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<document id="1A97090559DAFB9B609B4125B592ADB7" ID-DOI="10.1046/j.1096-3642.2002.00008.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5434168" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630432444395" checkinUser="carolina" docAuthor="Scoble, Malcolm J. &amp; Krüger, Martin" docDate="2002" docId="03F16A201E14FFAEFEC7A579FDC2DF72" docLanguage="en" docName="j.1096-3642.2002.00008.x.pdf" docOrigin="Zoological Journal of the Linnean Society 134 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00008.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Milocera Swinhoe 1904" docType="treatment" docVersion="5" lastPageNumber="269" masterDocId="FFC812581E1FFFA2FF9AA439FFEDDA11" masterDocTitle="A review of the genera of Macariini with a revised classification of the tribe (Geometridae: Ennominae)" masterLastPageNumber="315" masterPageNumber="257" pageNumber="268" updateTime="1699268739040" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="766BA81796404AD7FC0FB812B02B6CE7">A review of the genera of Macariini with a revised classification of the tribe (Geometridae: Ennominae)</mods:title>
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<mods:namePart id="09C736FADD9E6D1EA39C50344080C41F">Scoble, Malcolm J.</mods:namePart>
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<subSubSection id="C34288BD1E14FFA9FEC7A579FDC4DB6F" pageId="11" pageNumber="268" type="nomenclature">
<paragraph id="8BE7DB361E14FFA9FEC7A579FDA4DB47" blockId="11.[349,585,319,343]" box="[349,585,319,343]" pageId="11" pageNumber="268">
<heading id="D0AF6C5A1E14FFA9FEC7A579FDA4DB47" box="[349,585,319,343]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="268" reason="2">
<taxonomicName id="4C58A0B51E14FFA9FEC7A579FDA4DB47" ID-CoL="92FSN" authority="SWINHOE" authorityName="Swinhoe" authorityYear="1904" box="[349,585,319,343]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E14FFA9FEC7A579FE38DB47" box="[349,469,320,343]" italics="true" pageId="11" pageNumber="268">MILOCERA</emphasis>
SWINHOE
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE7DB361E14FFA9FEE7A551FDC4DB6F" blockId="11.[381,553,360,382]" box="[381,553,360,382]" pageId="11" pageNumber="268">(Figs 2, 6467)</paragraph>
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<paragraph id="8BE7DB361E14FFA9FF05A5AFFDA0DBDB" blockId="11.[159,775,406,459]" pageId="11" pageNumber="268">
<taxonomicName id="4C58A0B51E14FFA9FF05A5AFFE3DDBBA" authority="Swinhoe, 1904" authorityName="Swinhoe" authorityYear="1904" box="[159,464,406,428]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E14FFA9FF05A5AFFEEFDBBA" box="[159,258,406,427]" italics="true" pageId="11" pageNumber="268">Milocera</emphasis>
Swinhoe, 1904
</taxonomicName>
.
<typeStatus id="54E365941E14FFA9FE70A5AFFDCCDBBA" box="[490,545,406,427]" pageId="11" pageNumber="268">Type</typeStatus>
species:
<taxonomicName id="4C58A0B51E14FFA9FD3EA5AFFE5CDBDA" authority="Swinhoe (1904)" authorityName="Swinhoe" authorityYear="1904" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="species" species="horaria">
<emphasis id="B92C07241E14FFA9FD3EA5AFFF18DBD8" italics="true" pageId="11" pageNumber="268">Milocera horaria</emphasis>
Swinhoe (1904)
</taxonomicName>
.
<collectingCountry id="F34F9BA61E14FFA9FE27A58DFDA7DBDB" box="[445,586,436,458]" name="Madagascar" pageId="11" pageNumber="268">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BE7DB361E14FFA9FCADA4FCFC51DACD" blockId="11.[823,956,197,220]" box="[823,956,197,220]" pageId="11" pageNumber="268">
<emphasis id="B92C07241E14FFA9FCADA4FCFC51DACD" box="[823,956,197,220]" italics="true" pageId="11" pageNumber="268">Description</emphasis>
</paragraph>
<paragraph id="8BE7DB361E14FFA9FCADA4CDFC06DB38" blockId="11.[823,1439,244,297]" pageId="11" pageNumber="268">
<emphasis id="B92C07241E14FFA9FCADA4CDFC99DB18" box="[823,884,244,265]" italics="true" pageId="11" pageNumber="268">Head</emphasis>
. Chaetosemata extended. Antenna fasciculate, not bipectinate.
</paragraph>
<paragraph id="8BE7DB361E14FFA9FCADA57CFAFBDB68" blockId="11.[823,1438,325,377]" pageId="11" pageNumber="268">
<emphasis id="B92C07241E14FFA9FCADA57CFC96DB4B" box="[823,891,325,346]" italics="true" pageId="11" pageNumber="268">Wings</emphasis>
(Fig. 2). Often weakly falcate; ranging from off-white, through ochreous to red-brown.
</paragraph>
<paragraph id="8BE7DB361E14FFAEFCADA5AFFDEBDAE8" blockId="11.[823,1439,406,458]" lastBlockId="12.[145,761,197,250]" lastPageId="12" lastPageNumber="269" pageId="11" pageNumber="268">
<emphasis id="B92C07241E14FFA9FCADA5AFFC09DBBA" box="[823,996,406,427]" italics="true" pageId="11" pageNumber="268">Male genitalia</emphasis>
(
<figureCitation id="1363C7B31E14FFA9FC61A5AFFB60DBBD" box="[1019,1165,406,428]" captionStart="Figures 6467" captionStartId="11.[159,249,1853,1874]" captionTargetBox="[160,1440,594,1819]" captionTargetId="figure-104@11.[160,1440,594,1821]" captionTargetPageId="11" captionText="Figures 6467. Milocera species: 64, M. horaria, ♂ genitalia; 65, M. horaria, ♀ genitalia; 66, M. arcifera, ♂ genitalia; 67, M. arcifera, ♀ genitalia." figureDoi="http://doi.org/10.5281/zenodo.5434172" httpUri="https://zenodo.org/record/5434172/files/figure.png" pageId="11" pageNumber="268">Figs 64, 66</figureCitation>
). Uncus usually fairly narrow, pointed or truncated. Gnathos present. Valva divided; shape of costa and of sacculus varies; stout spine-like setae usually present.
</paragraph>
<caption id="DF278BBE1E14FFA9FF05A304FE2ADD7E" ID-DOI="http://doi.org/10.5281/zenodo.5434172" ID-Zenodo-Dep="5434172" httpUri="https://zenodo.org/record/5434172/files/figure.png" pageId="11" pageNumber="268" startId="11.[159,249,1853,1874]" targetBox="[160,1440,594,1819]" targetPageId="11">
<paragraph id="8BE7DB361E14FFA9FF05A304FE2ADD7E" blockId="11.[159,1439,1853,1904]" pageId="11" pageNumber="268">
<emphasis id="B92C07241E14FFA9FF05A304FEAADD43" bold="true" box="[159,327,1853,1874]" pageId="11" pageNumber="268">Figures 6467.</emphasis>
<taxonomicName id="4C58A0B51E14FFA9FEC8A304FDEEDD43" box="[338,515,1853,1874]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92C07241E14FFA9FEC8A304FE40DD43" box="[338,429,1853,1874]" italics="true" pageId="11" pageNumber="268">Milocera</emphasis>
species
</taxonomicName>
: 64,
<taxonomicName id="4C58A0B51E14FFA9FDACA307FD44DD43" authorityName="Swinhoe" authorityYear="1904" box="[566,681,1853,1875]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="species" species="horaria">
<emphasis id="B92C07241E14FFA9FDACA307FD44DD43" box="[566,681,1853,1875]" italics="true" pageId="11" pageNumber="268">M. horaria</emphasis>
</taxonomicName>
, ♂ genitalia; 65,
<taxonomicName id="4C58A0B51E14FFA9FCF3A307FC31DD43" authorityName="Swinhoe" authorityYear="1904" box="[873,988,1853,1875]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="species" species="horaria">
<emphasis id="B92C07241E14FFA9FCF3A307FC31DD43" box="[873,988,1853,1875]" italics="true" pageId="11" pageNumber="268">M. horaria</emphasis>
</taxonomicName>
, ♀ genitalia; 66,
<taxonomicName id="4C58A0B51E14FFA9FB03A307FAE3DD43" authorityName="Hampson" authorityYear="1910" box="[1177,1294,1853,1875]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="species" species="arcifera">
<emphasis id="B92C07241E14FFA9FB03A307FAE3DD43" box="[1177,1294,1853,1875]" italics="true" pageId="11" pageNumber="268">M. arcifera</emphasis>
</taxonomicName>
, ♂ genitalia; 67,
<taxonomicName id="4C58A0B51E14FFA9FF5CA362FED7DD7E" authorityName="Hampson" authorityYear="1910" box="[198,314,1882,1904]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="268" phylum="Arthropoda" rank="species" species="arcifera">
<emphasis id="B92C07241E14FFA9FF5CA362FED7DD7E" box="[198,314,1882,1904]" italics="true" pageId="11" pageNumber="268">M. arcifera</emphasis>
</taxonomicName>
, ♀ genitalia.
</paragraph>
</caption>
<paragraph id="8BE7DB361E13FFAEFF0BA534FE7ADB8F" blockId="12.[145,761,269,414]" pageId="12" pageNumber="269">
<emphasis id="B92C07241E13FFAEFF0BA534FE05DB33" box="[145,488,269,290]" italics="true" pageId="12" pageNumber="269">Pregenital abdomen of male</emphasis>
. Sternum A3 with or without a patch of bristles medially. Sternum A8 with margin weakly emarginated in
<typeStatus id="54E365941E13FFAEFD99A572FDD9DB71" box="[515,564,331,352]" pageId="12" pageNumber="269">type</typeStatus>
species, ranging to bilobed in some other species and with curved hornlike octavals in others.
</paragraph>
<paragraph id="8BE7DB361E13FFAEFF0BA58FFF13D836" blockId="12.[145,761,437,551]" pageId="12" pageNumber="269">
<emphasis id="B92C07241E13FFAEFF0BA58FFEBEDBDA" box="[145,339,438,459]" italics="true" pageId="12" pageNumber="269">Female genitalia</emphasis>
(
<figureCitation id="1363C7B31E13FFAEFEFCA58FFE00DBDA" box="[358,493,437,460]" captionStart="Figures 6467" captionStartId="11.[159,249,1853,1874]" captionTargetBox="[160,1440,594,1819]" captionTargetId="figure-104@11.[160,1440,594,1821]" captionTargetPageId="11" captionText="Figures 6467. Milocera species: 64, M. horaria, ♂ genitalia; 65, M. horaria, ♀ genitalia; 66, M. arcifera, ♂ genitalia; 67, M. arcifera, ♀ genitalia." figureDoi="http://doi.org/10.5281/zenodo.5434172" httpUri="https://zenodo.org/record/5434172/files/figure.png" pageId="12" pageNumber="269">Figs 65, 67</figureCitation>
). Corpus bursae small and usually densely covered with spines (as in
<taxonomicName id="4C58A0B51E13FFAEFF0BA5CAFEE7D819" authorityName="Warren" authorityYear="1900" box="[145,266,499,520]" class="Insecta" family="Geometridae" genus="Platypepla" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFF0BA5CAFEE7D819" box="[145,266,499,520]" italics="true" pageId="12" pageNumber="269">Platypepla</emphasis>
</taxonomicName>
), but spines absent in
<typeStatus id="54E365941E13FFAEFDFAA5CAFD7CD819" box="[608,657,499,520]" pageId="12" pageNumber="269">type</typeStatus>
species (
<taxonomicName id="4C58A0B51E13FFAEFF03A62BFF02D836" authorityName="Swinhoe" authorityYear="1904" box="[153,239,530,551]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="species" species="horaria">
<emphasis id="B92C07241E13FFAEFF03A62BFF02D836" box="[153,239,530,551]" italics="true" pageId="12" pageNumber="269">horaria</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE7DB361E13FFAEFF0BA679FD84D95B" blockId="12.[145,761,576,843]" pageId="12" pageNumber="269">
<emphasis id="B92C07241E13FFAEFF0BA679FEECD844" box="[145,257,576,597]" italics="true" pageId="12" pageNumber="269">Diagnosis</emphasis>
. The
<typeStatus id="54E365941E13FFAEFED6A679FE90D844" box="[332,381,576,597]" pageId="12" pageNumber="269">type</typeStatus>
species, which occurs in
<collectingCountry id="F34F9BA61E13FFAEFD2AA679FF31D865" name="Madagascar" pageId="12" pageNumber="269">Madagascar</collectingCountry>
, differs in the structure of the genitalia from all other species of
<taxonomicName id="4C58A0B51E13FFAEFED6A644FE42D883" authorityName="Swinhoe" authorityYear="1904" box="[332,431,637,658]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFED6A644FE42D883" box="[332,431,637,658]" italics="true" pageId="12" pageNumber="269">Milocera</emphasis>
</taxonomicName>
, notably in the shape of the valva (see
<figureCitation id="1363C7B31E13FFAEFE94A6A5FE8FD8A0" box="[270,354,667,690]" captionStart="Figures 6467" captionStartId="11.[159,249,1853,1874]" captionTargetBox="[160,1440,594,1819]" captionTargetId="figure-104@11.[160,1440,594,1821]" captionTargetPageId="11" captionText="Figures 6467. Milocera species: 64, M. horaria, ♂ genitalia; 65, M. horaria, ♀ genitalia; 66, M. arcifera, ♂ genitalia; 67, M. arcifera, ♀ genitalia." figureDoi="http://doi.org/10.5281/zenodo.5434172" httpUri="https://zenodo.org/record/5434172/files/figure.png" pageId="12" pageNumber="269">Fig. 64</figureCitation>
). The valva of
<taxonomicName id="4C58A0B51E13FFAEFD8BA6A5FD62D8A0" authorityName="Swinhoe" authorityYear="1904" box="[529,655,668,689]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="species" species="horaria">
<emphasis id="B92C07241E13FFAEFD8BA6A5FD62D8A0" box="[529,655,668,689]" italics="true" pageId="12" pageNumber="269">M. horaria</emphasis>
</taxonomicName>
does not bear stout setae, which differs from the situation in many species. In the female, the shape of the corpus bursae is small and round, typical of the
<taxonomicName id="4C58A0B51E13FFAEFDEDA6C1FD1DD91C" authorityName="Warren" authorityYear="1900" box="[631,752,760,781]" class="Insecta" family="Geometridae" genus="Platypepla" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFDEDA6C1FD1DD91C" box="[631,752,760,781]" italics="true" pageId="12" pageNumber="269">Platypepla</emphasis>
</taxonomicName>
- group of genera, but it lacks the dense arrangement of spines found in other species of
<taxonomicName id="4C58A0B51E13FFAEFE65A70CFD8FD95B" authorityName="Swinhoe" authorityYear="1904" box="[511,610,821,842]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFE65A70CFD8FD95B" box="[511,610,821,842]" italics="true" pageId="12" pageNumber="269">Milocera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE7DB361E13FFAEFF0BA75AFECCD989" blockId="12.[145,761,867,920]" pageId="12" pageNumber="269">
<emphasis id="B92C07241E13FFAEFF0BA75AFEF1D969" box="[145,284,867,888]" italics="true" pageId="12" pageNumber="269">Distribution</emphasis>
. Afrotropical region north of
<geoCoordinate id="EE6CBDF11E13FFAEFD1CA75AFD51D969" box="[646,700,867,888]" degrees="21" direction="south" orientation="latitude" pageId="12" pageNumber="269" precision="55555" value="-21.0">21°S</geoCoordinate>
, and
<collectingCountry id="F34F9BA61E13FFAEFF0BA7BBFEF3D989" box="[145,286,898,920]" name="Madagascar" pageId="12" pageNumber="269">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BE7DB361E13FFAEFF0BA789FD4DDF06" blockId="12.[145,762,944,1303]" pageId="12" pageNumber="269">
<emphasis id="B92C07241E13FFAEFF0BA789FEE4D9D4" box="[145,265,944,965]" italics="true" pageId="12" pageNumber="269">Comments</emphasis>
. The inclusion of species additional to the
<typeStatus id="54E365941E13FFAEFF56A7F6FF10D9F5" box="[204,253,975,996]" pageId="12" pageNumber="269">type</typeStatus>
species by
<bibRefCitation id="EFC9A6C71E13FFAEFE36A7F6FDA9D9F2" author="Prout LM" box="[428,580,974,996]" pageId="12" pageNumber="269" pagination="327 - 363" refId="ref25017" refString="Prout LM. 1922. New and little-known Geometridae. Novitates Zoologicae 29: 327 - 363." type="journal article" year="1922">Prout (1922</bibRefCitation>
,
<bibRefCitation id="EFC9A6C71E13FFAEFDC5A7F7FD75D9F2" author="Prout LB" box="[607,664,974,995]" pageId="12" pageNumber="269" pagination="375 - 512" refId="ref25035" refString="Prout LB. 1932. Voyage de Ch. Alluaud et R. Jeannel en Afrique orientale (1911 - 12). Memoires de la Societe Zoologique de France 29: 375 - 512." type="journal article" year="1932">1932</bibRefCitation>
,
<bibRefCitation id="EFC9A6C71E13FFAEFD29A7F7FD00D9F5" author="Prout LB" box="[691,749,974,996]" pageId="12" pageNumber="269" pagination="82 - 97" refId="ref25071" refString="Prout LB. 1934. New Congo Geometridae. Revue de Zoologie et de Botanique Africaines 26: 82 - 97." type="journal article" year="1934">1934</bibRefCitation>
),
<bibRefCitation id="EFC9A6C71E13FFAEFF0BA7D4FED2DE13" author="Herbulot C" box="[145,319,1005,1027]" pageId="12" pageNumber="269" pagination="67 - 76" refId="ref24385" refString="Herbulot C. 1973. Nouveaux Ennominae de l'Ouest du Cameroun (Lepidoptera Geometridae). Annales de la Faculte des Sciences du Cameroun 13: 67 - 76." type="journal article" year="1973">Herbulot (1973</bibRefCitation>
,
<bibRefCitation id="EFC9A6C71E13FFAEFED0A7D4FE6CDE12" author="Herbulot C" box="[330,385,1005,1027]" pageId="12" pageNumber="269" pagination="3 - 11" refId="ref24450" refString="Herbulot C. 1989. Nouveaux Ennominae d'Afrique intertropicale et de Madagascar (Lepidoptera Geometridae). Lambillionea 89: 3 - 11." type="journal article" year="1989">1989</bibRefCitation>
) and, particularly,
<bibRefCitation id="EFC9A6C71E13FFAEFDC3A7D4FD14DE12" author="Kruger M" box="[601,761,1005,1027]" pageId="12" pageNumber="269" pagination="1 - 502" refId="ref24679" refString="Kruger M. 2001. A revision of the tribe Macariini (Lepidoptera: Geometridae: Ennominae) of Africa, Madagascar and Arabia. Bulletin of the Natural History Museum (Entomology) 70 (1): 1 - 502." type="journal article" year="2001">Krüger (2001)</bibRefCitation>
is accepted here provisionally on grounds mainly of similarity in general appearance of the moths and, in particular, the weakly falcate shape of the forewings. An alternative approach would be to synonymize
<taxonomicName id="4C58A0B51E13FFAEFF0BA0BFFF19DE8A" authorityName="Swinhoe" authorityYear="1904" box="[145,244,1158,1179]" class="Insecta" family="Geometridae" genus="Milocera" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFF0BA0BFFF19DE8A" box="[145,244,1158,1179]" italics="true" pageId="12" pageNumber="269">Milocera</emphasis>
</taxonomicName>
with
<taxonomicName id="4C58A0B51E13FFAEFEDBA0BFFE57DE8A" authorityName="Warren" authorityYear="1900" box="[321,442,1158,1179]" class="Insecta" family="Geometridae" genus="Platypepla" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFEDBA0BFFE57DE8A" box="[321,442,1158,1179]" italics="true" pageId="12" pageNumber="269">Platypepla</emphasis>
</taxonomicName>
, thus expanding considerably the concept of this latter genus. The species of
<taxonomicName id="4C58A0B51E13FFAEFF0BA0FDFEE7DEC8" authorityName="Warren" authorityYear="1900" box="[145,266,1220,1241]" class="Insecta" family="Geometridae" genus="Platypepla" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B92C07241E13FFAEFF0BA0FDFEE7DEC8" box="[145,266,1220,1241]" italics="true" pageId="12" pageNumber="269">Platypepla</emphasis>
</taxonomicName>
(in the sense adopted here), however, form a monophyletic group based on characters from the wings and, particularly, of the male genitalia.
</paragraph>
<paragraph id="8BE7DB361E13FFAEFF0BA116FDC2DF72" blockId="12.[145,761,1327,1380]" pageId="12" pageNumber="269">
<emphasis id="B92C07241E13FFAEFF0BA116FE82DF54" box="[145,367,1327,1349]" italics="true" pageId="12" pageNumber="269">Number of species.</emphasis>
Twenty-six species are listed by
<bibRefCitation id="EFC9A6C71E13FFAEFF0BA177FED8DF75" author="Kruger M" box="[145,309,1358,1380]" pageId="12" pageNumber="269" pagination="1 - 502" refId="ref24679" refString="Kruger M. 2001. A revision of the tribe Macariini (Lepidoptera: Geometridae: Ennominae) of Africa, Madagascar and Arabia. Bulletin of the Natural History Museum (Entomology) 70 (1): 1 - 502." type="journal article" year="2001">Krüger (2001)</bibRefCitation>
, 16 of which are new.
</paragraph>
</subSubSection>
</treatment>
</document>