253 lines
37 KiB
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253 lines
37 KiB
XML
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<mods:title id="4831F9416464EAA455E4C2A079A3344C">Contributions to the knowledge of the “ Staphylinuscomplex ” (Coleoptera, Staphylinidae, Staphylinini) of China. Part 11. The genus Wasmannellus Bernhauer, 1920 and Nelmanwaslus gen. nov.</mods:title>
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<mods:namePart id="D2F9DBD4C19AEA09F35B654414B2881E">Smetana, Aleš</mods:namePart>
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<mods:date id="01C6611481072C50404DA95A44E4866F">2006</mods:date>
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<treatment id="03FE87ADEE3CFFB9FED29CB9FAB6FC09" ID-DOI="http://doi.org/10.5281/zenodo.6260697" ID-GBIF-Taxon="119349416" ID-Zenodo-Dep="6260697" LSID="urn:lsid:plazi:treatment:03FE87ADEE3CFFB9FED29CB9FAB6FC09" httpUri="http://treatment.plazi.org/id/03FE87ADEE3CFFB9FED29CB9FAB6FC09" lastPageId="3" lastPageNumber="36" pageId="1" pageNumber="34">
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<subSubSection id="C34D6530EE3CFFBBFED29CB9FE15FC66" pageId="1" pageNumber="34" type="nomenclature">
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<paragraph id="8BE836BBEE3CFFBBFED29CB9FD12FC09" blockId="1.[264,655,924,985]" box="[264,655,924,950]" pageId="1" pageNumber="34">
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<heading id="D0A081D7EE3CFFBBFED29CB9FD12FC09" bold="true" box="[264,655,924,950]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
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<taxonomicName id="4C574D38EE3CFFBBFED29CB9FD12FC09" ID-CoL="8XQPK" authority="Bernhauer, 1920" authorityName="Bernhauer" authorityYear="1920" box="[264,655,924,950]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3CFFBBFED29CB9FD12FC09" bold="true" box="[264,655,924,950]" pageId="1" pageNumber="34">
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<emphasis id="B923EAA9EE3CFFBBFED29CB9FE24FC09" bold="true" box="[264,441,924,950]" italics="true" pageId="1" pageNumber="34">Wasmannellus</emphasis>
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<bibRefCitation id="EFC64B4AEE3CFFBBFE1A9CB9FD12FC09" author="Bernhauer" box="[448,655,924,950]" pageId="1" pageNumber="42" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malayischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article" year="1920">Bernhauer, 1920</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE836BBEE3CFFBBFED29C9AFE15FC66" blockId="1.[264,655,924,985]" box="[264,392,959,985]" pageId="1" pageNumber="34">
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(
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<figureCitation id="136C2A3EEE3CFFBBFECB9C9AFEE2FC66" box="[273,383,959,985]" captionStart="FIGURES 1 – 6" captionStartId="5.[264,383,1771,1795]" captionTargetBox="[294,1305,328,1744]" captionTargetId="figure@5.[285,1310,326,1744]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Details of Wasmannellus tristis (1 – 2) and Nelmanwaslus ornatus (3 – 6). 1 — right lateral sclerite of tergite 9 and tergite 10 of female genital segment; 2 — first and second gonocoxites of female genital segment; 3 — tergite 10 of male genital segment; 4 — sternite 9 of male genital segment; 5 — aedoeagus, ventral view; 6 — underside of apical portion of paramere of aedoeagus." httpUri="https://zenodo.org/record/173962/files/figure.png" pageId="1" pageNumber="34">Figs. 1, 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D6530EE3CFFBBFED29B2BFB83FB96" box="[264,1054,1038,1065]" pageId="1" pageNumber="34" type="materials_examined">
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<paragraph id="8BE836BBEE3CFFBBFED29B2BFB83FB96" blockId="1.[264,1323,1038,1905]" box="[264,1054,1038,1065]" pageId="1" pageNumber="34">
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<emphasis id="B923EAA9EE3CFFBBFED29B2BFE37FB97" bold="true" box="[264,426,1038,1064]" pageId="1" pageNumber="34">
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<typeStatus id="54EC8819EE3CFFBBFED29B2BFED8FB97" box="[264,325,1038,1064]" pageId="1" pageNumber="34">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C574D38EE3CFFBBFE689B2AFCEDFB96" authority="Bernhauer, 1920" authorityName="Bernhauer" authorityYear="1920" box="[434,880,1039,1065]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B923EAA9EE3CFFBBFE689B2AFD39FB97" box="[434,676,1039,1064]" italics="true" pageId="1" pageNumber="34">Wasmannellus tristis</emphasis>
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<bibRefCitation id="EFC64B4AEE3CFFBBFD719B2AFCEDFB96" author="Bernhauer" box="[683,880,1039,1065]" pageId="1" pageNumber="42" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malayischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article" year="1920">Bernhauer, 1920</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D6530EE3CFFBBFEE29B13FCADFAFE" pageId="1" pageNumber="34" type="diagnosis">
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<paragraph id="8BE836BBEE3CFFBBFEE29B13FCADFAFE" blockId="1.[264,1323,1038,1905]" pageId="1" pageNumber="34">
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<emphasis id="B923EAA9EE3CFFBBFEE29B13FE2AFBEF" bold="true" box="[312,439,1078,1104]" pageId="1" pageNumber="34">Diagnosis.</emphasis>
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The members of
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<taxonomicName id="4C574D38EE3CFFBBFD519B12FCAEFBEF" box="[651,819,1079,1104]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3CFFBBFD519B12FCAEFBEF" box="[651,819,1079,1104]" italics="true" pageId="1" pageNumber="34">Wasmannellus</emphasis>
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</taxonomicName>
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differ from habitually similar members of the genus
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<taxonomicName id="4C574D38EE3CFFBBFE509B7AFD7EFBC6" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[394,739,1119,1145]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3CFFBBFE509B7AFD8AFBC7" box="[394,535,1119,1144]" italics="true" pageId="1" pageNumber="34">Platydracus</emphasis>
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<bibRefCitation id="EFC64B4AEE3CFFBBFDFF9B7AFD7EFBC6" author="Thomson" box="[549,739,1119,1145]" pageId="1" pageNumber="43" refString="Thomson, C. G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson, 1858</bibRefCitation>
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</taxonomicName>
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by the combination of the following character states: puncture bearing postocular seta on the head situated closer to posterior margin of head than to posterior margin of eye; hind tibia lacking spines on dorsolateral face; mandibles each with one simple tooth. The character state of the absence of spines on dorsolateral face of hind tibia will distinguish
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<taxonomicName id="4C574D38EE3CFFBBFCFA9BDAFC54FAA7" box="[800,969,1279,1304]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3CFFBBFCFA9BDAFC54FAA7" box="[800,969,1279,1304]" italics="true" pageId="1" pageNumber="34">Wasmannellus</emphasis>
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</taxonomicName>
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from the members of all other genera that may appear similar in their habitus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D6530EE3CFFB8FEE29A6BFDB4F969" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="description">
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<paragraph id="8BE836BBEE3CFFBBFEE29A6BFAB6F8CE" blockId="1.[264,1323,1038,1905]" pageId="1" pageNumber="34">
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<emphasis id="B923EAA9EE3CFFBBFEE29A6BFE52FAD7" bold="true" box="[312,463,1358,1384]" pageId="1" pageNumber="34">Description.</emphasis>
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Mandibles each with one tooth, delimited basally by a narrow, subbasal indentation. Mandibular prostheca with supporting structure long, lanceolate, with ciliae along medial margin short but becoming longer toward apex. Gular sutures approximate, contiguous. Maxillary palpus rather short, with last segment broadly fusiform, narrowly subtruncate apically, asetose, about as long as segment 3. Labial palpus short, with last segment setose, short, markedly widened anteriad and obliquely truncate apically. Puncture bearing postocular seta on the head situated closer to posterior margin of head than to posterior margin of eye. Superior line of pronotal hypomeron moderately, rather gradually deflected ventrad, anterior portion of superior line therefore situated only moderately below basal portion in lateral view; puncture bearing first lateral macroseta on pronotum situated close to superior line, separated from it by about two diameters of puncture; superior and inferior lines joined just behind anterior angles of pronotum and continuing together to anterior margin of pronotum; deflected portion of pronotal disc visible in ventral view at anterior angles of pronotum. Prothoracic epimeron absent.
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</paragraph>
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<paragraph id="8BE836BBEE3FFFB8FED29E39FDD9FDF1" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="35">Prosternum short, with deep, margined anterior depression, with sternacostal ridge almost straight. Mesosternum without ridges on intercoxal projection, intercoxal mesosternal process narrow, barely separating middle coxae. Metasternum short, evenly convex, with anterior projection short, middle coxal cavities margined posteriorly. Posterior basal line on first three visible abdominal tergites subangulately extended posteriad on each lateral portion, base of tergite between extensions somewhat depressed. Dorsolateral face of front tibia with a few spines on apical third, that of middle tibia with several spines, dorsolateral face of hind tibia aspinose.</paragraph>
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<paragraph id="8BE836BBEE3FFFB8FEE29D79FCD9FC81" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="35">
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Female genital segment with each of the two lateral sclerites of tergite 9 large, wide and flat, densely setose at apical margin and with long, stiff setae on apical portion and along lateral margin; tergite 10 pigmented apically, markedly narrowed into narrow, rodlike apical portion (
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<figureCitation id="136C2A3EEE3FFFB8FE349DF1FDAFFD51" box="[494,562,724,750]" captionStart="FIGURES 1 – 6" captionStartId="5.[264,383,1771,1795]" captionTargetBox="[294,1305,328,1744]" captionTargetId="figure@5.[285,1310,326,1744]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Details of Wasmannellus tristis (1 – 2) and Nelmanwaslus ornatus (3 – 6). 1 — right lateral sclerite of tergite 9 and tergite 10 of female genital segment; 2 — first and second gonocoxites of female genital segment; 3 — tergite 10 of male genital segment; 4 — sternite 9 of male genital segment; 5 — aedoeagus, ventral view; 6 — underside of apical portion of paramere of aedoeagus." httpUri="https://zenodo.org/record/173962/files/figure.png" pageId="2" pageNumber="35">Fig. 1</figureCitation>
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); first gonocoxites apparently fused basally, second gonocoxites each with dense, strong stiff setae on basal half of medial margin, gradually changing into regular setae toward apex of gonocoxite (
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<figureCitation id="136C2A3EEE3FFFB8FD379C01FCA8FC81" box="[749,821,804,830]" captionStart="FIGURES 1 – 6" captionStartId="5.[264,383,1771,1795]" captionTargetBox="[294,1305,328,1744]" captionTargetId="figure@5.[285,1310,326,1744]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Details of Wasmannellus tristis (1 – 2) and Nelmanwaslus ornatus (3 – 6). 1 — right lateral sclerite of tergite 9 and tergite 10 of female genital segment; 2 — first and second gonocoxites of female genital segment; 3 — tergite 10 of male genital segment; 4 — sternite 9 of male genital segment; 5 — aedoeagus, ventral view; 6 — underside of apical portion of paramere of aedoeagus." httpUri="https://zenodo.org/record/173962/files/figure.png" pageId="2" pageNumber="35">Fig. 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE836BBEE3FFFB8FEE29C69FDB4F969" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="35">
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<emphasis id="B923EAA9EE3FFFB8FEE29C69FD04FCD9" bold="true" box="[312,665,844,870]" pageId="2" pageNumber="35">Relationship and comments.</emphasis>
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<bibRefCitation id="EFC64B4AEE3FFFB8FD7B9C69FC77FCD9" author="Bernhauer" box="[673,1002,844,870]" pageId="2" pageNumber="42" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malayischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article" year="1920">Bernhauer (1920: 186–187)</bibRefCitation>
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, in the original description, compared the genus to the genera
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<taxonomicName id="4C574D38EE3FFFB8FD679C51FCA9FC32" box="[701,820,884,909]" class="Insecta" family="Staphylinidae" genus="Physetops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FD679C51FCA9FC32" box="[701,820,884,909]" italics="true" pageId="2" pageNumber="35">Physetops</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C574D38EE3FFFB8FC939C51FB28FC31" authority="Bernhauer, 1902" authorityName="Bernhauer" authorityYear="1902" box="[841,1205,884,910]" class="Insecta" family="Staphylinidae" genus="Barygnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FC939C51FC42FC32" box="[841,991,884,909]" italics="true" pageId="2" pageNumber="35">Barygnathus</emphasis>
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<bibRefCitation id="EFC64B4AEE3FFFB8FC369C51FB28FC31" author="Bernhauer" box="[1004,1205,884,910]" pageId="2" pageNumber="42" refString="Bernhauer, M. (1902) Zur Staphyliniden-Fauna von Ceylon. Deutsche Entomologische Zeitschrift, 1902, 17 - 45." type="journal article" year="1902">Bernhauer, 1902</bibRefCitation>
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</taxonomicName>
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and to a certain group of species that at that time were associated with the genus
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<taxonomicName id="4C574D38EE3FFFB8FB799CB9FE23FC61" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FB799CB9FAB7FC0A" box="[1187,1322,924,949]" italics="true" pageId="2" pageNumber="35">Staphylinus</emphasis>
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<bibRefCitation id="EFC64B4AEE3FFFB8FED29CE1FE23FC61" author="Linnaeus" box="[264,446,964,990]" pageId="2" pageNumber="43" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus I. Laurentii Salvii, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
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</taxonomicName>
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and discussed some distinguishing characters between these genera. As it turns out, the genus
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<taxonomicName id="4C574D38EE3FFFB8FE239CC9FD3CFBBA" box="[505,673,1004,1029]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FE239CC9FD3CFBBA" box="[505,673,1004,1029]" italics="true" pageId="2" pageNumber="35">Wasmannellus</emphasis>
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</taxonomicName>
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is not most closely related to any of these genera. The character state of the superior line of pronotal hypomeron being rather gradually deflected ventrad in front of anterior angles of pronotum (identical to that found in most species of the genus
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<taxonomicName id="4C574D38EE3FFFB8FE589B41FD8DFBC2" box="[386,528,1124,1149]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FE589B41FD8DFBC2" box="[386,528,1124,1149]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
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</taxonomicName>
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), combined with the character state of the position of the puncture bearing postocular seta on the head (situated closer to posterior margin of head than to posterior margin of eye) indicates that these two genera form a clade.
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<taxonomicName id="4C574D38EE3FFFB8FBE69B91FB79FB72" box="[1084,1252,1204,1229]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FBE69B91FB79FB72" box="[1084,1252,1204,1229]" italics="true" pageId="2" pageNumber="35">Wasmannellus</emphasis>
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</taxonomicName>
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is the only other Palaearctic genus, in addition to
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<taxonomicName id="4C574D38EE3FFFB8FCD29BF9FC08FB4A" box="[776,917,1244,1269]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B923EAA9EE3FFFB8FCD29BF9FC08FB4A" box="[776,917,1244,1269]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
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</taxonomicName>
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, with this position of the puncture bearing the postocular seta; however
|
||
<taxonomicName id="4C574D38EE3FFFB8FD139A21FCECFAA2" box="[713,881,1284,1309]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FD139A21FCECFAA2" box="[713,881,1284,1309]" italics="true" pageId="2" pageNumber="35">Wasmannellus</emphasis>
|
||
</taxonomicName>
|
||
does not share with
|
||
<taxonomicName id="4C574D38EE3FFFB8FBB59A21FB61FAA2" box="[1135,1276,1284,1309]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FBB59A21FB61FAA2" box="[1135,1276,1284,1309]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
|
||
</taxonomicName>
|
||
the character state of mandibular teeth appearing in two planes, which is believed to be a synapomorphy for all species of the genus
|
||
<taxonomicName id="4C574D38EE3FFFB8FCDE9A71FC0CFAD2" box="[772,913,1364,1389]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FCDE9A71FC0CFAD2" box="[772,913,1364,1389]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
|
||
</taxonomicName>
|
||
. The presence of a single tooth on each mandible in
|
||
<taxonomicName id="4C574D38EE3FFFB8FE059A59FD49FA2A" box="[479,724,1404,1429]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="tristis">
|
||
<emphasis id="B923EAA9EE3FFFB8FE059A59FD49FA2A" box="[479,724,1404,1429]" italics="true" pageId="2" pageNumber="35">Wasmannellus tristis</emphasis>
|
||
</taxonomicName>
|
||
indicates that this species is not a member of the genus
|
||
<taxonomicName id="4C574D38EE3FFFB8FE8D9A81FE79FA02" box="[343,484,1444,1469]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FE8D9A81FE79FA02" box="[343,484,1444,1469]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
|
||
</taxonomicName>
|
||
. In addition, unlike in most species of
|
||
<taxonomicName id="4C574D38EE3FFFB8FC189A81FBD2FA02" box="[962,1103,1444,1469]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FC189A81FBD2FA02" box="[962,1103,1444,1469]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
|
||
</taxonomicName>
|
||
, in which the last segment of labial palpus is moderately elongate and more or less fusiform, the last segment of labial palpus in
|
||
<taxonomicName id="4C574D38EE3FFFB8FD889AD1FCDBF9B2" box="[594,838,1524,1549]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="tristis">
|
||
<emphasis id="B923EAA9EE3FFFB8FD889AD1FCDBF9B2" box="[594,838,1524,1549]" italics="true" pageId="2" pageNumber="35">Wasmannellus tristis</emphasis>
|
||
</taxonomicName>
|
||
is short, widened anteriad and obliquely truncate apically. Unlike in all species of
|
||
<taxonomicName id="4C574D38EE3FFFB8FD349939FCE6F98A" box="[750,891,1564,1589]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FD349939FCE6F98A" box="[750,891,1564,1589]" italics="true" pageId="2" pageNumber="35">Platydracus</emphasis>
|
||
</taxonomicName>
|
||
, in which the hind tibiae bear spines on dorsolateral face, the hind tibiae of
|
||
<taxonomicName id="4C574D38EE3FFFB8FD3D9961FC40F9E2" box="[743,989,1604,1629]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="tristis">
|
||
<emphasis id="B923EAA9EE3FFFB8FD3D9961FC40F9E2" box="[743,989,1604,1629]" italics="true" pageId="2" pageNumber="35">Wasmannellus tristis</emphasis>
|
||
</taxonomicName>
|
||
lack spines on dorsolateral face. Unfortunately, the only known specimen of
|
||
<taxonomicName id="4C574D38EE3FFFB8FC8C9949FBD7F93A" box="[854,1098,1644,1669]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="tristis">
|
||
<emphasis id="B923EAA9EE3FFFB8FC8C9949FBD7F93A" box="[854,1098,1644,1669]" italics="true" pageId="2" pageNumber="35">Wasmannellus tristis</emphasis>
|
||
</taxonomicName>
|
||
is a female, so the possible differences in the development of the male genital segment and of the aedoeagus remain to be discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D6530EE3FFFB9FEE299C1FAB1FD21" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35" type="reference_group">
|
||
<paragraph id="8BE836BBEE3FFFB9FEE299C1FAB6FE11" blockId="2.[264,1324,284,1830]" lastBlockId="3.[264,1325,284,950]" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
|
||
<taxonomicName id="4C574D38EE3FFFB8FEE299C1FE7DF942" ID-CoL="8XQPK" box="[312,480,1764,1789]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3FFFB8FEE299C1FE7DF942" box="[312,480,1764,1789]" italics="true" pageId="2" pageNumber="35">Wasmannellus</emphasis>
|
||
</taxonomicName>
|
||
will key out in the key to the genera associated with
|
||
<emphasis id="B923EAA9EE3FFFB8FB8399C1FEA8F89A" italics="true" pageId="2" pageNumber="35">
|
||
<taxonomicName id="4C574D38EE3FFFB8FB8399C1FAB7F942" box="[1113,1322,1764,1789]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="sensu">
|
||
Staphylinus
|
||
<taxonomicNameLabel id="A21057D2EE3FFFB8FB3399C1FAB7F942" box="[1257,1322,1764,1789]" pageId="2" pageNumber="35" sensu="lato">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
lato
|
||
</emphasis>
|
||
in
|
||
<bibRefCitation id="EFC64B4AEE3FFFB8FE819829FD10F899" author="Smetana" box="[347,653,1804,1830]" pageId="2" pageNumber="43" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 88 pp." type="book" year="2000">Smetana & Davies (2000)</bibRefCitation>
|
||
to the couplet 3 which should be modified as follows: 3a. Puncture bearing postocular seta on head situated moderately to distinctly closer to posterior margin of head than to posterior margin of eye (Fig.
|
||
<quantity id="4CAF9B5EEE3EFFB9FB9D9E61FB0BFEE1" box="[1095,1174,324,350]" metricMagnitude="0" metricUnit="m" metricValue="2.5907999999999998" pageId="3" pageNumber="36" unit="in" value="102.0">102 in</quantity>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FB799E61FE4FFE39" author="Smetana" pageId="3" pageNumber="43" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 88 pp." type="book" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
). Head at least slightly dilated posteriad, very slightly so in
|
||
<taxonomicName id="4C574D38EE3EFFB9FB479E49FD2BFE11" ID-CoL="6VQ2C" authority="Motschulsky, 1862" authorityName="Motschulsky" authorityYear="1862" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="brevicornis">
|
||
<emphasis id="B923EAA9EE3EFFB9FB479E49FE20FE12" italics="true" pageId="3" pageNumber="36">Platydracus brevicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FE169EB1FD33FE11" author="Motschulsky" box="[460,686,404,430]" pageId="3" pageNumber="43" refString="Motschulsky, V. (1862) Entomologie speciale. Insectes du Japon. Etudes Entomologiques, 9 [1860], 39 - 41." type="journal article" year="1862">Motschulsky, 1862</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C574D38EE3EFFB9FD359EB1FB98FE11" ID-CoL="4JQT4" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[751,1029,404,430]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="juang">
|
||
<emphasis id="B923EAA9EE3EFFB9FD359EB1FD63FE12" box="[751,766,404,429]" italics="true" pageId="3" pageNumber="36">P</emphasis>
|
||
.
|
||
<emphasis id="B923EAA9EE3EFFB9FCD69EB1FCD2FE12" box="[780,847,404,429]" italics="true" pageId="3" pageNumber="36">juang</emphasis>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FC8C9EB1FB98FE11" author="Smetana" box="[854,1029,404,430]" pageId="3" pageNumber="43" refString="Smetana, A. (2005) Contributions to the knowledge of the " Staphylinus-complex " (Coleoptera, Staphylinidae, Staphylinini) of China. Part 10. A new species of the genus Platydracus Thomson, 1858. Zootaxa, 1048, 21 - 25." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
................................... 3b
|
||
</paragraph>
|
||
<paragraph id="8BE836BBEE3EFFB9FED29E99FAB1FD21" blockId="3.[264,1325,284,950]" pageId="3" pageNumber="36">
|
||
Puncture bearing postocular seta on head situated distinctly, or at least appreciably, closer to posterior margin of eye than to posterior margin of head (Figs.
|
||
<quantity id="4CAF9B5EEE3EFFB9FB789EC1FAB6FE41" box="[1186,1323,484,510]" metricMagnitude="0" metricUnit="m" metricValue="2.6188416" pageId="3" pageNumber="36" unit="in" value="103.104">103, 104 in</quantity>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FEE29D29FDC3FD99" author="Smetana" box="[312,606,524,550]" pageId="3" pageNumber="43" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 88 pp." type="book" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
). Head not appreciably dilated posteriad (slightly so in some
|
||
<taxonomicName id="4C574D38EE3EFFB9FEE29D11FD3FFDF1" ID-CoL="8VRX9" authority="Ganglbauer, 1895" authorityName="Ganglbauer" authorityYear="1895" box="[312,674,564,590]" class="Insecta" family="Staphylinidae" genus="Ontholestes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3EFFB9FEE29D11FE5FFDF2" box="[312,450,564,589]" italics="true" pageId="3" pageNumber="36">Ontholestes</emphasis>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FE169D11FD3FFDF1" author="Ganglbauer" box="[460,674,564,590]" pageId="3" pageNumber="43" refString="Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. Die Kafer der osterreich-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebiets. Band 2. Familienreihe Staphylinoidea. 1. Theil. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, vi + 880 pp." type="book" year="1895">Ganglbauer, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
), with posterior corners variably, obtusely rounded, or if dilated posteriad, then posterolateral portions of head each extended posteriad as rounded lobe (Fig.
|
||
<quantity id="4CAF9B5EEE3EFFB9FDC29DA1FDFFFD21" box="[536,610,644,670]" metricMagnitude="0" metricUnit="m" metricValue="2.667" pageId="3" pageNumber="36" unit="in" value="105.0">105 in</quantity>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FDB39DA1FC15FD21" author="Smetana" box="[617,904,644,670]" pageId="3" pageNumber="43" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 88 pp." type="book" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
)..................................................... 4
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D6530EE3EFFB9FED29D89FAB6FC09" pageId="3" pageNumber="36" type="description">
|
||
<paragraph id="8BE836BBEE3EFFB9FED29D89FAB7FC81" blockId="3.[264,1325,284,950]" pageId="3" pageNumber="36">
|
||
3b. Last segment of labial palpus asetose, moderately elongate, more or less fusiform (Fig.
|
||
<quantity id="4CAF9B5EEE3EFFB9FEE29DF1FE1FFD51" box="[312,386,724,750]" metricMagnitude="0" metricUnit="m" metricValue="3.429" pageId="3" pageNumber="36" unit="in" value="135.0">135 in</quantity>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FE539DF1FD31FD51" author="Smetana" box="[393,684,724,750]" pageId="3" pageNumber="43" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 88 pp." type="book" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
). Dorsolateral face of hind tibia with spines. Mandibular teeth, at least two on each mandible, appearing in two planes, dorsal and ventral (
|
||
<figureCitation id="136C2A3EEE3EFFB9FE9B9C01FE09FC81" box="[321,404,804,830]" captionStart="FIGURES 1 – 6" captionStartId="5.[264,383,1771,1795]" captionTargetBox="[294,1305,328,1744]" captionTargetId="figure@5.[285,1310,326,1744]" captionTargetPageId="5" captionText="FIGURES 1 – 6. Details of Wasmannellus tristis (1 – 2) and Nelmanwaslus ornatus (3 – 6). 1 — right lateral sclerite of tergite 9 and tergite 10 of female genital segment; 2 — first and second gonocoxites of female genital segment; 3 — tergite 10 of male genital segment; 4 — sternite 9 of male genital segment; 5 — aedoeagus, ventral view; 6 — underside of apical portion of paramere of aedoeagus." httpUri="https://zenodo.org/record/173962/files/figure.png" pageId="3" pageNumber="36">
|
||
Figs.
|
||
<quantity id="4CAF9B5EEE3EFFB9FE599C01FE09FC81" box="[387,404,804,830]" metricMagnitude="-1" metricUnit="m" metricValue="1.7018" pageId="3" pageNumber="36" unit="in" value="6.7">6</quantity>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="136C2A3EEE3EFFB9FE7A9C01FE32FC81" box="[416,431,804,830]" captionStart="FIGURES 7 – 8" captionStartId="6.[264,382,1386,1410]" captionTargetBox="[424,1164,323,1354]" captionTargetId="figure@6.[400,1186,308,1362]" captionTargetPageId="6" captionText="FIGURES 7 – 8. Wasmannellus tristis. 7 — habitus of the holotype; 8 — labels attached to the holotype." httpUri="https://zenodo.org/record/173963/files/figure.png" pageId="3" pageNumber="36">7</figureCitation>
|
||
in
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FE0F9C01FD68FC81" author="Smetana" box="[469,757,804,830]" pageId="3" pageNumber="43" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 88 pp." type="book" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
)
|
||
<emphasis id="B923EAA9EE3EFFB9FCDB9C01FC7DFC82" box="[769,992,804,829]" italics="true" pageId="3" pageNumber="36">..............................</emphasis>
|
||
<taxonomicName id="4C574D38EE3EFFB9FC3A9C01FAB7FC81" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[992,1322,804,830]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3EFFB9FC3A9C01FBF1FC82" box="[992,1132,804,829]" italics="true" pageId="3" pageNumber="36">Platydracus</emphasis>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FBA99C01FAB7FC81" author="Thomson" box="[1139,1322,804,830]" pageId="3" pageNumber="43" refString="Thomson, C. G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BE836BBEE3EFFB9FED29C69FAB6FC09" blockId="3.[264,1325,284,950]" pageId="3" pageNumber="36">
|
||
Last segment of labial palpus setose, short, markedly widened anteriad and obliquely truncate apically. Dorsolateral face of hind tibia lacking spines. Mandibles each with one tooth
|
||
<emphasis id="B923EAA9EE3EFFB9FE729CB9FC26FC0A" box="[424,955,924,949]" italics="true" pageId="3" pageNumber="36">........................................................................</emphasis>
|
||
<taxonomicName id="4C574D38EE3EFFB9FC669CB9FAB6FC09" authority="Bernhauer, 1920" authorityName="Bernhauer" authorityYear="1920" box="[956,1323,924,950]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923EAA9EE3EFFB9FC669CB9FBFCFC0A" box="[956,1121,924,949]" italics="true" pageId="3" pageNumber="36">Wasmannellus</emphasis>
|
||
<bibRefCitation id="EFC64B4AEE3EFFB9FBB29CB9FAB6FC09" author="Bernhauer" box="[1128,1323,924,950]" pageId="3" pageNumber="42" refString="Bernhauer, M. (1920) Neue Staphyliniden des indo-malayischen Gebietes. Archiv fur Naturgeschichte (A), 84, 177 - 188." type="journal article" year="1920">Bernhauer, 1920</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |