
||
), length ratio of polar cap to egg length = 0.39: 1. Micropyles are located around the base of the polar cap (
|
||
<figureCitation id="1304C583F208FF8D902CD76954BFFE73" captionStart="FIGURE 4" captionStartId="6.[151,250,1152,1176]" captionTargetBox="[151,1436,489,1128]" captionTargetId="figure-183@6.[151,1436,489,1128]" captionTargetPageId="6" captionText="FIGURE 4. Ephoron ookaewae sp. nov., eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide)." figureDoi="http://doi.org/10.5281/zenodo.4956277" httpUri="https://zenodo.org/record/4956277/files/figure.png" pageId="6" pageNumber="398">Fig. 4e</figureCitation>
|
||
). Chorion sculptured in hexagonal netted pattern, convex area inside each cell, about
|
||
<quantity id="4CC774E3F208FF8D9120D79550A1FE73" box="[1123,1195,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="398" unit="mm" value="12.0">12 µm</quantity>
|
||
in diameter (
|
||
<figureCitation id="1304C583F208FF8D907ED795519AFE70" box="[1341,1424,403,430]" captionStart="FIGURE 4" captionStartId="6.[151,250,1152,1176]" captionTargetBox="[151,1436,489,1128]" captionTargetId="figure-183@6.[151,1436,489,1128]" captionTargetPageId="6" captionText="FIGURE 4. Ephoron ookaewae sp. nov., eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide)." figureDoi="http://doi.org/10.5281/zenodo.4956277" httpUri="https://zenodo.org/record/4956277/files/figure.png" pageId="6" pageNumber="398">Fig. 4d</figureCitation>
|
||
). Eggs form two elongate oval pale-yellowish eggs masses. Length of eggs masses are
|
||
<quantity id="4CC774E3F208FF8D9117D7B050FDFE0F" box="[1108,1271,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.495" metricValueMax="5.53" metricValueMin="5.46" pageId="6" pageNumber="398" unit="mm" value="5.495" valueMax="5.53" valueMin="5.46">5.46–5.53 mm</quantity>
|
||
(n=10).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF40898EF208FF8D95D4D28650A8FB62" ID-DOI="http://doi.org/10.5281/zenodo.4956277" ID-Zenodo-Dep="4956277" httpUri="https://zenodo.org/record/4956277/files/figure.png" pageId="6" pageNumber="398" startId="6.[151,250,1152,1176]" targetBox="[151,1436,489,1128]" targetPageId="6">
|
||
<paragraph id="8B80D906F208FF8D95D4D28650A8FB62" blockId="6.[151,1436,1152,1212]" pageId="6" pageNumber="398">
|
||
<emphasis id="B94B0514F208FF8D95D4D286551EFB46" bold="true" box="[151,276,1152,1176]" pageId="6" pageNumber="398">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C3FA285F208FF8D9458D28655D5FB46" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[283,479,1152,1176]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
|
||
<emphasis id="B94B0514F208FF8D9458D28655D5FB46" box="[283,479,1152,1176]" italics="true" pageId="6" pageNumber="398">Ephoron ookaewae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94B0514F208FF8D94A5D2865632FB46" bold="true" box="[486,568,1152,1176]" pageId="6" pageNumber="398">
|
||
<taxonomicNameLabel id="A278B86FF208FF8D94A5D2865632FB46" box="[486,568,1152,1176]" pageId="6" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, eggs: a, egg (SEM); b, egg (light microscope); c, concave indented anterior area; d, detail of uncapped pole; e, micropylar devices (mc: micropylar canal, m: micropyle, sg: sperm guide).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3258A8DF208FF8D9584D2EC5570F9B2" pageId="6" pageNumber="398" type="etymology">
|
||
<paragraph id="8B80D906F208FF8D9584D2EC5601FA92" blockId="6.[151,1437,1258,1644]" pageId="6" pageNumber="398">
|
||
<emphasis id="B94B0514F208FF8D9584D2EC5544FADA" bold="true" box="[199,334,1258,1284]" pageId="6" pageNumber="398">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="4C3FA285F208FF8D9763D2EC569AFADA" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[544,656,1258,1284]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="ookaewae">
|
||
<emphasis id="B94B0514F208FF8D9763D2EC569AFADA" box="[544,656,1258,1284]" italics="true" pageId="6" pageNumber="398">ookaewae</emphasis>
|
||
</taxonomicName>
|
||
is dedicated to Assoc. Prof. Dr. Ookaew Prakobvitayakit (
|
||
<collectingRegion id="49FB17E4F208FF8D9055D2EC5196FADA" box="[1302,1436,1258,1284]" country="Thailand" name="Chiang Mai" pageId="6" pageNumber="398">Chiang Mai</collectingRegion>
|
||
University). Dr. Prakobvitayakit are well-known researcher on the aquatic ecology of
|
||
<collectingCountry id="F3289996F208FF8D9117D30850B0FAF6" box="[1108,1210,1294,1320]" name="Thailand" pageId="6" pageNumber="398">Thailand</collectingCountry>
|
||
, over 40 years, until retirement. The gender is female.
|
||
</paragraph>
|
||
<paragraph id="8B80D906F208FF8D9584D3505533FA4B" blockId="6.[151,1437,1258,1644]" pageId="6" pageNumber="398">
|
||
<emphasis id="B94B0514F208FF8D9584D3505599FAAE" bold="true" box="[199,403,1366,1392]" pageId="6" pageNumber="398">DNA barcoding.</emphasis>
|
||
Sequences of
|
||
<taxonomicName id="4C3FA285F208FF8D9710D350573FFAAE" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[595,821,1366,1392]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
|
||
<emphasis id="B94B0514F208FF8D9710D350573FFAAE" box="[595,821,1366,1392]" italics="true" pageId="6" pageNumber="398">Ephoron ookaewae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94B0514F208FF8D9605D35057A3FAAE" bold="true" box="[838,937,1366,1392]" pageId="6" pageNumber="398">
|
||
<taxonomicNameLabel id="A278B86FF208FF8D9605D35057A3FAAE" box="[838,937,1366,1392]" pageId="6" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were deposited in a GenBank database (
|
||
<accessionNumber id="946C44E5F208FF8D95DCD37C5527FA4B" box="[159,301,1402,1429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW168821" pageId="6" pageNumber="398">MW168821</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B80D906F208FF8D9584D3985570F9B2" blockId="6.[151,1437,1258,1644]" pageId="6" pageNumber="398">
|
||
<emphasis id="B94B0514F208FF8D9584D39855B5FA66" bold="true" box="[199,447,1438,1464]" pageId="6" pageNumber="398">Habitat and ecology.</emphasis>
|
||
<taxonomicName id="4C3FA285F208FF8D9484D39856ABFA66" authority="Techakijvej & Sareein & Hwang & Bae & Phalaraksh, 2021" authorityName="Techakijvej & Sareein & Hwang & Bae & Phalaraksh" authorityYear="2021" box="[455,673,1438,1464]" class="Insecta" family="Polymitarcyidae" genus="Ephoron" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="398" phylum="Arthropoda" rank="species" species="ookaewae" status="sp. nov.">
|
||
<emphasis id="B94B0514F208FF8D9484D39856ABFA66" box="[455,673,1438,1464]" italics="true" pageId="6" pageNumber="398">Ephoron ookaewae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94B0514F208FF8D97EAD398570EFA66" bold="true" box="[681,772,1438,1464]" pageId="6" pageNumber="398">
|
||
<taxonomicNameLabel id="A278B86FF208FF8D97EAD398570EFA66" box="[681,772,1438,1464]" pageId="6" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
larvae were found in U-shaped burrows in hard clay (
|
||
<figureCitation id="1304C583F208FF8D902CD39854C5FA03" captionStart="FIGURE 5" captionStartId="7.[151,250,1714,1738]" captionTargetBox="[151,1436,741,1691]" captionTargetId="figure-313@7.[151,1436,741,1691]" captionTargetPageId="7" captionText="FIGURE 5. Habitat of Ephoron ookaewae sp. nov.: a, Ping River, Nakonsawan Province, Thailand; b, light trap near a Ping River; c, larva inside a broken U-shaped cavity; d) broken hard clay substrate." figureDoi="http://doi.org/10.5281/zenodo.4956281" httpUri="https://zenodo.org/record/4956281/files/figure.png" pageId="6" pageNumber="398">Fig. 5c–d</figureCitation>
|
||
) in moderately deep water (
|
||
<quantity id="4CC774E3F208FF8D9740D3C4567FFA03" box="[515,629,1474,1501]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="6" pageNumber="398" unit="cm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 cm</quantity>
|
||
) and moderate flow (
|
||
<quantity id="4CC774E3F208FF8D9623D3C457D8FA02" box="[864,978,1474,1500]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="398" unit="m" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 m</quantity>
|
||
/sec) of a large river. Some physicochemical variables of water at the moment of collection: temperature 31°C, pH 8.56, Conductivity 265 µS/cm, dissolved oxygen
|
||
<quantity id="4CC774E3F208FF8D95B3D00C5537F9FA" box="[240,317,1546,1572]" metricMagnitude="-6" metricUnit="kg" metricValue="9.2" pageId="6" pageNumber="398" unit="mg" value="9.2">9.2 mg</quantity>
|
||
/L and turbidity 11 NTU. Adult swarm densities increase between 7:00 p.m. and 8:00 p.m. (GMT +7 hrs). Emergence of male imagos occurred about one hour after sunset and female subimagos emerged about 15–20 minutes after males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |