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<document id="424BFDD858678C2A8F4B63C3E38D03C9" ID-DOI="10.11646/zootaxa.3957.2.3" ID-GBIF-Dataset="cce31970-05a8-4d3e-b727-2efac5c4f4f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="243146" ID-ZooBank="E19B5C42-F9CD-4123-8138-249C3CBEEFC7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461060656996" checkinUser="plazi" docAuthor="Kubendran, T., Balasubramanian, C., Selvakumar, C., Gattolliat, J. L. &amp; Sivaramakrishnan, K. G." docDate="2015" docId="03FE87D163500855FF38F997DC01FDB5" docLanguage="en" docName="zt03957p200.pdf" docOrigin="Zootaxa 3957 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Labiobaetis jacobusi Kubendran &amp; Balasubramanian, n. sp." docType="treatment" docVersion="8" lastPageNumber="199" masterDocId="FFC7FFA96357085EFFAFFFD3DD79FF88" masterDocTitle="Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India)" masterLastPageNumber="200" masterPageNumber="188" pageNumber="195" updateTime="1698582100768" updateUser="plazi">
<mods:mods id="8228F5046941A8EAC86D371659AFA5D5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1CE357861C01B4430C4C0E070A1D8B8D">
<mods:title id="208391F7C27A85629A9020B0FBBBC16A">Contribution to the knowledge of Tenuibaetis Kang &amp; Yang 1994, Nigrobaetis Novikova &amp; Kluge 1987 and Labiobaetis Novikova &amp; Kluge 1987 (Ephemeroptera: Baetidae) from the Western Ghats (India)</mods:title>
</mods:titleInfo>
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<mods:role id="494C86A616359C181FB4CE27B2B650DF">
<mods:roleTerm id="45C4CAF44C7DD113968255B9E1E22840">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A3E971367C9653E19D8354E8832AF45">Kubendran, T.</mods:namePart>
</mods:name>
<mods:name id="BA90513CDE34FBDA5725B39D300AEC67" type="personal">
<mods:role id="BCFBA95251FCC2859144218221598B7B">
<mods:roleTerm id="EE7550AAFB3BA9F37C04EC16E0365122">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25FCACFF19EBAA67CD6B3FA342C49660">Balasubramanian, C.</mods:namePart>
</mods:name>
<mods:name id="C219C381687FE0AB9878246F203A720D" type="personal">
<mods:role id="36FF931A9488746C746E5CB43C295E48">
<mods:roleTerm id="B6F2174C380CA5390AEB312F9A59183A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9DB374CABB10F83E665C81CD8D930B1">Selvakumar, C.</mods:namePart>
</mods:name>
<mods:name id="9C39F7354EC9376F41CA989287154471" type="personal">
<mods:role id="68D35B868FD9AB4C294E0A8DB2191B44">
<mods:roleTerm id="A3D89136154B9BF273BA5A1F074CFD27">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9591712E33E9D78FAB7628F00486513">Gattolliat, J. L.</mods:namePart>
</mods:name>
<mods:name id="C444D73B40BB5EEFFFF40C6483A8D947" type="personal">
<mods:role id="DE4659F76BBF8C1DDC19B7A947DC191D">
<mods:roleTerm id="8C4E7D1CE0D3B85BEBE321D50E6B00B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F68365D3183B261B2EEBF5397F347B19">Sivaramakrishnan, K. G.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CDEBB4821FE37AB532E665D204879B9A">text</mods:typeOfResource>
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<mods:titleInfo id="C4EDE7AE61FE754CDD5402CF6B0BE96F">
<mods:title id="1018495F7717FFE50CF0A8C128E39FE3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9EADF425B026330513AEA29DC5EB78D8">
<mods:date id="E1F93AF65ADC7CEDF76978F7B571A475">2015</mods:date>
<mods:detail id="87AEA63B9C9D3EAEA30179E065103931" type="volume">
<mods:number id="7CBC60B1693C82A387E8013C4F9D3BEE">3957</mods:number>
</mods:detail>
<mods:detail id="A435768C0D3E55CCA7854F9B64C95EEE" type="issue">
<mods:number id="F9883AF6AA3799F5886023D251857AA2">2</mods:number>
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<mods:start id="131B49D7A24CC52AACE9A02D2B7D99E4">188</mods:start>
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<mods:classification id="37A6D7A20CE5166069B89F4503A490A9">journal article</mods:classification>
<mods:identifier id="A8C48E5BB0A5856C122AF8C036405635" type="DOI">10.11646/zootaxa.3957.2.3</mods:identifier>
<mods:identifier id="FF14C0D0525D1C0780D12F86B6575E72" type="GBIF-Dataset">cce31970-05a8-4d3e-b727-2efac5c4f4f9</mods:identifier>
<mods:identifier id="DED1A7A677CFBB664753C37718168B12" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0ACD5A59C697AC699C6CE16914782176" type="Zenodo-Dep">243146</mods:identifier>
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</mods:mods>
<treatment id="03FE87D163500855FF38F997DC01FDB5" ID-DOI="http://doi.org/10.5281/zenodo.6113279" ID-GBIF-Taxon="119614176" ID-Zenodo-Dep="6113279" LSID="urn:lsid:plazi:treatment:03FE87D163500855FF38F997DC01FDB5" httpUri="http://treatment.plazi.org/id/03FE87D163500855FF38F997DC01FDB5" lastPageId="11" lastPageNumber="199" pageId="7" pageNumber="195">
<subSubSection id="C34D654C63500859FF38F997DC62F9F7" pageId="7" pageNumber="195" type="nomenclature">
<paragraph id="8BE836C763500859FF38F997DE0EF9D6" blockId="7.[151,887,1604,1663]" box="[151,887,1604,1630]" pageId="7" pageNumber="195">
<heading id="D0A081AB63500859FF38F997DE0EF9D6" bold="true" box="[151,887,1604,1630]" fontSize="11" level="1" pageId="7" pageNumber="195" reason="1">
<emphasis id="B923EAD563500859FF38F997DE0EF9D6" bold="true" box="[151,887,1604,1630]" pageId="7" pageNumber="195">
<taxonomicName id="4C574D4463500859FF38F997DE54F9D6" authority="Kubendran &amp; Balasubramanian" authorityName="Kubendran &amp; Balasubramanian" box="[151,813,1604,1630]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="jacobusi" status="sp. nov.">
<emphasis id="B923EAD563500859FF38F997DCF6F9D6" bold="true" box="[151,399,1604,1630]" italics="true" pageId="7" pageNumber="195">Labiobaetis jacobusi</emphasis>
Kubendran &amp; Balasubramanian
</taxonomicName>
<taxonomicNameLabel id="A21057AE63500859FC9AF997DE0EF9D6" box="[821,887,1604,1630]" pageId="7" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE836C763500859FF38F9B5DC62F9F7" blockId="7.[151,887,1604,1663]" box="[151,283,1638,1663]" pageId="7" pageNumber="195">
<figureCitation id="136C2A4263500859FF38F9B5DC62F9F7" box="[151,283,1638,1663]" captionStart-0="FIGURES 40 – 48" captionStart-1="FIGURES 49 – 53" captionStart-2="FIGURES 54 – 58" captionStartId-0="8.[151,264,1577,1600]" captionStartId-1="9.[151,264,1356,1379]" captionStartId-2="10.[151,264,1597,1620]" captionTargetBox-0="[400,1187,308,1545]" captionTargetBox-1="[399,1205,568,1324]" captionTargetBox-2="[398,1187,418,1562]" captionTargetId-0="figure@8.[376,1210,301,1557]" captionTargetId-1="figure@9.[339,1248,553,1336]" captionTargetId-2="figure@10.[389,1197,409,1576]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." captionText-1="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." captionText-2="FIGURES 54 – 58. Imago of Labiobaetis jacobusi n. sp. 54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia." httpUri-0="https://zenodo.org/record/243152/files/figure.png" httpUri-1="https://zenodo.org/record/243153/files/figure.png" httpUri-2="https://zenodo.org/record/243154/files/figure.png" pageId="7" pageNumber="195">Figs. 40–58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34D654C63500857FF38F97DDE65F9DE" lastPageId="9" lastPageNumber="197" pageId="7" pageNumber="195" type="materials_examined">
<paragraph id="8BE836C763500859FF38F97DD998F89A" blockId="7.[151,1437,1710,2037]" pageId="7" pageNumber="195">
<emphasis id="B923EAD563500859FF38F97DDCFAF94F" bold="true" box="[151,387,1710,1735]" pageId="7" pageNumber="195">Material examined</emphasis>
.
<typeStatus id="54EC886563500859FE34F97DDF71F94F" box="[411,520,1710,1735]" pageId="7" pageNumber="195" type="holotype">Holotype</typeStatus>
: male larva:
<collectingCountry id="F340765763500859FD1CF97CDE7CF94F" box="[691,773,1711,1735]" name="India" pageId="7" pageNumber="195">INDIA</collectingCountry>
, Tamil Nadu, Theni, Valipparai stream, Vaigai River,
<geoCoordinate id="EE63500063500859FF38F905DC45F965" box="[151,316,1746,1774]" direction="north" orientation="latitude" pageId="7" pageNumber="195" precision="1" value="9.726575">9o43’35.67” N</geoCoordinate>
,
<geoCoordinate id="EE63500063500859FEE6F905DC81F965" box="[329,504,1746,1774]" direction="east" orientation="longitude" pageId="7" pageNumber="195" precision="1" value="77.51674">77o31’00.24” E</geoCoordinate>
,
<quantity id="4CAF9B2263500859FDA9F905DF27F965" box="[518,606,1750,1774]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="195" unit="m" value="1300.0">1300 m</quantity>
,
<date id="FFE9100763500859FDC4F906DF92F966" box="[619,747,1749,1774]" pageId="7" pageNumber="195" value="2012-07-29">29.vii.2012</date>
(ZSI), Colls. Balasubramanian, Kubendran and Selvakumar.
<typeStatus id="54EC886563500859FF38F929DC70F89A" box="[151,265,1786,1810]" pageId="7" pageNumber="195" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D51FD4E63500859FEBCF929DC24F89A" box="[275,349,1785,1810]" pageId="7" pageNumber="195" type="male">4 male</specimenCount>
larvae,
<specimenCount id="9D51FD4E63500859FE18F92ADF61F89A" box="[439,536,1785,1810]" pageId="7" pageNumber="195" type="female">6 female</specimenCount>
larvae and
<specimenCount id="9D51FD4E63500859FD35F92ADF9FF89A" box="[666,742,1785,1810]" pageId="7" pageNumber="195" type="male">6 male</specimenCount>
imagos, same data as
<typeStatus id="54EC886563500859FC4FF92AD93BF89A" box="[992,1090,1785,1810]" pageId="7" pageNumber="195" type="holotype">holotype</typeStatus>
(ZSI,
<collectionCode id="ED46AE0263500859FB24F929D9A2F89A" box="[1163,1243,1786,1810]" pageId="7" pageNumber="195">MZTC</collectionCode>
)
</paragraph>
<paragraph id="8BE836C763500859FF68F8CED9EAF87D" blockId="7.[151,1437,1710,2037]" pageId="7" pageNumber="195">
<emphasis id="B923EAD563500859FF68F8CEDF46F8BE" bold="true" box="[199,575,1821,1846]" pageId="7" pageNumber="195">Additional material examined</emphasis>
.
<specimenCount id="9D51FD4E63500859FDFAF8CDDFD3F8BE" box="[597,682,1821,1846]" pageId="7" pageNumber="195" type="male">1 male</specimenCount>
larva (on slide) and one female larva,
<collectingCountry id="F340765763500859FB0EF8CDD98AF8BE" box="[1185,1267,1822,1846]" name="India" pageId="7" pageNumber="195">INDIA</collectingCountry>
, Tamil Nadu, Kodaikanal, Bas. Manjalaru, Riv. Mulaiyaru,
<geoCoordinate id="EE63500063500859FD64F896DE0FF8D5" box="[715,886,1857,1885]" direction="north" orientation="latitude" pageId="7" pageNumber="195" precision="1" value="10.238887">10o14’19.99”N</geoCoordinate>
,
<geoCoordinate id="EE63500063500859FC23F896D94DF8D5" box="[908,1076,1857,1885]" direction="east" orientation="longitude" pageId="7" pageNumber="195" precision="1" value="77.48886">77o29’19.90”E</geoCoordinate>
,
<quantity id="4CAF9B2263500859FBE5F897D9D2F8D4" box="[1098,1195,1860,1885]" metricMagnitude="3" metricUnit="m" metricValue="1.216" pageId="7" pageNumber="195" unit="m" value="1216.0">1216 m</quantity>
,
<date id="FFE9100763500859FB6EF897D83DF8D5" box="[1217,1348,1860,1885]" pageId="7" pageNumber="195" value="2012-07-29">29.vii.2012</date>
, Colls. Balasubramanian., Kubendran and Selvakumar.
<specimenCount id="9D51FD4E63500859FD6EF8BBDE56F809" box="[705,815,1896,1921]" pageId="7" pageNumber="195" type="larva">16 larvae</specimenCount>
,
<collectingCountry id="F340765763500859FC91F8BADEE9F809" box="[830,912,1897,1921]" name="India" pageId="7" pageNumber="195">INDIA</collectingCountry>
, Tamil Nadu, Chinnasuruli, Bas. Vaigai, Riv. Suruli falls,
<geoCoordinate id="EE63500063500859FE95F85CDCAEF820" box="[314,471,1932,1960]" direction="north" orientation="latitude" pageId="7" pageNumber="195" precision="1" value="9.709769">9o42’35.17”N</geoCoordinate>
,
<geoCoordinate id="EE63500063500859FE40F843DFE1F820" box="[495,664,1932,1960]" direction="east" orientation="longitude" pageId="7" pageNumber="195" precision="1" value="77.424614">77o25’28.62”E</geoCoordinate>
,
<quantity id="4CAF9B2263500859FD1FF85CDE7CF82F" box="[688,773,1935,1960]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="7" pageNumber="195" unit="m" value="605.0">605 m</quantity>
,
<date id="FFE9100763500859FCB1F85CDEE1F820" box="[798,920,1935,1960]" pageId="7" pageNumber="195" value="2012-09-15">15.ix.2012</date>
, Colls. Balasubramanian, Kubendran and Selvakumar.
<specimenCount id="9D51FD4E63500859FE9EF864DCDDF847" box="[305,420,1974,1999]" pageId="7" pageNumber="195" type="larva">20 larvae</specimenCount>
,
<collectingCountry id="F340765763500859FE18F864DF70F847" box="[439,521,1975,1999]" name="India" pageId="7" pageNumber="195">INDIA</collectingCountry>
, Tamil Nadu, Megamalai, Bas. Highwavys, Riv. Megamalai,
<geoCoordinate id="EE63500063500859FB55F864D8EEF847" box="[1274,1431,1971,1999]" direction="north" orientation="latitude" pageId="7" pageNumber="195" precision="1" value="9.727366">9o43’38.52”N</geoCoordinate>
,
<geoCoordinate id="EE63500063500859FF38F80DDC46F87D" box="[151,319,2009,2037]" direction="east" orientation="longitude" pageId="7" pageNumber="195" precision="1" value="77.36092">77o21’39.34”E</geoCoordinate>
,
<quantity id="4CAF9B2263500859FEE5F80FDCD8F87D" box="[330,417,2012,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.4849999999999999" pageId="7" pageNumber="195" unit="m" value="1485.0">1485 m</quantity>
,
<date id="FFE9100763500859FE02F80FDF56F87D" box="[429,559,2012,2037]" pageId="7" pageNumber="195" value="2012-12-20">20.xii.2012</date>
, Colls. Balasubramanian, Kubendran and Selvakumar.
</paragraph>
<paragraph id="8BE836C7635F0856FF68FF44D8F1FF70" blockId="8.[151,1436,151,248]" pageId="8" pageNumber="196">
<emphasis id="B923EAD5635F0856FF68FF44DC09FF38" bold="true" box="[199,368,151,176]" pageId="8" pageNumber="196">Mature larva.</emphasis>
Maximal length: Body 5.0–6.0 mm; cerci 2.5–3.0 mm; terminal filament 2.0–
<quantity id="4CAF9B22635F0856FAACFF44D827FF27" box="[1283,1374,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="196" unit="mm" value="2.5">2.5 mm</quantity>
. Size variable between different populations. Colouration: Head uniformly brown without distinct pattern; thorax light brown, legs yellowish brown; abdominal tergites uniformly light brown (
<figureCitation id="136C2A42635F0856FC66FF0CD964FF70" box="[969,1053,223,248]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 40</figureCitation>
); sternites pale yellow (
<figureCitation id="136C2A42635F0856FA88FF0CD803FF70" box="[1319,1402,223,248]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 41</figureCitation>
).
</paragraph>
<caption id="DF28664F635F0856FF38F9FAD9B3F9D7" httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196" targetBox="[400,1187,308,1545]" targetPageId="8">
<paragraph id="8BE836C7635F0856FF38F9FAD9B3F9D7" blockId="8.[151,1436,1577,1631]" pageId="8" pageNumber="196">
<emphasis id="B923EAD5635F0856FF38F9FADC29F9B7" bold="true" box="[151,336,1577,1600]" pageId="8" pageNumber="196">FIGURES 40–48</emphasis>
. Larva of
<taxonomicName id="4C574D44635F0856FE6DF9F9DFE0F9C8" box="[450,665,1578,1600]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="196" phylum="Arthropoda" rank="species" species="jacobusi" status="sp. nov.">
<emphasis id="B923EAD5635F0856FE6DF9F9DFE0F9C8" box="[450,665,1578,1600]" italics="true" pageId="8" pageNumber="196">Labiobaetis jacobusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21057AE635F0856FD0EF9F9DFA4F9B7" box="[673,733,1578,1599]" pageId="8" pageNumber="196" rank="species">
<emphasis id="B923EAD5635F0856FD0EF9F9DFA4F9B7" bold="true" box="[673,733,1578,1599]" pageId="8" pageNumber="196">n. sp.</emphasis>
</taxonomicNameLabel>
40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium.
</paragraph>
</caption>
<paragraph id="8BE836C7635F0857FF68F958D81CFEED" blockId="8.[151,1437,1675,2024]" lastBlockId="9.[151,1437,151,501]" lastPageId="9" lastPageNumber="197" pageId="8" pageNumber="196">
<emphasis id="B923EAD5635F0856FF68F958DC7EF92C" bold="true" box="[199,263,1675,1700]" pageId="8" pageNumber="196">Head</emphasis>
: Antennae (
<figureCitation id="136C2A42635F0856FE36F958DC8AF92C" box="[409,499,1675,1700]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 42</figureCitation>
): Scape and pedicel bare; distolateral notch on scape absent. Labrum (
<figureCitation id="136C2A42635F0856FA99F958D8F6F92C" box="[1334,1423,1675,1700]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 43</figureCitation>
): rectangular, approximately 0.8 times as long as wide, anterior margin with a small median notch; dorsal surface with about 12 spatulate submarginal setae in contiguous row, fine and simple setae scattered medially; ventrally bordered with feathered setae along margin and a distomedial arc of fine setae. Hypopharynx (
<figureCitation id="136C2A42635F0856FB48F924D839F898" box="[1255,1344,1783,1808]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 44</figureCitation>
): lingua covered with short thin setae; stout and longer medially; superlingua with thin setae apically and laterally. Left mandible (
<figureCitation id="136C2A42635F0856FEBDF8ECDC13F8D0" box="[274,362,1855,1880]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 45</figureCitation>
): with incisors composed of 7 denticles; prostheca with 5 broad denticles and comb-shaped structure; tuft of setae at apex of mola absent. Right mandible (
<figureCitation id="136C2A42635F0856FCDBF8B0DEB5F8F4" box="[884,972,1891,1916]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 46</figureCitation>
): with incisors composed of 7 denticles; prostheca with 10 denticles; margin between prostheca and mola and without tubercle; tuft of setae at apex of mola present. Maxillae (
<figureCitation id="136C2A42635F0856FEC6F878DCC2F84C" box="[361,443,1963,1988]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="8" pageNumber="196">Fig. 47</figureCitation>
) with 4 caninae; lacinia with 2 rows of setae, one row with abundant small setae ending with stout and long setae, second row with 2 long stout dentisetae; two thin setae at base of caninae; palp twosegmented, longer than galea-lacinia; segment 2 longer than segment 1, apically rounded with broadly concave subapical margin, lateral margin with about 9–10 thin setae. Labium (
<figureCitation id="136C2A42635E0857FC32FF6FDE89FF5C" box="[925,1008,188,213]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="9" pageNumber="197">Fig. 48</figureCitation>
): glossae with one row of simple setae on inner margin; glossae shorter than paraglossae, paraglossae with 2 rows of apical feathered setae; labial palp three-segmented; segment 1 slender, subequal in length to segment 2 and 3 combined; segment 2 with a broad falcated and apically pointed distomedial projection covered with short thin and simple setae scattered over surface; segment 3 somewhat globular, with stout and pointed setae and thin setae scattered on dorsal surface.
</paragraph>
<paragraph id="8BE836C7635E0857FF68FEBCD9FBFE7D" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="197">
<emphasis id="B923EAD5635E0857FF68FEBCDC59FE00" bold="true" box="[199,288,367,392]" pageId="9" pageNumber="197">Thorax</emphasis>
: Foreleg (
<figureCitation id="136C2A42635E0857FE3DFEBCDC9CFE00" box="[402,485,367,392]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="9" pageNumber="197">Fig. 49</figureCitation>
): dorsal margin of femora with a row of about 7 long stout pointed setae, without row of setae subparallel to margin; ventral margin with about 11 pointed stout setae; dorsal margin of tibiae with a row of setae; dorsal margin of tarsi bare; ventral margin with about 11 pointed stout setae; tarsal claw with one row of about 11 acute teeth (
<figureCitation id="136C2A42635E0857FE29FE0FDCA3FE7D" box="[390,474,476,501]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="9" pageNumber="197">Fig. 50</figureCitation>
); mid and hind leg similar to foreleg. Hindwing pads absent.
</paragraph>
<caption id="DF28664F635E0857FF38FA9FDC48FA0A" httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="9" pageNumber="197" targetBox="[399,1205,568,1324]" targetPageId="9">
<paragraph id="8BE836C7635E0857FF38FA9FDC48FA0A" blockId="9.[151,1436,1356,1410]" pageId="9" pageNumber="197">
<emphasis id="B923EAD5635E0857FF38FA9FDC2AFAEA" bold="true" box="[151,339,1356,1379]" pageId="9" pageNumber="197">FIGURES 49–53.</emphasis>
Larva of
<taxonomicName id="4C574D44635E0857FE19FA9EDFF3FAEB" box="[438,650,1357,1379]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="197" phylum="Arthropoda" rank="species" species="jacobusi" status="sp. nov.">
<emphasis id="B923EAD5635E0857FE19FA9EDFF3FAEB" box="[438,650,1357,1379]" italics="true" pageId="9" pageNumber="197">Labiobaetis jacobusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21057AE635E0857FD3FFA9DDFB0FAEB" box="[656,713,1358,1379]" pageId="9" pageNumber="197" rank="species">
<emphasis id="B923EAD5635E0857FD3FFA9DDFB0FAEB" bold="true" box="[656,713,1358,1379]" pageId="9" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga.
</paragraph>
</caption>
<paragraph id="8BE836C7635E0857FF68FA7EDE65F9DE" blockId="9.[151,1437,1453,2018]" pageId="9" pageNumber="197">
<emphasis id="B923EAD5635E0857FF68FA7EDC45FA4E" bold="true" box="[199,316,1453,1478]" pageId="9" pageNumber="197">Abdomen</emphasis>
: Single lamellate gills on segments 2–7 (
<figureCitation id="136C2A42635E0857FC8FFA7EDE0FFA4E" box="[800,886,1453,1478]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="9" pageNumber="197">Fig. 51</figureCitation>
), elongated with serrated margins; trachea well visible; each gill with a dark band surrounding margin. Paraproct (
<figureCitation id="136C2A42635E0857FCD2FA02DEB7FA62" box="[893,974,1489,1514]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="9" pageNumber="197">Fig. 52</figureCitation>
) margin with numerous small to medium pointed spines, with scale bases; posterolateral extension with scale bases, margin with irregular spines. Tergites with numerous scale bases; distal-margin with broad triangular spines (
<figureCitation id="136C2A42635E0857FC7BF9CAD952F9BA" box="[980,1067,1561,1586]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="9" pageNumber="197">Fig. 53</figureCitation>
). Caudal filaments light brown; terminal filament shorter than cerci and with lateral hairs.
</paragraph>
</subSubSection>
<subSubSection id="C34D654C635E0857FF68F9B2DE3EF835" pageId="9" pageNumber="197" type="description">
<paragraph id="8BE836C7635E0857FF68F9B2DE3EF835" blockId="9.[151,1437,1453,2018]" pageId="9" pageNumber="197">
<emphasis id="B923EAD5635E0857FF68F9B2DC2CF9F2" bold="true" box="[199,341,1633,1658]" pageId="9" pageNumber="197">Male imago</emphasis>
: Length: body 9.0 mm; fore wing
<quantity id="4CAF9B22635E0857FD4BF9B2DE47F9F1" box="[740,830,1633,1658]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="197" unit="mm" value="7.5">7.5 mm</quantity>
. Foreleg: femur
<quantity id="4CAF9B22635E0857FC53F9B1D92FF9F1" box="[1020,1110,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="197" unit="mm" value="1.2">1.2 mm</quantity>
; tibia
<quantity id="4CAF9B22635E0857FB31F9B2D981F9F1" box="[1182,1272,1633,1658]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="197" unit="mm" value="1.5">1.5 mm</quantity>
; tarsi
<quantity id="4CAF9B22635E0857FA91F9B1D8E0F9F1" box="[1342,1433,1634,1658]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="197" unit="mm" value="0.4">0.4 mm</quantity>
. Cerci
<quantity id="4CAF9B22635E0857FF75F956DC2CF915" box="[218,341,1669,1694]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="9" pageNumber="197" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
. Colouration: Head (
<figureCitation id="136C2A42635E0857FD92F956DFC5F916" box="[573,700,1669,1694]" captionStart="FIGURES 54 – 58" captionStartId="10.[151,264,1597,1620]" captionTargetBox="[398,1187,418,1562]" captionTargetId="figure@10.[389,1197,409,1576]" captionTargetPageId="10" captionText="FIGURES 54 – 58. Imago of Labiobaetis jacobusi n. sp. 54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia." httpUri="https://zenodo.org/record/243154/files/figure.png" pageId="9" pageNumber="197">Figs 55, 56</figureCitation>
) dark brown; scape and pedicel medium to dark brown; flagellum medium brown; facetted surface of compound eyes reddish brown; lateral face reddish brown. Thorax: dark brown. Legs: femora light brown, tibiae light yellowish brown, tarsi light brown. Forewings (
<figureCitation id="136C2A42635E0857FBF9F91ED9D1F96E" box="[1110,1192,1741,1766]" captionStart="FIGURES 54 – 58" captionStartId="10.[151,264,1597,1620]" captionTargetBox="[398,1187,418,1562]" captionTargetId="figure@10.[389,1197,409,1576]" captionTargetPageId="10" captionText="FIGURES 54 – 58. Imago of Labiobaetis jacobusi n. sp. 54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia." httpUri="https://zenodo.org/record/243154/files/figure.png" pageId="9" pageNumber="197">Fig. 54</figureCitation>
) hyaline; pterostigma with 4 cross-veins generally reaching subcostal vein; double intercalary veins shorter than distance between corresponding main veins (
<figureCitation id="136C2A42635E0857FE68F8C6DF65F8A6" box="[455,540,1813,1838]" captionStart="FIGURES 54 – 58" captionStartId="10.[151,264,1597,1620]" captionTargetBox="[398,1187,418,1562]" captionTargetId="figure@10.[389,1197,409,1576]" captionTargetPageId="10" captionText="FIGURES 54 – 58. Imago of Labiobaetis jacobusi n. sp. 54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia." httpUri="https://zenodo.org/record/243154/files/figure.png" pageId="9" pageNumber="197">Fig. 57</figureCitation>
). Hindwings absent. Abdomen (
<figureCitation id="136C2A42635E0857FC2BF8C6DEA0F8A6" box="[900,985,1813,1838]" captionStart="FIGURES 54 – 58" captionStartId="10.[151,264,1597,1620]" captionTargetBox="[398,1187,418,1562]" captionTargetId="figure@10.[389,1197,409,1576]" captionTargetPageId="10" captionText="FIGURES 54 – 58. Imago of Labiobaetis jacobusi n. sp. 54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia." httpUri="https://zenodo.org/record/243154/files/figure.png" pageId="9" pageNumber="197">Fig. 54</figureCitation>
): Tergites I–VI whitish without marking or pattern; tergites VII-X medium brown without marking or pattern, sternites colouration similar to corresponding tergites. Genitalia (
<figureCitation id="136C2A42635E0857FEDAF88EDCB7F8FE" box="[373,462,1885,1910]" captionStart="FIGURES 54 – 58" captionStartId="10.[151,264,1597,1620]" captionTargetBox="[398,1187,418,1562]" captionTargetId="figure@10.[389,1197,409,1576]" captionTargetPageId="10" captionText="FIGURES 54 – 58. Imago of Labiobaetis jacobusi n. sp. 54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia." httpUri="https://zenodo.org/record/243154/files/figure.png" pageId="9" pageNumber="197">Fig. 58</figureCitation>
): segment 1 and 2 completely fused; inner margin of segment 1 expanded basally, progressively narrower with 6–8 thin setae; segment 2 with parallel margins; segment 3 globular; well developed sclerotized triangular process between forceps, without setae.
</paragraph>
</subSubSection>
<subSubSection id="C34D654C635E0854FF68F81ADCCCFEEC" lastPageId="10" lastPageNumber="198" pageId="9" pageNumber="197" type="diagnosis">
<paragraph id="8BE836C7635E0854FF68F81ADCCCFEEC" blockId="9.[151,1437,1453,2018]" lastBlockId="10.[151,1436,151,357]" lastPageId="10" lastPageNumber="198" pageId="9" pageNumber="197">
<emphasis id="B923EAD5635E0857FF68F81ADC43F86A" bold="true" box="[199,314,1993,2018]" pageId="9" pageNumber="197">Diagnosis</emphasis>
.
<taxonomicName id="4C574D44635E0857FEE6F81ADF4DF86A" box="[329,564,1993,2018]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="197" phylum="Arthropoda" rank="species" species="jacobusi" status="sp. nov.">
<emphasis id="B923EAD5635E0857FEE6F81ADF4DF86A" box="[329,564,1993,2018]" italics="true" pageId="9" pageNumber="197">Labiobaetis jacobusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21057AE635E0857FD93F81ADF05F86A" box="[572,636,1993,2018]" pageId="9" pageNumber="197" rank="species">
<emphasis id="B923EAD5635E0857FD93F81ADF05F86A" bold="true" box="[572,636,1993,2018]" pageId="9" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
can be differentiated from other species of the genus by the following combination of characters: (i) distolateral notch on antennal scape absent (
<figureCitation id="136C2A42635D0854FBADFF44D923FF38" box="[1026,1114,151,176]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="10" pageNumber="198">Fig. 42</figureCitation>
); (ii) a row of 12 spatulate submarginal setae on labrum (
<figureCitation id="136C2A42635D0854FE5FFF6FDF3EFF5C" box="[496,583,188,213]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="10" pageNumber="198">Fig. 43</figureCitation>
); (iii) right mandible without tubercle at inner margin between incisor and mola (
<figureCitation id="136C2A42635D0854FF70FF0CDC4CFF70" box="[223,309,223,248]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="10" pageNumber="198">Fig. 46</figureCitation>
); (iv) broadly concave subapical inner margin present on segment 2 of maxillary palp (
<figureCitation id="136C2A42635D0854FABEFF0CD81FFF70" box="[1297,1382,223,248]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="10" pageNumber="198">Fig. 47</figureCitation>
); (v) segment 2 of labial palp with broad falcate distomedial projection (
<figureCitation id="136C2A42635D0854FC19FED7D976FE94" box="[950,1039,260,285]" captionStart="FIGURES 40 – 48" captionStartId="8.[151,264,1577,1600]" captionTargetBox="[400,1187,308,1545]" captionTargetId="figure@8.[376,1210,301,1557]" captionTargetPageId="8" captionText="FIGURES 40 – 48. Larva of Labiobaetis jacobusi n. sp. 40, Dorsal view; 41, Ventral view; 42, Antenna; 43, Labrum (left: dorsal; right: ventral); 44, Hypopharynx; 45, Left mandible; 46, Right mandible; 47, Maxilla; 48, Labium." httpUri="https://zenodo.org/record/243152/files/figure.png" pageId="10" pageNumber="198">Fig. 48</figureCitation>
); (vi) hindwing pads absent; (vii) serrated edges of abdominal gills with well visible trachea on segment 2–7 (
<figureCitation id="136C2A42635D0854FC5CFEF4D93EFEC8" box="[1011,1095,295,320]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="10" pageNumber="198">Fig. 51</figureCitation>
); (viii) dorsal margin of tibiae with stout setae (
<figureCitation id="136C2A42635D0854FEFBFE9FDCD0FEEC" box="[340,425,332,357]" captionStart="FIGURES 49 – 53" captionStartId="9.[151,264,1356,1379]" captionTargetBox="[399,1205,568,1324]" captionTargetId="figure@9.[339,1248,553,1336]" captionTargetPageId="9" captionText="FIGURES 49 – 53. Larva of Labiobaetis jacobusi n. sp. 49, Foreleg; 50, Claw; 51, Gill 4; 52, Paraproct; 53, Posterior marginal spines of terga." httpUri="https://zenodo.org/record/243153/files/figure.png" pageId="10" pageNumber="198">Fig. 49</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF28664F635D0854FF38F9EEDCF6F9FA" httpUri="https://zenodo.org/record/243154/files/figure.png" pageId="10" pageNumber="198" targetBox="[398,1187,418,1562]" targetPageId="10">
<paragraph id="8BE836C7635D0854FF38F9EEDCF6F9FA" blockId="10.[151,1436,1597,1650]" pageId="10" pageNumber="198">
<emphasis id="B923EAD5635D0854FF38F9EEDC2EF9DB" bold="true" box="[151,343,1597,1620]" pageId="10" pageNumber="198">FIGURES 54–58.</emphasis>
Imago of
<taxonomicName id="4C574D44635D0854FE67F9EDDFE6F9DC" box="[456,671,1598,1620]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="jacobusi" status="sp. nov.">
<emphasis id="B923EAD5635D0854FE67F9EDDFE6F9DC" box="[456,671,1598,1620]" italics="true" pageId="10" pageNumber="198">Labiobaetis jacobusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21057AE635D0854FD07F9EDDF9DF9DB" box="[680,740,1598,1619]" pageId="10" pageNumber="198" rank="species">
<emphasis id="B923EAD5635D0854FD07F9EDDF9DF9DB" bold="true" box="[680,740,1598,1619]" pageId="10" pageNumber="198">n. sp.</emphasis>
</taxonomicNameLabel>
54, Male; 55, Lateral view of head; 56, Dorsal view of head; 57, Forewing; 58, Genitalia.
</paragraph>
</caption>
<subSubSection id="C34D654C635D0854FF68F973DC59F955" pageId="10" pageNumber="198" type="etymology">
<paragraph id="8BE836C7635D0854FF68F973DC59F955" blockId="10.[151,1436,1696,2009]" pageId="10" pageNumber="198">
<emphasis id="B923EAD5635D0854FF68F973DC3FF931" bold="true" box="[199,326,1696,1721]" pageId="10" pageNumber="198">Etymology</emphasis>
. The species is named in honour of Dr. Luke M. Jacobus for his outstanding contribution to mayfly systematics.
</paragraph>
</subSubSection>
<subSubSection id="C34D654C635D0855FF68F93BDC01FDB5" lastPageId="11" lastPageNumber="199" pageId="10" pageNumber="198" type="discussion">
<paragraph id="8BE836C7635D0855FF68F93BDFE0FE24" blockId="10.[151,1436,1696,2009]" lastBlockId="11.[151,1437,151,573]" lastPageId="11" lastPageNumber="199" pageId="10" pageNumber="198">
<emphasis id="B923EAD5635D0854FF68F93BDC3CF889" bold="true" box="[199,325,1768,1793]" pageId="10" pageNumber="198">Discussion</emphasis>
.
<taxonomicName id="4C574D44635D0854FEF6F93BDF33F889" box="[345,586,1768,1793]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="jacobusi" status="sp. nov.">
<emphasis id="B923EAD5635D0854FEF6F93BDF33F889" box="[345,586,1768,1793]" italics="true" pageId="10" pageNumber="198">Labiobaetis jacobusi</emphasis>
</taxonomicName>
<emphasis id="B923EAD5635D0854FDF7F93BDFE4F889" bold="true" box="[600,669,1768,1793]" pageId="10" pageNumber="198">
<taxonomicNameLabel id="A21057AE635D0854FDF7F93BDFE4F889" box="[600,669,1768,1793]" pageId="10" pageNumber="198" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C574D44635D0854FD4EF93BD851F889" authority="Muller-Liebenau 1982" authorityName="Muller-Liebenau" authorityYear="1982" box="[737,1320,1768,1793]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="sumigarensis">
<emphasis id="B923EAD5635D0854FD4EF93BD97FF889" box="[737,1030,1768,1793]" italics="true" pageId="10" pageNumber="198">Labiobaetis sumigarensis</emphasis>
(
<bibRefCitation id="EFC64B36635D0854FBB5F93BD859F889" author="Muller-Liebenau" box="[1050,1312,1768,1793]" pageId="10" pageNumber="198" refString="Muller-Liebenau, I. (1982) New species of the family Baetidae from the Philippines (Insecta, Ephemeroptera). Archiv fur Hydrobiologie, 94, 70 - 82." type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
)
</taxonomicName>
from the
<collectingCountry id="F3407657635D0854FF38F8DFDC6CF8AD" box="[151,277,1804,1829]" name="Philippines" pageId="10" pageNumber="198">Philippines</collectingCountry>
are the only species of
<taxonomicName id="4C574D44635D0854FDB5F8DFDFE7F8AD" box="[538,670,1804,1829]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD5635D0854FDB5F8DFDFE7F8AD" box="[538,670,1804,1829]" italics="true" pageId="10" pageNumber="198">Labiobaetis</emphasis>
</taxonomicName>
from Oriental realm with six pairs of gills, the labrum with spatulate setae, absence of hindwings and absence of antennal scape (
<bibRefCitation id="EFC64B36635D0854FCFEF8E3D958F8C1" author="Shi" box="[849,1057,1840,1865]" pageId="10" pageNumber="198" refString="Shi, W. F. &amp; Tong, X. L. (2014) The genus Labiobaetis (Ephemeroptera: Baetidae) in China, with description of a new species. Zootaxa, 3815 (3), 397 - 408. http: // dx. doi. org / 10.11646 / zootaxa. 3815.3.5" type="journal article" year="2014">Shi &amp; Tong 2014</bibRefCitation>
). The two species also possess a similar distomedial projection in segment 2 of labial palp (although more arched in
<emphasis id="B923EAD5635D0854FBCDF885D865F8E4" box="[1122,1308,1877,1901]" italics="true" pageId="10" pageNumber="198">
<taxonomicName id="4C574D44635D0854FBCDF885D861F8E4" box="[1122,1304,1877,1901]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="sumigarensis">L. sumigarensis</taxonomicName>
,
</emphasis>
<figureCitation id="136C2A42635D0854FA89F887D813F8E5" box="[1318,1386,1876,1901]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="10" pageNumber="198">Fig. 5</figureCitation>
B in
<bibRefCitation id="EFC64B36635D0854FF38F8ABDCE3F819" author="Muller-Liebenau" box="[151,410,1912,1937]" pageId="10" pageNumber="198" refString="Muller-Liebenau, I. (1982) New species of the family Baetidae from the Philippines (Insecta, Ephemeroptera). Archiv fur Hydrobiologie, 94, 70 - 82." type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
). They differ by the setation of the dorsal margin of femora (numerous short setae in
<taxonomicName id="4C574D44635D0854FA2AF8A9DC53F83D" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="sumigarensis">
<emphasis id="B923EAD5635D0854FA2AF8A9DC53F83D" italics="true" pageId="10" pageNumber="198">L. sumigarensis</emphasis>
</taxonomicName>
(
<figureCitation id="136C2A42635D0854FE95F84FDC04F83D" box="[314,381,1948,1973]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1660,1683]" captionTargetBox="[386,1191,456,1628]" captionTargetId="figure@2.[376,1211,445,1639]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Larva of Tenuibaetis frequentus (Müller-Liebenau &amp; Hubbard 1985) n. comb. 1, Dorsal view of male; 2, Ventral view of male; 3, Antenna; 4, Labrum (left: dorsal; right: ventral); 5, Hypopharynx; 6, Left mandible; 7, Right mandible; 8, Maxilla; 9, Labium." httpUri="https://zenodo.org/record/243147/files/figure.png" pageId="10" pageNumber="198">Fig. 5</figureCitation>
G in
<bibRefCitation id="EFC64B36635D0854FE16F84FDFC1F83D" author="Muller-Liebenau" box="[441,696,1948,1973]" pageId="10" pageNumber="198" refString="Muller-Liebenau, I. (1982) New species of the family Baetidae from the Philippines (Insecta, Ephemeroptera). Archiv fur Hydrobiologie, 94, 70 - 82." type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
) and long and stout setae in
<taxonomicName id="4C574D44635D0854FBABF84DD9FBF83D" box="[1028,1154,1948,1973]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="jacobusi">
<emphasis id="B923EAD5635D0854FBABF84DD9FBF83D" box="[1028,1154,1948,1973]" italics="true" pageId="10" pageNumber="198">L. jacobusi</emphasis>
</taxonomicName>
), the size (
<taxonomicName id="4C574D44635D0854FAACF84DD8F8F83D" box="[1283,1409,1948,1973]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="jacobusi">
<emphasis id="B923EAD5635D0854FAACF84DD8F8F83D" box="[1283,1409,1948,1973]" italics="true" pageId="10" pageNumber="198">L. jacobusi</emphasis>
</taxonomicName>
is more than twice the size of
<taxonomicName id="4C574D44635D0854FE5CF811DFD3F851" box="[499,682,1986,2009]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="198" phylum="Arthropoda" rank="species" species="sumigarensis">
<emphasis id="B923EAD5635D0854FE5CF811DFD3F851" box="[499,682,1986,2009]" italics="true" pageId="10" pageNumber="198">L. sumigarensis</emphasis>
</taxonomicName>
) and the number of spatulate setae on the labrum. The genus
<taxonomicName id="4C574D44635C0855FF38FF44DC62FF38" box="[151,283,151,176]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD5635C0855FF38FF44DC62FF38" box="[151,283,151,176]" italics="true" pageId="11" pageNumber="199">Labiobaetis</emphasis>
</taxonomicName>
has a wide distribution all over the world except in Central and South
<collectingCountry id="F3407657635C0855FBD8FF44D9A2FF38" box="[1143,1243,151,176]" name="United States of America" pageId="11" pageNumber="199">America</collectingCountry>
.
<bibRefCitation id="EFC64B36635C0855FB45FF44DC17FF5C" author="Lugo-Ortiz" pageId="11" pageNumber="199" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1997) Labiobaetis Novikova and Kluge (Ephemeroptera: Baetidae) from the Afrotropical region. African Entomology, 5, 241 - 260." type="journal article" year="1997">Lugo-Ortiz and McCafferty (1997)</bibRefCitation>
have expanded the concept of
<taxonomicName id="4C574D44635C0855FD4FFF6FDE1DFF5D" box="[736,868,188,213]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD5635C0855FD4FFF6FDE1DFF5D" box="[736,868,188,213]" italics="true" pageId="11" pageNumber="199">Labiobaetis</emphasis>
</taxonomicName>
to include species with or without a distolateral notch on the antennal scape, with the distomedial modification of segment 2 of maxillary palps ranging from a constriction to a pronounced concavity, with or without villopore, with gills present or absent on abdominal segment 1 and with or without hindwing pads. Genetic analysis including Afrotropical and Asian species proved that the present concept of
<taxonomicName id="4C574D44635C0855FE63FE9FDF29FEED" box="[460,592,332,357]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAD5635C0855FE63FE9FDF29FEED" box="[460,592,332,357]" italics="true" pageId="11" pageNumber="199">Labiobaetis</emphasis>
</taxonomicName>
is polyphyletic (
<bibRefCitation id="EFC64B36635C0855FCBFFE9FD976FEED" author="Monaghan" box="[784,1039,332,357]" pageId="11" pageNumber="199" refString="Monaghan, M. T., Gattolliat, J. L., Sartori, M., Elouard, J. M., James, H., Derleth, P., Glaizot, O., De Moor, F. &amp; Vogler, A. P. (2005) Trans-oceanic and endemic origins of the small minnow mayflies (Ephemeroptera, Baetidae) of Madagascar. Proceedings of the Royal Society B-Biological Sciences, 272, 1829 - 1836. http: // dx. doi. org / 10.1098 / rspb. 2005.3139" type="journal article" year="2005">
Monaghan
<emphasis id="B923EAD5635C0855FC3DFE9EDEBDFEED" box="[914,964,332,357]" italics="true" pageId="11" pageNumber="199">et al</emphasis>
. 2005
</bibRefCitation>
;
<bibRefCitation id="EFC64B36635C0855FBB3FE9FD874FEEC" author="Gattolliat" box="[1052,1293,332,357]" pageId="11" pageNumber="199" refString="Gattolliat, J. - L., Monaghan, M. T., Sartori, M., Barber-James, H. M., Elouard, J. - M., Derleth, P., Glaizot, O., de Moor, F. C. &amp; Vogler, A. P. (2008) A molecular analysis of the Afrotropical Baetidae. In: Hauer, F. R., Stanford, J. A. &amp; Newell, R. L. (Eds.), International Advances in the Ecology, Zoogeography and Systematics of Mayflies and Stoneflies. University of California Press. Berkeley, California, pp. 219 - 232. http: // dx. doi. org / 10.1525 / california / 9780520098688.003.0016" type="book chapter" year="2008">
Gattolliat
<emphasis id="B923EAD5635C0855FB21FE9ED9B1FEED" box="[1166,1224,332,357]" italics="true" pageId="11" pageNumber="199">et al.</emphasis>
2008
</bibRefCitation>
). This riddle can be only solved by comprehensive combined morphological and molecular phylogenetic and phylogeographic studies including all the different known taxa.
</paragraph>
<paragraph id="8BE836C7635C0855FF68FE64DC01FDB5" blockId="11.[151,1437,151,573]" pageId="11" pageNumber="199">
<emphasis id="B923EAD5635C0855FF68FE64DC5DFE58" bold="true" box="[199,292,439,464]" pageId="11" pageNumber="199">Ecology</emphasis>
. The
<typeStatus id="54EC8865635C0855FECAFE6BDCECFE58" box="[357,405,440,464]" pageId="11" pageNumber="199">type</typeStatus>
locality is a perennial stream of Vaigai basin with an average of
<quantity id="4CAF9B22635C0855FBDEFE6BD9E5FE47" box="[1137,1180,440,464]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="199" unit="m" value="4.0">4 m</quantity>
wide and
<quantity id="4CAF9B22635C0855FABEFE6BD82FFE47" box="[1297,1366,440,464]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="199" unit="cm" value="40.0">40 cm</quantity>
depth and moderate water current (
<quantity id="4CAF9B22635C0855FE77FE0FDF63FE7C" box="[472,538,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="199" unit="m" value="0.9">0.9 m</quantity>
/sec) on the southern Western Ghats. The water temperature ranged between 21– 24˚C (seasonal variations) and the pH between 6.7 and 7.3. Larvae were found among grasses in stream edges of medium water flow.
</paragraph>
</subSubSection>
</treatment>
</document>
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