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<document id="206FBF29FBFEBD02161884A366E353FD" ID-CLB-Dataset="31156" ID-DOI="10.11646/zootaxa.4365.1.2" ID-GBIF-Dataset="e96f0a31-8cfd-4f96-96d1-189ad5392f31" ID-ISSN="1175-5326" ID-Zenodo-Dep="1116510" ID-ZooBank="B6516910-3B14-4E5B-AF4F-38BC8B7DBAF3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513314681040" checkinUser="plazi" docAuthor="Nestor-Arriola, Jorge Ismael &amp; Toledo-Hernández, Víctor Hugo" docDate="2017" docId="03F1E8391B442611C3E89588792D7B58" docLanguage="en" docName="zootaxa.4365.1.2.pdf" docOrigin="Zootaxa 4365 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brachiacantha brevihamata Nestor-Arriola &amp; Toledo-Hernández 2017, sp. nov." docType="treatment" docVersion="7" masterDocId="FFC890411B452612C37F90317B157D50" masterDocTitle="New species of Brachiacantha Dejean, 1837 (Coleoptera: Coccinellidae) from Mexico and Central America" updateTime="1698510066118" updateUser="ExternalLinkService">
<mods:mods id="434AADBFFFF78EE73E68C76C81459902" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C401DD45E9FB7E108BE9DCB561F5D2D8">
<mods:title id="4D2C43B67D1E02108FDF4DD4F71F476B">New species of Brachiacantha Dejean, 1837 (Coleoptera: Coccinellidae) from Mexico and Central America</mods:title>
</mods:titleInfo>
<mods:name id="67BD558469946FBEF55C47BABF42D72B" type="personal">
<mods:role id="19D5361792FCA1BD98518F53AA1A1FA5">
<mods:roleTerm id="9E0CB8200DEAC8F572BA766F3EBC556C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC5C50BDBAF387B511C2003FD6CD3112">Nestor-Arriola, Jorge Ismael</mods:namePart>
</mods:name>
<mods:name id="891ADA7A32AF5E6FF3DA829D1FE43350" type="personal">
<mods:role id="B1A0C7A4EB719D408817B7AD5D6F34AE">
<mods:roleTerm id="099E2D861EA681B58657618B16BFAD32">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF2EE46CF920E484FF035818F60C60F8">Toledo-Hernández, Víctor Hugo</mods:namePart>
</mods:name>
<mods:typeOfResource id="771FD2A30EBCDF20D06F17179E9426BB">text</mods:typeOfResource>
<mods:relatedItem id="961E5ABEDF281D6C2724A92A291B4DA0" type="host">
<mods:titleInfo id="26A4CFD576B723E54D9F1A6CC78ABFDA">
<mods:title id="1EDCC498DB86AF14575373A6C95B5FFA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="44DB045D0470751D21EC9D66DC1446F4">
<mods:date id="92EB077F8C74FEB4BF2E7079EC0D7935">2017</mods:date>
<mods:detail id="C8B6BEC132AC00D798E7A78C7B2F615C" type="pubDate">
<mods:number id="C99BA38D7BE79D1187D15BEE9C7DDF56">2017-12-15</mods:number>
</mods:detail>
<mods:detail id="8378152C78813229A55E35BE047B0311" type="volume">
<mods:number id="5CB12D6BC620F376D7C37E06C7E399F4">4365</mods:number>
</mods:detail>
<mods:detail id="933C61396253514BF6C7CF7A73218EC2" type="issue">
<mods:number id="8A05742EC8555929EF1B38F31E95BBBD">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="BCE9801E216398AF795942CB0490370A">journal volume</mods:classification>
<mods:identifier id="F71414100468BA6DD0D36EBBE78B51BC" type="CLB-Dataset">31156</mods:identifier>
<mods:identifier id="729DB8415EF4082355E077DD480E7A03" type="DOI">10.11646/zootaxa.4365.1.2</mods:identifier>
<mods:identifier id="BB847719A6B63A002E54CEF8033FF165" type="GBIF-Dataset">e96f0a31-8cfd-4f96-96d1-189ad5392f31</mods:identifier>
<mods:identifier id="8B8F652A60007EF3C6408D32741ED008" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="579D84A55AF876228A365DC69AD71E5F" type="Zenodo-Dep">1116510</mods:identifier>
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</mods:mods>
<treatment id="03F1E8391B442611C3E89588792D7B58" ID-DOI="http://doi.org/10.5281/zenodo.6001884" ID-GBIF-Taxon="139328571" ID-Zenodo-Dep="6001884" LSID="urn:lsid:plazi:treatment:03F1E8391B442611C3E89588792D7B58" httpUri="http://treatment.plazi.org/id/03F1E8391B442611C3E89588792D7B58" lastPageId="3" pageId="1">
<subSubSection id="C3420AA41B442613C3E895887A0778A4" pageId="1" type="nomenclature">
<paragraph id="8BE7592F1B442613C3E89588795A7883" blockId="1.[151,591,1465,1525]" box="[151,591,1465,1491]" pageId="1">
<heading id="D0AFEE431B442613C3E89588795A7883" bold="true" box="[151,591,1465,1491]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="B92C853D1B442613C3E89588795A7883" bold="true" box="[151,591,1465,1491]" pageId="1">
<taxonomicName id="4C5822AC1B442613C3E895887AFF7883" authority="Nestor-Arriola &amp; Toledo-Hernández, 2017" authorityName="Nestor-Arriola &amp; Toledo-Hernández" authorityYear="2017" box="[151,490,1465,1491]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="brevihamata" status="sp. nov.">
<emphasis id="B92C853D1B442613C3E895887AFF7883" bold="true" box="[151,490,1465,1491]" italics="true" pageId="1">Brachiacantha brevihamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F38461B442613C28E9588795A7883" box="[497,591,1465,1491]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7592F1B442613C3E895ED7A0778A4" blockId="1.[151,591,1465,1525]" box="[151,274,1500,1525]" pageId="1">
(
<figureCitation id="136345AA1B442613C3E095ED7A1C78A4" box="[159,265,1500,1525]" captionStart="FIGURES 14" captionStartId="2.[151,264,1973,1996]" captionTargetBox="[223,1324,220,1925]" captionTargetId="figure@2.[207,1379,193,1952]" captionTargetPageId="2" captionText="FIGURES 14. Brachiacantha brevihamata (images by Jorge Valdez Carrasco). 1) dorsal habitus; 2) tegmen, concave side vieW;3) tegmen, lateral vieW; 4) sipho." httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="1">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3420AA41B442613C3E896157E957B30" pageId="1" type="diagnosis">
<paragraph id="8BE7592F1B442613C3E896157E957B30" blockId="1.[151,1437,1572,2029]" pageId="1">
<emphasis id="B92C853D1B442613C3E896157A047B6D" bold="true" box="[151,273,1572,1597]" pageId="1">Diagnosis.</emphasis>
Five yellowish orange spots on each elytron. Male with ventrites 2 to 5 emarginate and depressed, forming a longitudinal fossa on abdomen. Basal lobe of male genitalia with a small lateral hook at apeX (
<figureCitation id="136345AA1B442613C65296767E677B30" box="[1325,1394,1607,1632]" captionStart="FIGURES 14" captionStartId="2.[151,264,1973,1996]" captionTargetBox="[223,1324,220,1925]" captionTargetId="figure@2.[207,1379,193,1952]" captionTargetPageId="2" captionText="FIGURES 14. Brachiacantha brevihamata (images by Jorge Valdez Carrasco). 1) dorsal habitus; 2) tegmen, concave side vieW;3) tegmen, lateral vieW; 4) sipho." httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="1">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3420AA41B442611C3B8965D78887E68" lastPageId="3" pageId="1" type="description">
<paragraph id="8BE7592F1B442613C3B8965D7E887ABC" blockId="1.[151,1437,1572,2029]" pageId="1">
<emphasis id="B92C853D1B442613C3B8965D7A4C7BD5" bold="true" box="[199,345,1644,1669]" pageId="1">Description.</emphasis>
<emphasis id="B92C853D1B442613C221965D7ADB7BD5" box="[350,462,1644,1669]" italics="true" pageId="1">
<typeStatus id="54E3E78D1B442613C221965D7AD27BD5" box="[350,455,1644,1669]" pageId="1" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male: Length
<quantity id="4CA0F4CA1B442613C10D965D79DF7BD4" box="[626,714,1644,1668]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" unit="mm" value="3.8">3.8 mm</quantity>
, width
<quantity id="4CA0F4CA1B442613C064965D78667BD4" box="[795,883,1644,1669]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" unit="mm" value="2.5">2.5 mm</quantity>
; body oval, slightly elongated. Dorsal color black eXcept head yellow; pronotum with anterolateral angles and the anterior and lateral borders yellowish orange, a mark of the same color eXtends over the middle line from apeX to half the length of the pronotum; each elytron with five yellowish-orange spots: a small triangular humeral spot, a rounded scutellar spot, a rounded middle-lateral spot, a rounded discal spot and a transversely oval subapical spot (
<figureCitation id="136345AA1B442613C0F596CD78D87A44" box="[906,973,1788,1813]" captionStart="FIGURES 14" captionStartId="2.[151,264,1973,1996]" captionTargetBox="[223,1324,220,1925]" captionTargetId="figure@2.[207,1379,193,1952]" captionTargetPageId="2" captionText="FIGURES 14. Brachiacantha brevihamata (images by Jorge Valdez Carrasco). 1) dorsal habitus; 2) tegmen, concave side vieW;3) tegmen, lateral vieW; 4) sipho." httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="1">Fig. 1</figureCitation>
). Ventral surface with head, prosternum, metaventrite and abdomen black; mouthparts and antenna yellow. Legs yellow, basal third of femur black. Head punctures small, separated by one diameter, each puncture slightly larger than an eye facet; pronotal punctures smaller than head punctures, separated by one and a half diameters; elytral punctures larger than pronotum and head punctures, separated by two diameters; metaventral punctures as large as elytron punctures, separated by one and a half diameters. Clypeus weakly emarginate apically, nearly truncate, lateral angle straight. Pronotum narrowed from base to apeX, sides curved. Prosternal carinae short, straight, divergent to prosternal anterior part.
</paragraph>
<caption id="DF2709A71B472610C3E8978479307ABA" httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="2" targetBox="[223,1324,220,1925]" targetPageId="2">
<paragraph id="8BE7592F1B472610C3E8978479307ABA" blockId="2.[151,1436,1973,2027]" pageId="2">
<emphasis id="B92C853D1B472610C3E897847A297A9B" bold="true" box="[151,316,1973,1996]" pageId="2">FIGURES 14.</emphasis>
<taxonomicName id="4C5822AC1B472610C23A978779767A9C" authorityName="Nestor-Arriola &amp; Toledo-Hernández" authorityYear="2017" box="[325,611,1974,1996]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="brevihamata">
<emphasis id="B92C853D1B472610C23A978779767A9C" box="[325,611,1974,1996]" italics="true" pageId="2">Brachiacantha brevihamata</emphasis>
</taxonomicName>
(images by Jorge Valdez Carrasco).
<emphasis id="B92C853D1B472610C09B978478EC7A9C" bold="true" box="[996,1017,1973,1996]" pageId="2">1)</emphasis>
dorsal habitus;
<emphasis id="B92C853D1B472610C7DF97847FA17A9C" bold="true" box="[1184,1204,1973,1996]" pageId="2">2)</emphasis>
tegmen, concave side vieW;
<emphasis id="B92C853D1B472610C3AA97E57BFE7ABB" bold="true" box="[213,235,2004,2027]" pageId="2">3)</emphasis>
tegmen, lateral vieW;
<emphasis id="B92C853D1B472610C2B397E57AF57ABB" bold="true" box="[460,480,2004,2027]" pageId="2">4)</emphasis>
sipho.
</paragraph>
</caption>
<paragraph id="8BE7592F1B462611C3E890A67BE77F6C" blockId="3.[151,1437,151,1544]" pageId="3">
Protibial tooth small, protibia not flanged. Epipleuron narrow, emarginate for reception of femoral apices. Abdomen with postcoXal line on basal ventrite slightly flattened along posterior ventrite margin, ventrite with sparse, short pubescence and dense, small punctures; ventrites 25 emarginated and depressed at the middle, pubescent and densely punctate at sides, glabrous and almost without punctures at the middle; ventrites 35 with a tubercle on each side of the emargination; siXth ventrite flattened, slightly depressed and glabrous at the middle, with lateral projections on sides, the projections tuberculate with abundant punctures and bristles. Genitalia with basal lobe slightly longer than the paramere, asymmetrical, sides almost parallel, slightly narrowed at the end of the second third, rounded at apeX (
<figureCitation id="136345AA1B462611C28E91A579237CFC" box="[497,566,404,429]" captionStart="FIGURES 14" captionStartId="2.[151,264,1973,1996]" captionTargetBox="[223,1324,220,1925]" captionTargetId="figure@2.[207,1379,193,1952]" captionTargetPageId="2" captionText="FIGURES 14. Brachiacantha brevihamata (images by Jorge Valdez Carrasco). 1) dorsal habitus; 2) tegmen, concave side vieW;3) tegmen, lateral vieW; 4) sipho." httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="3">Fig. 2</figureCitation>
), with a small apical hook on a side; paramere scimitar shaped, wide, curved, long, rounded at apeX, with a small tooth on the concave margin and abundant long bristles on the conveX and apical margins, shorter bristles on the apical half of the concave margin (
<figureCitation id="136345AA1B462611C09291ED7F217CA4" box="[1005,1076,476,501]" captionStart="FIGURES 14" captionStartId="2.[151,264,1973,1996]" captionTargetBox="[223,1324,220,1925]" captionTargetId="figure@2.[207,1379,193,1952]" captionTargetPageId="2" captionText="FIGURES 14. Brachiacantha brevihamata (images by Jorge Valdez Carrasco). 1) dorsal habitus; 2) tegmen, concave side vieW;3) tegmen, lateral vieW; 4) sipho." httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="3">Fig. 3</figureCitation>
); sipho with lateral, elongated, rounded alae at the apeX; siphonal capsule crested, heavily sclerotized, inner arm short and wide with apeX acute (
<figureCitation id="136345AA1B462611C3E092157BF07F6C" box="[159,229,548,573]" captionStart="FIGURES 14" captionStartId="2.[151,264,1973,1996]" captionTargetBox="[223,1324,220,1925]" captionTargetId="figure@2.[207,1379,193,1952]" captionTargetPageId="2" captionText="FIGURES 14. Brachiacantha brevihamata (images by Jorge Valdez Carrasco). 1) dorsal habitus; 2) tegmen, concave side vieW;3) tegmen, lateral vieW; 4) sipho." httpUri="https://zenodo.org/record/1116512/files/figure.png" pageId="3">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BE7592F1B462611C3B8927678467FA0" blockId="3.[151,1437,151,1544]" pageId="3">
<emphasis id="B92C853D1B462611C3B892767A307F30" bold="true" box="[199,293,583,608]" pageId="3">Female.</emphasis>
Length
<quantity id="4CA0F4CA1B462611C2F592767AF37F0F" box="[394,486,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" unit="mm" value="4.5">4.5 mm</quantity>
, width
<quantity id="4CA0F4CA1B462611C13F927979887F0F" box="[576,669,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" unit="mm" value="3.3">3.3 mm</quantity>
. Similar to male eXcept head black with an orange spot on front, pronotum without yellow anterior border but with the medial line yellowish orange, legs black or dark brown eXcept the tarsi and the tibial apeX yellow; abdomen without modifications. Genitalia with spermathecal capsule short, thick, with narrowed apeX; cornu, nodulus and ramus differentiated; bursa copulatriX subtriangular with two sclerotized arms; apical strut short, thin and laterally flattened.
</paragraph>
<paragraph id="8BE7592F1B462611C3B892CD78887E68" blockId="3.[151,1437,151,1544]" pageId="3">
<emphasis id="B92C853D1B462611C3B892CD7A297E45" bold="true" box="[199,316,764,789]" pageId="3">Variation.</emphasis>
Length
<quantity id="4CA0F4CA1B462611C2DD92CD79267E44" box="[418,563,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="3" unit="mm" value="3.85" valueMax="4.0" valueMin="3.7">3.7 to 4 mm</quantity>
width
<quantity id="4CA0F4CA1B462611C1F992CD783A7E44" box="[646,815,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="3" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3 to 2.8 mm</quantity>
. Scutellar and discal spots may be fused in hourglass shape. The elytral spots and pronotal marks can be pale yellow color.
</paragraph>
</subSubSection>
<subSubSection id="C3420AA41B462611C3B8937578BD7E0C" box="[199,936,836,861]" pageId="3" type="distribution">
<paragraph id="8BE7592F1B462611C3B8937578BD7E0C" blockId="3.[151,1437,151,1544]" box="[199,936,836,861]" pageId="3">
<emphasis id="B92C853D1B462611C3B893757A757E0D" bold="true" box="[199,352,836,861]" pageId="3">Distribution.</emphasis>
<collectingRegion id="499C97CD1B462611C21893757AD47E0D" box="[359,449,836,861]" country="Mexico" name="Mexico" pageId="3">
<collectingCountry id="F34F19BF1B462611C21893757AD47E0D" box="[359,449,836,861]" name="Mexico" pageId="3">MeXico</collectingCountry>
</collectingRegion>
; in
<collectingRegion id="499C97CD1B462611C2979375792B7E0D" box="[488,574,836,861]" country="Mexico" name="Mexico" pageId="3">
<collectingCountry id="F34F19BF1B462611C2979375792B7E0D" box="[488,574,836,861]" name="Mexico" pageId="3">MeXico</collectingCountry>
</collectingRegion>
State,
<collectingRegion id="499C97CD1B462611C1F5937578137E0D" box="[650,774,836,861]" country="Mexico" name="Michoacan de Ocampo" pageId="3">Michoacan</collectingRegion>
and
<collectingRegion id="499C97CD1B462611C042937478B17E0C" box="[829,932,837,860]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3">Veracruz</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3420AA41B462611C3B8935678357965" pageId="3" type="materials_examined">
<paragraph id="8BE7592F1B462611C3B893567AAD7EF4" blockId="3.[151,1437,151,1544]" pageId="3">
<emphasis id="B92C853D1B462611C3B893567A607ED0" bold="true" box="[199,373,871,896]" pageId="3">
<typeStatus id="54E3E78D1B462611C3B893567A177ED0" box="[199,258,871,896]" pageId="3">Type</typeStatus>
material.
</emphasis>
<emphasis id="B92C853D1B462611C20393567AF97ED0" box="[380,492,871,896]" italics="true" pageId="3">
<typeStatus id="54E3E78D1B462611C20393567AF37ED0" box="[380,486,871,896]" pageId="3" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingRegion id="499C97CD1B462611C28B935979727ED0" box="[500,615,872,896]" country="Mexico" name="Mexico" pageId="3">
<collectingCountry id="F34F19BF1B462611C28B935979727ED0" box="[500,615,872,896]" name="Mexico" pageId="3">MEXICO</collectingCountry>
</collectingRegion>
: “Estado de
<collectingRegion id="499C97CD1B462611C18C935678597ED0" box="[755,844,871,896]" country="Mexico" name="Mexico" pageId="3">
<collectingCountry id="F34F19BF1B462611C18C935678597ED0" box="[755,844,871,896]" name="Mexico" pageId="3">MéXico</collectingCountry>
</collectingRegion>
,
<collectingRegion id="499C97CD1B462611C028935678847ED0" box="[855,913,871,896]" country="Honduras" name="Valle" pageId="3">Valle</collectingRegion>
de Bravo,
<date id="FFE67FEF1B462611C77293567F937ED0" box="[1037,1158,871,896]" pageId="3" value="1962-06-01">1-VI-1962</date>
, col. Balderas and Ortiz, cat. 0576” (
<collectionCode id="ED49C1EA1B462611C26693BD7A787EF4" box="[281,365,908,932]" country="Mexico" httpUri="http://grbio.org/cool/ff28-ftqi" name="Centro de Entomologica y Acarologia" pageId="3">CEAM</collectionCode>
) (
<specimenCount id="9D5E92A61B462611C2F993BD7ABF7EF4" box="[390,426,908,932]" pageId="3" type="male">1♂</specimenCount>
).
</paragraph>
<paragraph id="8BE7592F1B462611C3B893807A867940" blockId="3.[151,1437,151,1544]" pageId="3">
<emphasis id="B92C853D1B462611C3B893807A547E98" box="[199,321,945,968]" italics="true" pageId="3">
<typeStatus id="54E3E78D1B462611C3B893807A297E98" box="[199,316,945,968]" pageId="3" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingRegion id="499C97CD1B462611C23693817AA97E98" box="[329,444,944,968]" country="Mexico" name="Mexico" pageId="3">
<collectingCountry id="F34F19BF1B462611C23693817AA97E98" box="[329,444,944,968]" name="Mexico" pageId="3">MEXICO</collectingCountry>
</collectingRegion>
: same data as the
<typeStatus id="54E3E78D1B462611C1F9939E79FD7E98" box="[646,744,943,968]" pageId="3" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED49C1EA1B462611C1889381785E7E98" box="[759,843,944,968]" country="Mexico" httpUri="http://grbio.org/cool/ff28-ftqi" name="Centro de Entomologica y Acarologia" pageId="3">CEAM</collectionCode>
) (
<specimenCount id="9D5E92A61B462611C01B9381789C7E98" box="[868,905,944,968]" pageId="3" type="male">1♂</specimenCount>
); “
<collectingRegion id="499C97CD1B462611C0D6939E7F3E7E98" box="[937,1067,943,968]" country="Mexico" name="Michoacan de Ocampo" pageId="3">Michoacán</collectingRegion>
, Zitácuaro, Presa El Bosque,
<date id="FFE67FEF1B462611C60893817BED7EBD" pageId="3" value="1986-06-23">23- VI-1986</date>
, col. E. Muriño” (
<collectionCode id="ED49C1EA1B462611C2BD93E5791F7EBC" box="[450,522,980,1004]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="3">CNIN</collectionCode>
) (
<specimenCount id="9D5E92A61B462611C15B93E579517EBC" box="[548,580,980,1004]" pageId="3" type="female">1♀</specimenCount>
); “
<collectingRegion id="499C97CD1B462611C11893E579C57EBC" box="[615,720,980,1004]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3">Veracruz</collectingRegion>
, Huatusco, 9.3 Miles south of Huatusco,
<date id="FFE67FEF1B462611C7E193E57E317EBC" box="[1182,1316,980,1004]" pageId="3" value="1983-06-18">18-VI-1983</date>
, col. B. K. Dozier” (
<collectionCode id="ED49C1EA1B462611C38193C97A527940" box="[254,327,1016,1040]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3">FSCA</collectionCode>
) (
<specimenCount id="9D5E92A61B462611C21D93C97A967940" box="[354,387,1016,1040]" pageId="3" type="female">1♀</specimenCount>
).
</paragraph>
<paragraph id="8BE7592F1B462611C3B8942D78357965" blockId="3.[151,1437,151,1544]" box="[199,800,1052,1077]" pageId="3">
<materialsCitation id="3B3053721B462611C3B8942D79007964" ID-GBIF-Occurrence="1707598862" box="[199,533,1052,1077]" location="Valle" pageId="3" specimenCount="1" stateProvince="Valle" typeStatus="holotype">
<emphasis id="B92C853D1B462611C3B8942D7A707965" bold="true" box="[199,357,1052,1077]" pageId="3">
<typeStatus id="54E3E78D1B462611C3B8942D7A177965" box="[199,258,1052,1077]" pageId="3">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="499C97CD1B462611C213942D7AB37965" box="[364,422,1052,1077]" country="Honduras" name="Valle" pageId="3">Valle</collectingRegion>
de Bravo
</materialsCitation>
, MeXico State, MeXico.
</paragraph>
</subSubSection>
<subSubSection id="C3420AA41B462611C3B8940E7E887891" pageId="3" type="discussion">
<paragraph id="8BE7592F1B462611C3B8940E7E887891" blockId="3.[151,1437,151,1544]" pageId="3">
<emphasis id="B92C853D1B462611C3B8940E7A2E7908" bold="true" box="[199,315,1087,1112]" pageId="3">Remarks.</emphasis>
In coloration, this species is very similar to that of the
<emphasis id="B92C853D1B462611C0D1947078E37908" box="[942,1014,1089,1112]" italics="true" pageId="3">ursina</emphasis>
group, but differs in having the male abdomen with several depressed ventrites and the asymmetrical basal lobe of the male genitalia. For the “
<emphasis id="B92C853D1B462611C3DC94B67A9679F0" box="[163,387,1159,1184]" italics="true" pageId="3">longitudinal fossa”</emphasis>
this species belongs to “group 5” of
<bibRefCitation id="EFC924DE1B462611C02794B878FE79F0" author="Leng" box="[856,1003,1160,1185]" pageId="3" refString="Leng, C. W. (1911) The species of Brachyacantha of North and South America. Bulletin American Museum of Natural History, 30, 279 - 333." type="journal article" year="1911">Leng (1911)</bibRefCitation>
, a Central American species group. Similarities in the male abdomen have been described in South American species such as
<taxonomicName id="4C5822AC1B462611C7C0949D7AE779B8" authority="Gordon, Canepari &amp; Hanley" authorityName="Gordon, Canepari &amp; Hanley" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="gail">
<emphasis id="B92C853D1B462611C7C0949D7E897995" box="[1215,1436,1196,1221]" italics="true" pageId="3">Brachiacantha gail</emphasis>
Gordon, Canepari &amp; Hanley
</taxonomicName>
, from the group
<emphasis id="B92C853D1B462611C1AD94FE782979B8" box="[722,828,1231,1256]" italics="true" pageId="3">bahiensis</emphasis>
, and
<taxonomicName id="4C5822AC1B462611C0F294E07E7A79B8" authority="Gordon, Canepari &amp; Hanley" authorityName="Gordon, Canepari &amp; Hanley" box="[909,1391,1231,1256]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="valerie">
<emphasis id="B92C853D1B462611C0F294E07F1679B8" box="[909,1027,1231,1256]" italics="true" pageId="3">B. valerie</emphasis>
Gordon, Canepari &amp; Hanley
</taxonomicName>
,
<taxonomicName id="4C5822AC1B462611C6FB94E07A6B785D" authority="Weise" authorityName="Weise" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B92C853D1B462611C6FB94E07A3A785C" italics="true" pageId="3">B. tucumanensis</emphasis>
Weise
</taxonomicName>
and
<taxonomicName id="4C5822AC1B462611C2C394C4789C785D" authority="Gordon, Canepari &amp; Hanley" authorityName="Gordon, Canepari &amp; Hanley" box="[444,905,1268,1293]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="eleanor">
<emphasis id="B92C853D1B462611C2C394C47921785D" box="[444,564,1268,1293]" italics="true" pageId="3">B. eleanor</emphasis>
Gordon, Canepari &amp; Hanley
</taxonomicName>
, from the group
<taxonomicName id="4C5822AC1B462611C72C94C47BC97860" authority="(Gordon et al. 2014)" authorityName="Gordon et al." authorityYear="2014" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B92C853D1B462611C72C94C47FF9785C" box="[1107,1260,1269,1292]" italics="true" pageId="3">tucumanensis</emphasis>
(
<bibRefCitation id="EFC924DE1B462611C78194C57BC17860" author="Gordon" pageId="3" refString="Gordon, R. D., Canepari, C. &amp; Hanley, G. (2014) South American Coccinellidae (Coleoptera) Part XVI: systematic revision of Brachiacantha Dejean (Coccinellinae: Hyperaspidini). Insecta Mundi, 390, 1 - 76." type="journal article" year="2014">
Gordon
<emphasis id="B92C853D1B462611C61F94C47E89785D" box="[1376,1436,1268,1293]" italics="true" pageId="3">et al.</emphasis>
2014
</bibRefCitation>
)
</taxonomicName>
. The genitalia of
<taxonomicName id="4C5822AC1B462611C2DC952879457860" authorityName="Nestor-Arriola &amp; Toledo-Hernández" authorityYear="2017" box="[419,592,1303,1328]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brevihamata">
<emphasis id="B92C853D1B462611C2DC952879457860" box="[419,592,1303,1328]" italics="true" pageId="3">B. brevihamata</emphasis>
</taxonomicName>
have only a few similarities with the South American species, such as the elongated shape of the basal lobe and the paramere shape, but the asymmetrical apeX and the small hook differentiates it from the majority of the South American groups. The asymmetrical male genitalia are characteristic of the
<emphasis id="B92C853D1B462611C20695B57AC378CD" box="[377,470,1412,1437]" italics="true" pageId="3">dentipes</emphasis>
group, but
<taxonomicName id="4C5822AC1B462611C12995B4781478CD" authorityName="Nestor-Arriola &amp; Toledo-Hernández" authorityYear="2017" box="[598,769,1412,1437]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brevihamata">
<emphasis id="B92C853D1B462611C12995B4781478CD" box="[598,769,1412,1437]" italics="true" pageId="3">B. brevihamata</emphasis>
</taxonomicName>
lacks the characteristic cusps in the third male ventrite. The relationships with other species are not clear and it is possible that
<taxonomicName id="4C5822AC1B462611C00095987F3E7890" authorityName="Nestor-Arriola &amp; Toledo-Hernández" authorityYear="2017" box="[895,1067,1447,1472]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="brevihamata">
<emphasis id="B92C853D1B462611C00095987F3E7890" box="[895,1067,1447,1472]" italics="true" pageId="3">B. brevihamata</emphasis>
</taxonomicName>
must be placed in its own group.
</paragraph>
</subSubSection>
<subSubSection id="C3420AA41B462611C3B895FD792D7B58" pageId="3" type="etymology">
<paragraph id="8BE7592F1B462611C3B895FD792D7B58" blockId="3.[151,1437,151,1544]" pageId="3">
<emphasis id="B92C853D1B462611C3B895FD7A5B78B5" bold="true" box="[199,334,1484,1509]" pageId="3">Etymology.</emphasis>
From Latin “brevis” (short) and “hamatus” (hooked). The name refers to the small hook in the basal lobe apeX of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>