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<document id="5A7E3047F5DA6B4D5FBEF8BFC7048632" ID-DOI="10.11646/zootaxa.4039.3.4" ID-GBIF-Dataset="01d8c41d-28d4-4d9b-99e7-c525e3d1bbbb" ID-ISSN="1175-5326" ID-Zenodo-Dep="243160" ID-ZooBank="3924A813-5BCE-4DED-8257-1664BC133DF9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461332093721" checkinUser="plazi" docAuthor="Gardzińska, Joanna" docDate="2015" docId="03FE8792FFE28C69FF1EAAEF0FD9FB54" docLanguage="en" docName="zt04039p455.pdf" docOrigin="Zootaxa 4039 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Udvardya Proszynski 1992" docType="treatment" docVersion="7" lastPageNumber="448" masterDocId="FFC7FFEAFFE18C6AFF89AA770D22FFEC" masterDocTitle="Revision of Tarodes Pocock, 1899 and Udvardya Prószyński, 1992 (Araneae: Salticidae), with descriptions of two new species of Udvardya from New Guinea" masterLastPageNumber="455" masterPageNumber="445" pageNumber="448" updateTime="1698618589457" updateUser="plazi">
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<mods:title id="4881C14A3B043D453D759AB80E3E6689">Revision of Tarodes Pocock, 1899 and Udvardya Prószyński, 1992 (Araneae: Salticidae), with descriptions of two new species of Udvardya from New Guinea</mods:title>
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<treatment id="03FE8792FFE28C69FF1EAAEF0FD9FB54" ID-DOI="http://doi.org/10.5281/zenodo.6098441" ID-GBIF-Taxon="119644152" ID-Zenodo-Dep="6098441" LSID="urn:lsid:plazi:treatment:03FE8792FFE28C69FF1EAAEF0FD9FB54" httpUri="http://treatment.plazi.org/id/03FE8792FFE28C69FF1EAAEF0FD9FB54" lastPageNumber="448" pageId="3" pageNumber="448">
<subSubSection id="C34D650FFFE28C69FF1EAAEF0C24FF3F" pageId="3" pageNumber="448" type="nomenclature">
<paragraph id="8BE83684FFE28C69FF1EAAEF0F1FFF5E" blockId="3.[151,573,152,211]" box="[151,573,152,178]" pageId="3" pageNumber="448">
<heading id="D0A081E8FFE28C69FF1EAAEF0F1FFF5E" bold="true" box="[151,573,152,178]" fontSize="11" level="1" pageId="3" pageNumber="448" reason="1">
<emphasis id="B923EA96FFE28C69FF1EAAEF0F1FFF5E" bold="true" box="[151,573,152,178]" pageId="3" pageNumber="448">
Genus
<taxonomicName id="4C574D07FFE28C69FF67AAEF0F1FFF5E" ID-CoL="84HR" authority="Proszynski, 1992" authorityName="Proszynski" authorityYear="1992" box="[238,573,152,178]" class="Arachnida" family="Salticidae" genus="Udvardya" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B923EA96FFE28C69FF67AAEF0C41FF5E" bold="true" box="[238,355,152,178]" italics="true" pageId="3" pageNumber="448">Udvardya</emphasis>
<bibRefCitation id="EFC64B75FFE28C69FEE3AAEF0F1FFF5E" author="Proszynski" box="[362,573,152,178]" pageId="3" pageNumber="448" refString="Proszynski, J. (1992) Salticidae (Araneae) of the Old World and Pacific Islands in several US collections. Annales Zoologici, 44, 87 - 163." type="journal article" year="1992">Prószyński, 1992</bibRefCitation>
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<figureCitation id="136C2A01FFE28C69FF1EAACD0C24FF3F" box="[151,262,186,211]" captionStart-0="FIGURES 8 – 17" captionStart-1="FIGURES 18 – 27" captionStart-2="FIGURES 28 – 36" captionStart-3="FIGURES 37 – 45" captionStart-4="FIGURES 46 – 51" captionStart-5="FIGURE 52" captionStartId-0="4.[151,264,1901,1923]" captionStartId-1="6.[151,264,1919,1941]" captionStartId-2="7.[151,264,1911,1933]" captionStartId-3="8.[151,264,1855,1877]" captionStartId-4="9.[151,264,1166,1188]" captionStartId-5="9.[151,250,2001,2023]" captionTargetBox-0="[151,1436,193,1879]" captionTargetBox-1="[202,1385,193,1886]" captionTargetBox-2="[215,1376,199,1881]" captionTargetBox-3="[183,1436,198,1829]" captionTargetBox-4="[151,1436,193,1144]" captionTargetBox-5="[151,1436,1268,1979]" captionTargetId-0="figure@4.[151,1436,193,1880]" captionTargetId-1="figure@6.[202,1385,193,1887]" captionTargetId-2="figure@7.[208,1377,193,1882]" captionTargetId-3="figure@8.[151,1436,193,1833]" captionTargetId-4="figure@9.[151,1436,193,1145]" captionTargetId-5="figure@9.[151,1436,1268,1979]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="9" captionText-0="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." captionText-1="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." captionText-2="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." captionText-3="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." captionText-4="FIGURES 46 – 51. Udvardya, comparison of female abdominal color patterns (dorsal and lateral views). 46 – 47 U. elegans; 48 – 49 U. fortis; 50 – 51 U. bellatrix." captionText-5="FIGURE 52. Distributional records of species of Tarodes (squares) and Udvardya (circles)." httpUri-0="https://zenodo.org/record/243162/files/figure.png" httpUri-1="https://zenodo.org/record/243163/files/figure.png" httpUri-2="https://zenodo.org/record/243164/files/figure.png" httpUri-3="https://zenodo.org/record/243165/files/figure.png" httpUri-4="https://zenodo.org/record/243166/files/figure.png" httpUri-5="https://zenodo.org/record/243167/files/figure.png" pageId="3" pageNumber="448">Figs 8‒52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34D650FFFE28C69FF1EAB750CCBFEF4" box="[151,489,258,280]" pageId="3" pageNumber="448" type="reference_group">
<paragraph id="8BE83684FFE28C69FF1EAB750CCBFEF4" blockId="3.[151,657,258,311]" box="[151,489,258,280]" pageId="3" pageNumber="448">
<treatmentCitationGroup id="AB4711AAFFE28C69FF1EAB750CCBFEF4" box="[151,489,258,280]" pageId="3" pageNumber="448">
<taxonomicName id="4C574D07FFE28C69FF1EAB750CC7FEF4" ID-CoL="84HR" authority="Proszynski, 1992: 113" authorityName="Proszynski" authorityPageNumber="113" authorityYear="1992" box="[151,485,258,280]" class="Arachnida" family="Salticidae" genus="Udvardya" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B923EA96FFE28C69FF1EAB750DD9FEF4" box="[151,251,258,280]" italics="true" pageId="3" pageNumber="448">Udvardya</emphasis>
<treatmentCitation id="0AF61095FFE28C69FE88AB750CC7FEF4" author="Proszynski" box="[257,485,258,280]" page="113" pageId="3" pageNumber="448" year="1992">
<bibRefCitation id="EFC64B75FFE28C69FE88AB750C91FEF4" author="Proszynski" box="[257,435,258,280]" pageId="3" pageNumber="448" refString="Proszynski, J. (1992) Salticidae (Araneae) of the Old World and Pacific Islands in several US collections. Annales Zoologici, 44, 87 - 163." type="journal article" year="1992">Prószyński, 1992</bibRefCitation>
: 113
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.
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</paragraph>
</subSubSection>
<subSubSection id="C34D650FFFE28C69FF1EAB560FB3FEDB" box="[151,657,289,311]" pageId="3" pageNumber="448" type="materials_examined">
<paragraph id="8BE83684FFE28C69FF1EAB560FB3FEDB" blockId="3.[151,657,258,311]" box="[151,657,289,311]" pageId="3" pageNumber="448">
<emphasis id="B923EA96FFE28C69FF1EAB560C07FEDB" bold="true" box="[151,293,289,311]" pageId="3" pageNumber="448">
<typeStatus id="54EC8826FFE28C69FF1EAB560DEEFEDB" box="[151,204,289,311]" pageId="3" pageNumber="448">Type</typeStatus>
species:
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<taxonomicName id="4C574D07FFE28C69FEA2AB560FAFFEDB" authority="Szombathy, 1915" authorityName="Szombathy" authorityYear="1915" box="[299,653,289,311]" class="Arachnida" family="Salticidae" genus="Silerella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B923EA96FFE28C69FEA2AB560CF1FEDB" box="[299,467,289,311]" italics="true" pageId="3" pageNumber="448">Silerella elegans</emphasis>
<bibRefCitation id="EFC64B75FFE28C69FE53AB560FAFFEDB" author="Szombathy" box="[474,653,289,311]" pageId="3" pageNumber="448" refString="Szombathy, K. (1915) Attides nouveaux appartenant aux collections du Musee National Hongrois. Annales Musei Nationalis Hungarici, 13, 468 - 490." type="journal article" year="1915">Szombathy, 1915</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="C34D650FFFE28C69FF1EAB1308BEFE04" pageId="3" pageNumber="448" type="diagnosis">
<paragraph id="8BE83684FFE28C69FF1EAB1308BEFE04" blockId="3.[151,1437,356,1208]" pageId="3" pageNumber="448">
<emphasis id="B923EA96FFE28C69FF1EAB130C28FE91" bold="true" box="[151,266,356,381]" pageId="3" pageNumber="448">Diagnosis</emphasis>
. Compared with the closest
<emphasis id="B923EA96FFE28C69FDC0AB130F87FE90" box="[585,677,356,381]" italics="true" pageId="3" pageNumber="448">
<taxonomicName id="4C574D07FFE28C69FDC0AB130FBCFE91" box="[585,670,356,381]" class="Arachnida" family="Salticidae" genus="Tarodes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="448" phylum="Arthropoda" rank="genus">Tarodes</taxonomicName>
,
</emphasis>
males of
<taxonomicName id="4C574D07FFE28C69FC9EAB130EA4FE91" box="[791,902,356,381]" class="Arachnida" family="Salticidae" genus="Udvardya" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B923EA96FFE28C69FC9EAB130EA4FE91" box="[791,902,356,381]" italics="true" pageId="3" pageNumber="448">Udvardya</emphasis>
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have mastidia projecting forward from anterior surface of chelicerae; cephalothorax of both sexes with more or less distinct cheek swellings; epigyne with copulatory openings oriented anteriorly, copulatory ducts with longitudinal chambers, accompanied by accessory glands; typical secondary spermathecae lacking; embolus relatively longer; retrolateral tibial apophysis spade-like.
</paragraph>
</subSubSection>
<subSubSection id="C34D650FFFE28C69FF4EAB830CD5FB78" pageId="3" pageNumber="448" type="description">
<paragraph id="8BE83684FFE28C69FF4EAB830CD5FB78" blockId="3.[151,1437,356,1208]" pageId="3" pageNumber="448">
<emphasis id="B923EA96FFE28C69FF4EAB830C7BFDE1" bold="true" box="[199,345,500,525]" pageId="3" pageNumber="448">Description.</emphasis>
Cephalothorax of both sexes of trapezoid shape, with more or less distinct cheek swellings (CS) and posterior slope more gentle, relatively low, with sides slightly curved; dorsal surface with granulation, especially on eye field (
<figureCitation id="136C2A01FFE28C69FE2FA84B0CCDFDB8" box="[422,495,572,597]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 8</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FE74A84B0F64FDB8" box="[509,582,572,596]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">18, 21</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FDDAA84B0FBFFDB8" box="[595,669,572,596]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">28, 31</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FD23A84B0FD7FDB8" box="[682,757,572,596]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">37, 39</figureCitation>
). Anterior eye row wider than posterior one; eye field takes more than 50% of CL length. Posterior eyes on relatively small protuberances. Clypeus very narrow (about 10% of AME diameter), densely covered with pallid hairs, forming transverse bands on cheeks (
<figureCitation id="136C2A01FFE28C69FB1BA8F309C8FD70" box="[1170,1258,644,669]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 11</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FB70A8F30860FD70" box="[1273,1346,644,668]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">21‒22</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FAC6A8F308BAFD70" box="[1359,1432,644,668]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">31‒32</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FF1EA8DF0DC0FD2C" box="[151,226,680,704]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">39‒41</figureCitation>
). Chelicerae with fissident pattern (
<figureCitation id="136C2A01FFE28C69FDFBA8D00FEBFD2C" box="[626,713,679,704]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 13</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FD5FA8DF0FD6FD2C" box="[726,756,680,704]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">23</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FC88A8DF0E3DFD2C" box="[769,799,680,704]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">33</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FCA2A8DF0E6BFD2C" box="[811,841,680,704]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">42</figureCitation>
) and distinct lateral edges and transverse furrows on anterior surface (
<figureCitation id="136C2A01FFE28C69FEDEA8BB0C8DFD08" box="[343,431,716,741]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 12</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FE35A8BB0CF8FD08" box="[444,474,716,740]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">22</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FE6EA8BB0F27FD08" box="[487,517,716,740]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">32</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FD9BA8BB0F13FD08" box="[530,561,716,740]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">41</figureCitation>
); males with mastidia (Ma), as in
<figureCitation id="136C2A01FFE28C69FC3EA8BB091AFD08" box="[951,1080,716,741]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">Figs 21‒22</figureCitation>
. Fangs relatively short. Endites divergent, without modifications; labium longer than wide. Sternum narrow, with anterior margin as wide as labium (
<figureCitation id="136C2A01FFE28C69FF72A9630C6EFCC0" box="[251,332,788,813]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 9</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FEE9A9630C5CFCC0" box="[352,382,788,812]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">19</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FE1BA9630C92FCC0" box="[402,432,788,812]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">29</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FE4DA9630CC1FCC0" box="[452,483,788,812]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">38</figureCitation>
). Pedicel moderately long, visible in dorsal view. Abdomen elongate-ovoid or cylindrical, not constricted, with anterior hair tuft and pattern of transverse and longitudinal pallid stripes and patches on dark background (
<figureCitation id="136C2A01FFE28C69FE6EA92B0F7EFC98" box="[487,604,860,885]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 8‒10</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FDE0A92B0F90FC98" box="[617,690,860,884]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">18‒20</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FD36A92B0E2AFC98" box="[703,776,860,884]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">28‒30</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FC9CA92B0E7CFC98" box="[789,862,860,884]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">37‒39</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FCE3A92B0E97FC99" box="[874,949,860,885]" captionStart="FIGURES 46 – 51" captionStartId="9.[151,264,1166,1188]" captionTargetBox="[151,1436,193,1144]" captionTargetId="figure@9.[151,1436,193,1145]" captionTargetPageId="9" captionText="FIGURES 46 – 51. Udvardya, comparison of female abdominal color patterns (dorsal and lateral views). 46 – 47 U. elegans; 48 – 49 U. fortis; 50 – 51 U. bellatrix." httpUri="https://zenodo.org/record/243166/files/figure.png" pageId="3" pageNumber="448">46‒51</figureCitation>
); males with dorsal and ventral scuta (
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) (
<figureCitation id="136C2A01FFE28C69FF16A9080C04FC74" box="[159,294,895,920]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">Figs 18‒20</figureCitation>
). First legs distinctly more robust and longer than others, femora flattened laterally, with distinct retroventral edges.
<collectingCountry id="F3407614FFE28C69FEFBA9D30C8EFC51" box="[370,428,932,957]" name="Malta" pageId="3" pageNumber="448">Male</collectingCountry>
trochanters I somewhat longer than coxae, in females relatively shorter. Leg formula: I- IV-II-III. Tibiae I armed with 7‒8 proventral and 6‒7 retroventral spines; metatarsi I with 3 pairs of ventral spines. Ventral surface of tibiae I and patellae I fringed with dense setae. Epigyne with copulatory ducts (CD) sclerotized and relatively short, running obliquely from the copulatory openings (CO) towards the midline; spermathecae (S) from almost oval through baggy to pear-shaped; accessory glands (AG) located on CD and walls of S (
<figureCitation id="136C2A01FFE28C69FA9FAE43084FFBA0" box="[1302,1389,1076,1101]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,264,1901,1923]" captionTargetBox="[151,1436,193,1879]" captionTargetId="figure@4.[151,1436,193,1880]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Udvardya elegans (Szombathy, 1915), female holotype. 8 general appearance, dorsal view; 9 same, ventral view; 10 same, lateral view; 11 cephalothorax, frontal view; 12 chelicerae, frontal view; 13 same, ventral view; 14 right leg I, prolateral view; 15 epigyne, ventral view; 16 same, cleared, ventral view; 17 same, internal structures. Scale bars: 8, 10, 14: 1 mm; 15 – 17: 0.2 mm." httpUri="https://zenodo.org/record/243162/files/figure.png" pageId="3" pageNumber="448">Figs 17</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FAF0AE4308B5FBA1" box="[1401,1431,1076,1101]" captionStart="FIGURES 28 – 36" captionStartId="7.[151,264,1911,1933]" captionTargetBox="[215,1376,199,1881]" captionTargetId="figure@7.[208,1377,193,1882]" captionTargetPageId="7" captionText="FIGURES 28 – 36. Udvardya bellatrix sp. nov., female paratype. 28 general appearance, dorsal view; 29 same, ventral view; 30 same, dorso-lateral view; 31 cephalothorax, lateral view; 32 same, frontal view; 33 chelicerae, endites and labium, ventral view; 34 epigyne, ventral view; 35 same, cleared, ventral view; 36 same, internal structures. Scale bars: 28, 30 – 31: 1 mm; 34 – 36: 0.2 mm." httpUri="https://zenodo.org/record/243164/files/figure.png" pageId="3" pageNumber="448">36</figureCitation>
,
<figureCitation id="136C2A01FFE28C69FF1EAE200D95FB9C" box="[151,183,1111,1136]" captionStart="FIGURES 37 – 45" captionStartId="8.[151,264,1855,1877]" captionTargetBox="[183,1436,198,1829]" captionTargetId="figure@8.[151,1436,193,1833]" captionTargetPageId="8" captionText="FIGURES 37 – 45. Udvardya fortis sp. nov., female holotype. 37 general appearance, dorsal view; 38 same, ventral view; 39 same, dorso-lateral view; 40 cephalothorax, lateral view; 41 same, frontal view; 42 cheliceral dentition, retrolateral view; 43 epigyne, ventral view; 44 same, cleared, ventral view; 45 same, internal structures. Scale bars: 37, 39 – 40: 1 mm; 43 – 45: 0.2 mm." httpUri="https://zenodo.org/record/243165/files/figure.png" pageId="3" pageNumber="448">45</figureCitation>
). Palpal organ with embolus (E) moderately long, located apically at bulb; retrolateral tibial apophysis (RTA) broad, not forked (
<figureCitation id="136C2A01FFE28C69FEEEAE0B0CC9FB78" box="[359,491,1148,1173]" captionStart="FIGURES 18 – 27" captionStartId="6.[151,264,1919,1941]" captionTargetBox="[202,1385,193,1886]" captionTargetId="figure@6.[202,1385,193,1887]" captionTargetPageId="6" captionText="FIGURES 18 – 27. Udvardya bellatrix sp. nov., male holotype. 18 general appearance, dorsal view; 19 same, ventral view; 20 abdomen, lateral view; 21 cephalothorax, lateral view; 22 same, fronto-lateral view; 23 chelicera, retrolateral view; 24, 26 palpal organ, ventral view; 25, 27 same, retrolateral view. Scale bars: 18, 21: 1 mm; 24, 26: 0.2 mm." httpUri="https://zenodo.org/record/243163/files/figure.png" pageId="3" pageNumber="448">Figs 24‒27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D650FFFE28C69FF4EAEE80FD9FB54" box="[199,763,1183,1208]" pageId="3" pageNumber="448" type="distribution">
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<emphasis id="B923EA96FFE28C69FF4EAEE80C42FB54" bold="true" box="[199,352,1183,1208]" pageId="3" pageNumber="448">Distribution.</emphasis>
Known from New
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(
<figureCitation id="136C2A01FFE28C69FD13AEE80FCCFB54" box="[666,750,1183,1208]" captionStart="FIGURE 52" captionStartId="9.[151,250,2001,2023]" captionTargetBox="[151,1436,1268,1979]" captionTargetId="figure@9.[151,1436,1268,1979]" captionTargetPageId="9" captionText="FIGURE 52. Distributional records of species of Tarodes (squares) and Udvardya (circles)." httpUri="https://zenodo.org/record/243167/files/figure.png" pageId="3" pageNumber="448">Fig. 52</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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