167 lines
18 KiB
XML
167 lines
18 KiB
XML
<document id="09B2202620AE90CA8E70FA27B9207C93" ID="10.11646/zootaxa.4196.1.9" ID-DOI="10.11646/zootaxa.4196.1.9" ID-GBIF-Dataset="d535b664-dea2-43a9-a058-a69c175912f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="167723" ID-ZooBank="71F26B38-19B0-4402-BDF3-FDF6378E3223" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479714024570" checkinUser="plazi" docAuthor="Nel, Andre, Perreau, Ziann & Doitteau, Gaelle" docDate="2016" docId="03FED014FF80FF89FF60FE69FC3737C8" docLanguage="en" docName="zootaxa.4196.1.9.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Symphoromyia eocenica Nel, Perreau & Doitteau, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="145" masterDocId="FFC7A86CFF81FF88FFF7FF9BFFBE312B" masterDocTitle="The oldest representative of the modern snipe fly genus Symphoromyia (Diptera: Rhagionidae)" masterLastPageNumber="150" masterPageNumber="144" pageNumber="145" updateTime="1698394045248" updateUser="plazi">
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<mods:titleInfo id="662F32167F17D13E56B81EE07FE32D78">
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<mods:title id="49E1755E2BA3B8ABB7A762A9FFFDCC41">The oldest representative of the modern snipe fly genus Symphoromyia (Diptera: Rhagionidae)</mods:title>
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<mods:name id="65EFF8CED64BC580B0D4F5A0E27549C2" type="personal">
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<mods:roleTerm id="5E226F96FB5410BA6CDEA17A2B18167A">Author</mods:roleTerm>
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<mods:namePart id="20E9292A63F62A805EF5E09EA7C7CACB">Nel, Andre</mods:namePart>
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<mods:roleTerm id="8B65836C17584D66FD00D5F8DDA87FC2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6DBCD87884CC15EF51A7ABDA5731525B">Perreau, Ziann</mods:namePart>
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</mods:name>
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<mods:name id="52449797ED5EBCC975226AC336FA3EF8" type="personal">
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<mods:roleTerm id="B10171840A192C81E029C47D33928CC4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A2587A65F0B2312BFCB7956BA6D38FAF">Doitteau, Gaelle</mods:namePart>
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<mods:typeOfResource id="B1A0556391D721C0E6B6C4C89DDFD350">text</mods:typeOfResource>
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<mods:title id="6153DBE94789C5F1B1AFD4D9FA55D74C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A7E8FB8383D00CD27B01C6E6ABEFD003">
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<mods:date id="8CD03C2CE2AF0E9E592F13E377FFE57B">2016</mods:date>
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<mods:detail id="5045D4D3E4175AD74B7B471DF94ED2AD" type="volume">
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<mods:number id="ACCF3C426F0E504B8B3FB9B447952E5B">4196</mods:number>
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</mods:detail>
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<mods:detail id="327F90C91E49E7A8E11ECFDB268CA9E4" type="issue">
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<mods:number id="1C29F9A331EF6E7D3B38D1C953BC165C">1</mods:number>
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<mods:start id="1A770A69A863A650A6DC8E24BAD4EC90">144</mods:start>
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<mods:end id="F319C744BA134C519196984C580C3BB3">150</mods:end>
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<mods:classification id="F8F5D405EBA2603A59CDA607B538621C">journal article</mods:classification>
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<mods:identifier id="4ABB5519CFAC5B07182C1FC3DD955B44" type="DOI">10.11646/zootaxa.4196.1.9</mods:identifier>
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<mods:identifier id="8C7C4382329F28E733542A81B7E1F707" type="GBIF-Dataset">d535b664-dea2-43a9-a058-a69c175912f2</mods:identifier>
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<mods:identifier id="4DF3F6F3579CCC44563DE6974C24BB36" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5192504525D8E23EAB77C9B1D8887D70" type="Zenodo-Dep">167723</mods:identifier>
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<mods:identifier id="88D4982A307DAADEBF533523F5F2F9AE" type="ZooBank">71F26B38-19B0-4402-BDF3-FDF6378E3223</mods:identifier>
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<treatment id="03FED014FF80FF89FF60FE69FC3737C8" ID-DOI="http://doi.org/10.5281/zenodo.6057816" ID-GBIF-Taxon="124814376" ID-Zenodo-Dep="6057816" LSID="urn:lsid:plazi:treatment:03FED014FF80FF89FF60FE69FC3737C8" httpUri="http://treatment.plazi.org/id/03FED014FF80FF89FF60FE69FC3737C8" lastPageNumber="145" pageId="1" pageNumber="145">
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<subSubSection id="C34D3289FF80FF89FF60FE69FEAF3305" pageId="1" pageNumber="145" type="nomenclature">
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<paragraph id="8BE86102FF80FF89FF60FE69FD9D3327" blockId="1.[151,547,498,558]" box="[151,547,498,524]" pageId="1" pageNumber="145">
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<heading id="D0A0D66EFF80FF89FF60FE69FD9D3327" bold="true" box="[151,547,498,524]" fontSize="11" level="1" pageId="1" pageNumber="145" reason="1">
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<emphasis id="B923BD10FF80FF89FF60FE69FD9D3327" bold="true" box="[151,547,498,524]" pageId="1" pageNumber="145">
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<taxonomicName id="4C571A81FF80FF89FF60FE69FE003327" box="[151,446,498,524]" class="Insecta" family="Rhagionidae" genus="Symphoromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="145" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="B923BD10FF80FF89FF60FE69FE003327" bold="true" box="[151,446,498,524]" italics="true" pageId="1" pageNumber="145">Symphoromyia eocenica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A210006BFF80FF89FE32FE69FD9D3327" box="[453,547,498,524]" pageId="1" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BE86102FF80FF89FF60FD8EFEAF3305" blockId="1.[151,547,498,558]" box="[151,273,533,558]" pageId="1" pageNumber="145">
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(
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<figureCitation id="136C7D87FF80FF89FF68FD8EFEB73305" box="[159,265,533,558]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1951,1973]" captionStartId-1="3.[151,250,1960,1982]" captionStartId-2="4.[151,250,1104,1126]" captionTargetBox-0="[151,1436,195,1926]" captionTargetBox-1="[178,1413,194,1916]" captionTargetBox-2="[193,1397,193,1079]" captionTargetId-0="figure@2.[151,1436,193,1929]" captionTargetId-1="figure@3.[178,1414,193,1916]" captionTargetId-2="figure@4.[193,1398,193,1079]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Symphoromyia eocenica sp. nov., holotype MNHN. F. A 57537. Habitus. A, left side; B, right side." captionText-1="FIGURE 2. Symphoromyia eocenica sp. nov., holotype MNHN. F. A 57537. A, head, white arrow first flagellomere; black arrows palpomeres; B, thorax, abdomen, and wing." captionText-2="FIGURE 3. Symphoromyia eocenica sp. nov., holotype MNHN. F. A 57537. Wing." httpUri-0="https://zenodo.org/record/167724/files/figure.png" httpUri-1="https://zenodo.org/record/167725/files/figure.png" httpUri-2="https://zenodo.org/record/167726/files/figure.png" pageId="1" pageNumber="145">Figs. 1–3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34D3289FF80FF89FF60FDC6FD36335D" box="[151,648,605,630]" pageId="1" pageNumber="145" type="etymology">
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<paragraph id="8BE86102FF80FF89FF60FDC6FD36335D" blockId="1.[151,1437,605,1763]" box="[151,648,605,630]" pageId="1" pageNumber="145">
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<emphasis id="B923BD10FF80FF89FF60FDC6FEA8335D" bold="true" box="[151,278,605,630]" pageId="1" pageNumber="145">Etymology</emphasis>
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. Named after the Eocene period.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D3289FF80FF89FF30FD1AFE383265" pageId="1" pageNumber="145" type="materials_examined">
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<paragraph id="8BE86102FF80FF89FF30FD1AFE843395" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
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<emphasis id="B923BD10FF80FF89FF30FD1AFE9133B1" bold="true" box="[199,303,641,666]" pageId="1" pageNumber="145">Material</emphasis>
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. Holotype MNHN.F.
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<accessionNumber id="9404FCE1FF80FF89FDD4FD1AFDC033B1" box="[547,638,641,666]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A57537" pageId="1" pageNumber="145" type="EnaNcbi">A57537</accessionNumber>
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(PA 2496, sex unknown), stored in the laboratory of palaeontology, MNHN, Paris.
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</paragraph>
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<paragraph id="8BE86102FF80FF89FF30FD52FEDD322D" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
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<materialsCitation id="3B3F6B5FFF80FF89FF30FD52FEDD322D" ID-GBIF-Occurrence="1341384676" collectorName="Farm Le Quesnoy" country="France" county="Oise Department" latitude="49.325275" location="Chevriere" longLatPrecision="18" longitude="2.672778" municipality="Creil" pageId="1" pageNumber="145" specimenCount="1" typeStatus="holotype">
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<emphasis id="B923BD10FF80FF89FF30FD52FEDC33C9" bold="true" box="[199,354,713,738]" pageId="1" pageNumber="145">
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<typeStatus id="54ECDFA0FF80FF89FF30FD52FEBC33C9" box="[199,258,713,738]" pageId="1" pageNumber="145">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectorName id="26A204D4FF80FF89FE82FD51FDF433C9" box="[373,586,714,738]" pageId="1" pageNumber="145">Farm Le Quesnoy</collectorName>
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,
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<geoCoordinate id="EE6307C5FF80FF89FDADFD51FD5233CA" box="[602,748,714,738]" direction="north" orientation="latitude" pageId="1" pageNumber="145" precision="15" value="49.325275">49°19'31" N</geoCoordinate>
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,
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<geoCoordinate id="EE6307C5FF80FF89FD0BFD51FCC333CA" box="[764,893,714,738]" direction="east" orientation="longitude" pageId="1" pageNumber="145" precision="15" value="2.672778">2°40'22" E</geoCoordinate>
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,
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<location id="8E8837D9FF80FF89FC7AFD52FC4033C9" LSID="urn:lsid:plazi:treatment:03FED014FF80FF89FF60FE69FC3737C8:8E8837D9FF80FF89FC7AFD52FC4033C9" box="[909,1022,713,738]" country="France" county="Oise Department" latitude="49.325275" longLatPrecision="18" longitude="2.672778" municipality="Creil" name="Chevriere" pageId="1" pageNumber="145">Chevrière</location>
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,
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<location id="8E8837D9FF80FF89FBF8FD52FBDE33C9" LSID="urn:lsid:plazi:treatment:03FED014FF80FF89FF60FE69FC3737C8:8E8837D9FF80FF89FBF8FD52FBDE33C9" box="[1039,1120,713,738]" country="France" county="Oise Department" latitude="49.325275" longLatPrecision="18" longitude="2.672778" municipality="Creil" name="Region" pageId="1" pageNumber="145">Region</location>
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of
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<collectingMunicipality id="6B8CFB78FF80FF89FB79FD52FB7733C9" box="[1166,1225,713,738]" pageId="1" pageNumber="145">Creil</collectingMunicipality>
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||
,
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<collectingCounty id="6289198EFF80FF89FB2EFD52FA2233CA" box="[1241,1436,713,738]" pageId="1" pageNumber="145">Oise Department</collectingCounty>
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(northern
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<collectingCountry id="F3402192FF80FF89FEF1FD75FEE9322D" box="[262,343,750,774]" name="France" pageId="1" pageNumber="145">France</collectingCountry>
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||
).
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8BE86102FF80FF89FF30FC8AFE383265" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
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||
<emphasis id="B923BD10FF80FF89FF30FC8AFED83201" bold="true" box="[199,358,785,810]" pageId="1" pageNumber="145">
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<typeStatus id="54ECDFA0FF80FF89FF30FC8AFEBC3201" box="[199,258,785,810]" pageId="1" pageNumber="145">Type</typeStatus>
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horizon
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</emphasis>
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. Lowermost Eocene amber, Sparnacian (53 Ma), level MP7 of the mammal fauna of Dormaal (
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<bibRefCitation id="EFC61CF3FF80FF89FF69FCADFEC43265" author="Brasero" box="[158,378,821,846]" pageId="1" pageNumber="145" refString="Brasero, N., Nel, A. & Michez, D. (2009) Insects from the Early Eocene amber of Oise (France): diversity and palaeontological significance. Denisia, 26, 41 - 52." type="journal article" year="2009">
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Brasero
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<emphasis id="B923BD10FF80FF89FF09FCADFE913266" box="[254,303,821,845]" italics="true" pageId="1" pageNumber="145">et al</emphasis>
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. 2009
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D3289FF80FF89FF30FCC2FAEF32F5" pageId="1" pageNumber="145" type="diagnosis">
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<paragraph id="8BE86102FF80FF89FF30FCC2FAEF32F5" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
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<emphasis id="B923BD10FF80FF89FF30FCC2FE843259" bold="true" box="[199,314,857,882]" pageId="1" pageNumber="145">Diagnosis</emphasis>
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. Clypeus bulbous; postspiracular sclerite smooth and flat; hind tibial spur one; R2+3 well separated from R1 at apices; M3 present, reaching wing margin; laterotergite setose; macrochaetae of hind tibia absent; palpus two-segmented; scale-like thoracic setae absent; first antennal flagellomere kidney-shaped, convex below arista, arista subterminal and anterodorsal; mandibles present; wing ca.
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<quantity id="4CAFCCE7FF80FF89FC4EFC5DFBB132F6" box="[953,1039,966,990]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="145" unit="mm" value="4.2">4.2 mm</quantity>
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long; body ca.
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<quantity id="4CAFCCE7FF80FF89FB4BFC5EFAAC32F6" box="[1212,1298,965,990]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="145" unit="mm" value="5.1">5.1 mm</quantity>
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long.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D3289FF80FF89FF30FC72FC3737C8" pageId="1" pageNumber="145" type="description">
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<paragraph id="8BE86102FF80FF89FF30FC72FC05359E" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
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<emphasis id="B923BD10FF80FF89FF30FC72FEEF3529" bold="true" box="[199,337,1001,1026]" pageId="1" pageNumber="145">Description</emphasis>
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||
. Body ca.
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||
<quantity id="4CAFCCE7FF80FF89FE2FFC72FD8D352A" box="[472,563,1001,1026]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="145" unit="mm" value="5.1">5.1 mm</quantity>
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||
long. Head
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<quantity id="4CAFCCE7FF80FF89FD3EFC71FC8C352A" box="[713,818,1002,1026]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="145" unit="mm" value="0.71">0.71 mm</quantity>
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||
long, ca. 1.0 mm high. Clypeus bulbous. Face bare between eyes. Scape
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||
<quantity id="4CAFCCE7FF80FF89FE64FB95FE42350E" box="[403,508,1038,1062]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="145" unit="mm" value="0.07">0.07 mm</quantity>
|
||
long,
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||
<quantity id="4CAFCCE7FF80FF89FDBBFB95FD0B350E" box="[588,693,1038,1062]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="145" unit="mm" value="0.08">0.08 mm</quantity>
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||
high, clearly larger than pedicel. First flagellomere of antenna kidney-shaped, convex below arista, laterally compressed, enlarged, bearing a long subterminal anterodorsal arista. Eyes large, without visible setae; distinctly separated. Labella ca.
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||
<quantity id="4CAFCCE7FF80FF89FC65FBCDFC473546" box="[914,1017,1110,1134]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="145" unit="mm" value="0.79">0.79 mm</quantity>
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||
long, slightly shorter than height of head. Palpus two-segmented, distal segment setose,
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||
<quantity id="4CAFCCE7FF80FF89FD2BFBE1FCFE35BA" box="[732,832,1146,1170]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="145" unit="mm" value="0.24">0.24 mm</quantity>
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||
long, longer than proximal segment, ca.
|
||
<quantity id="4CAFCCE7FF80FF89FAFFFBE1FAE335BA" box="[1288,1373,1146,1170]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="145" unit="mm" value="0.2">0.2 mm</quantity>
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||
long. Lacinia longer than palpus, with serrated tip. Mandibles long and acute.
|
||
</paragraph>
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||
<paragraph id="8BE86102FF80FF89FF30FB5AFA8237B0" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
|
||
Thorax
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||
<quantity id="4CAFCCE7FF80FF89FEDEFB5AFE3435F2" box="[297,394,1217,1242]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="145" unit="mm" value="1.6">1.6 mm</quantity>
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||
long. Dorsocentral bristles of mesonotum absent; all dorsal setae of equal length. Anepisternum setulose throughout posterior half. Laterotergite setose. Postspiracular sclerite smooth and flat. Prescutellum present. Subscutellum inconspicuous. Wing apparently hyaline, without markings or membrane lightly to moderately infuscate, brownish. Lower calypter reduced. Upper calypter well developed, with broad curvature, lobe-like. Wing ca.
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||
<quantity id="4CAFCCE7FF80FF89FE04FAC9FDF53442" box="[499,587,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="145" unit="mm" value="4.2">4.2 mm</quantity>
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||
long,
|
||
<quantity id="4CAFCCE7FF80FF89FD62FACAFD533442" box="[661,749,1361,1386]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="145" unit="mm" value="1.6">1.6 mm</quantity>
|
||
wide. Apex of costa not preserved. Humeral crossvein well developed. Crossvein sc-r weakly developed, positioned distal to h a little less than length of h. R1 and R2+3 separated at wing margin. Dorsal side of R1 setulose, ventral side bare. All other wing veins without setulae. Apical third of R2+3 ultimately bends anteriorly slightly, toward leading edge of wing margin. Length of R2+3 probably shorter than R5. Base of R4+5 fork proximal distal end of cell dm. R4 nearly straight apically. R5 probably posterior to wing tip, clearly longer than R4+5. Crossvein r-m at proximal one-third of discal cell. M3 present. Crossvein m-cu present. Origin of CuA1 at cell bm near discal cell. M3 cell at margin convergent. CuA2 length ca. 2/3 the length of posterior vein of cell bm. Alula with broad, evenly rounded curvature. Anal lobe well developed. Cell cup open. Halter knob approximately 3/5 length of stem. Tibial spur formula 0:2:1. Hind tibial macrochaetae absent.
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||
</paragraph>
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||
<paragraph id="8BE86102FF80FF89FF30F93DFC3737C8" blockId="1.[151,1437,605,1763]" pageId="1" pageNumber="145">
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Abdomen ca.
|
||
<quantity id="4CAFCCE7FF80FF89FE9AF93CFE793795" box="[365,455,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="145" unit="mm" value="2.4">2.4 mm</quantity>
|
||
long,
|
||
<quantity id="4CAFCCE7FF80FF89FDE0F93CFDCC3795" box="[535,626,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="145" unit="mm" value="1.2">1.2 mm</quantity>
|
||
high. Abdominal segments evenly tapered. Segmentation not clearly visible, last 3 abdominal segments telescoped; genitalia not visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |