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<mods:title id="844C72B2FDF60B78A3A67776D7125084">New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic</mods:title>
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<mods:namePart id="21F470E0A81A08D4D3520668A48F54F9">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="7558B0D026D393B176EAF7DAA6D8A96B">Ng, Peter K. L.</mods:namePart>
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<paragraph id="8BFD36CC3B79E314FC4EFAB4FAD398D3" blockId="66.[904,1360,1405,1429]" box="[904,1360,1405,1429]" pageId="66" pageNumber="462">
<heading id="D0B581A03B79E314FC4EFAB4FAD398D3" bold="true" box="[904,1360,1405,1429]" centered="true" fontSize="10" level="2" pageId="66" pageNumber="462" reason="2">
<taxonomicName id="4C424D4F3B79E314FC4EFAB4FAD398D3" ID-CoL="7PMH5" authority="Huang, Hsueh &amp; Ng, 2002" authorityName="Huang, Hsueh &amp; Ng" authorityYear="2002" box="[904,1360,1405,1429]" class="Malacostraca" family="Hexapodidae" genus="Latohexapus" kingdom="Animalia" order="Decapoda" pageId="66" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B79E314FC4EFAB4FAD398D3" bold="true" box="[904,1360,1405,1429]" pageId="66" pageNumber="462">
<emphasis id="B936EADE3B79E314FC4EFAB4FB9598D3" bold="true" box="[904,1046,1405,1429]" italics="true" pageId="66" pageNumber="462">Latohexapus</emphasis>
<bibRefCitation id="EFD34B3D3B79E314FBD9FAB4FAD398D3" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" box="[1055,1360,1405,1429]" pageId="66" pageNumber="462" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang, Hsueh &amp; Ng, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C35865473B79E314FCEBFA75FB729894" box="[813,1265,1468,1490]" pageId="66" pageNumber="462" type="reference_group">
<paragraph id="8BFD36CC3B79E314FCEBFA75FB729894" blockId="66.[813,1265,1468,1490]" box="[813,1265,1468,1490]" pageId="66" pageNumber="462">
<treatmentCitationGroup id="AB5211E23B79E314FCEBFA75FB729894" box="[813,1265,1468,1490]" pageId="66" pageNumber="462">
<treatmentCitation id="0AE310DD3B79E314FCEBFA75FB6E9894" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" box="[813,1261,1468,1490]" page="652" pageId="66" pageNumber="462" year="2002">
<taxonomicName id="4C424D4F3B79E314FCEBFA75FB6E9894" ID-CoL="7PMH5" authority="Huang, Hsueh &amp; Ng, 2002: 652" authorityName="Huang, Hsueh &amp; Ng" authorityPageNumber="652" authorityYear="2002" box="[813,1261,1468,1490]" class="Malacostraca" family="Hexapodidae" genus="Latohexapus" kingdom="Animalia" order="Decapoda" pageId="66" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B79E314FCEBFA75FC2B9897" box="[813,936,1468,1489]" italics="true" pageId="66" pageNumber="462">Latohexapus</emphasis>
<bibRefCitation id="EFD34B3D3B79E314FC69FA75FB6E9894" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" box="[943,1261,1468,1490]" pageId="66" pageNumber="462" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang, Hsueh &amp; Ng, 2002: 652</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C35865473B79E312FCEBFA33FDFC9DB3" lastPageId="68" lastPageNumber="464" pageId="66" pageNumber="462" type="diagnosis">
<paragraph id="8BFD36CC3B79E314FCEBFA33FA939574" blockId="66.[813,1452,1530,2098]" pageId="66" pageNumber="462">
<emphasis id="B936EADE3B79E314FCEBFA33FB9A9B54" bold="true" box="[813,1049,1530,1554]" pageId="66" pageNumber="462">Emended diagnosis.</emphasis>
Carapace much broader than long; regions distinct, separated by deep groove, strongly granulated. Eye movable; anterolateral margin arcuate. Pterygostomial region with row of striae, granular; ischium of third maxillipeds expanded distally, longer than merus; exopod with well developed flagellum. Chelipeds asymmetrical, with prominent teeth at base of fingers. Surfaces of P2P4 granular, dactylus upcurved. Male thoracic sternum very broad; surface prominently granular; sternites 1 and 2 fused, strongly bent backward, separated from sternite 3 by distinct transverse ridge; deep transverse groove separating thoracic sternites 3 and 4; sternoabdominal cavity extending from edge of abdominal cavity to base of chelipeds; sternite 8 exposed. Male abdomen with somites 35 fused; somite 6 appearing hexagonal, with median part prominently expanded into triangular projection; telson subpentagonal. G1 very slender, directed anterolaterally, distal quarter not concealed under abdomen.
</paragraph>
<caption id="DF3D66443B78E315FF4EF7C6FC799504" ID-DOI="http://doi.org/10.5281/zenodo.5354065" ID-Zenodo-Dep="5354065" httpUri="https://zenodo.org/record/5354065/files/figure.png" pageId="67" pageNumber="463" startId="67.[136,171,2063,2085]" targetBox="[136,1474,336,2030]" targetPageId="67">
<paragraph id="8BFD36CC3B78E315FF4EF7C6FC799504" blockId="67.[136,1475,2063,2114]" pageId="67" pageNumber="463">
Fig. 54.
<taxonomicName id="4C424D4F3B78E315FF1CF7C6FD419562" authority="Huang, Hsueh &amp; Ng, 2002" authorityName="Huang, Hsueh &amp; Ng" authorityYear="2002" box="[218,706,2063,2085]" class="Malacostraca" family="Hexapodidae" genus="Latohexapus" kingdom="Animalia" order="Decapoda" pageId="67" pageNumber="463" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B936EADE3B78E315FF1CF7C6FE379563" box="[218,436,2063,2085]" italics="true" pageId="67" pageNumber="463">Latohexapus granosus</emphasis>
<bibRefCitation id="EFD34B3D3B78E315FE7DF7C6FD419562" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" box="[443,706,2063,2085]" pageId="67" pageNumber="463" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang, Hsueh &amp; Ng, 2002</bibRefCitation>
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, holotype male (16.3 × 8.2 mm) (ZRC 2002.001). A, overall view of carapace, chelipeds and P2P4; B, frontal view of cephalothorax; C, ventral view of cephalothorax.
</paragraph>
</caption>
<paragraph id="8BFD36CC3B7FE312FFB7FF75FDFC9DB3" blockId="68.[113,752,188,245]" pageId="68" pageNumber="464">
<emphasis id="B936EADE3B7FE312FFB7FF75FEB59D92" bold="true" box="[113,310,188,212]" pageId="68" pageNumber="464">Species included.</emphasis>
<taxonomicName id="4C424D4F3B7FE312FEF8FF74FF559DB3" authority="Huang, Hsueh &amp; Ng, 2002" authorityName="Huang, Hsueh &amp; Ng" authorityYear="2002" class="Malacostraca" family="Hexapodidae" genus="Latohexapus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="464" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B936EADE3B7FE312FEF8FF74FDB39D92" box="[318,560,189,213]" italics="true" pageId="68" pageNumber="464">Latohexapus granosus</emphasis>
<bibRefCitation id="EFD34B3D3B7FE312FDFFFF74FF559DB3" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" pageId="68" pageNumber="464" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang, Hsueh &amp; Ng, 2002</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54F9886E3B7FE312FF21FF14FE959DB3" box="[231,278,221,245]" pageId="68" pageNumber="464">type</typeStatus>
species by original designation).
</paragraph>
</subSubSection>
<subSubSection id="C35865473B7FE312FFB7FED4FD729FF3" pageId="68" pageNumber="464" type="discussion">
<paragraph id="8BFD36CC3B7FE312FFB7FED4FD729FF3" blockId="68.[113,753,285,693]" pageId="68" pageNumber="464">
<emphasis id="B936EADE3B7FE312FFB7FED4FF639C73" bold="true" box="[113,224,285,309]" pageId="68" pageNumber="464">Remarks.</emphasis>
So far this is the only genus in the family that has the carapace much broader than long and with the regions distinctly demarcated. As discussed by
<bibRefCitation id="EFD34B3D3B7FE312FDABFE94FF389CD3" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" pageId="68" pageNumber="464" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang et al. (2002)</bibRefCitation>
, the extension of the male thoracic sternal groove is comparable to the condition in
<taxonomicName id="4C424D4F3B7FE312FE0CFE54FDF39CF3" authorityName=", Alcock" authorityYear="1902" box="[458,624,413,437]" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B7FE312FE0CFE54FDF39CF3" box="[458,624,413,437]" italics="true" pageId="68" pageNumber="464">Lambdophallus</emphasis>
</taxonomicName>
but several differences such as movable eyes (versus immovable eyes in
<taxonomicName id="4C424D4F3B7FE312FF4BFE14FEB09CB3" authorityName=", Alcock" authorityYear="1902" box="[141,307,477,501]" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B7FE312FF4BFE14FEB09CB3" box="[141,307,477,501]" italics="true" pageId="68" pageNumber="464">Lambdophallus</emphasis>
</taxonomicName>
), the shapes of the third maxillipeds with the ischium expanded distally in
<taxonomicName id="4C424D4F3B7FE312FE16FE34FDDB9F53" authorityName="Huang, Hsueh &amp; Ng" authorityYear="2002" box="[464,600,509,533]" class="Malacostraca" family="Hexapodidae" genus="Latohexapus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B7FE312FE16FE34FDDB9F53" box="[464,600,509,533]" italics="true" pageId="68" pageNumber="464">Latohexapus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD34B3D3B7FE312FDA1FE34FF289F73" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" pageId="68" pageNumber="464" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang et al., 2002</bibRefCitation>
: fig. 4D) (versus slightly convex in
<taxonomicName id="4C424D4F3B7FE312FD85FDD4FE0E9F13" authority=", Manning &amp; Holthuis, 1981" authorityName=", Manning &amp; Holthuis" authorityYear="1981" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B7FE312FD85FDD4FD699F73" box="[579,746,541,565]" italics="true" pageId="68" pageNumber="464">Lambdophallus</emphasis>
,
<bibRefCitation id="EFD34B3D3B7FE312FFB7FDF4FE0E9F13" author="Manning RB &amp; Holthuis LB" box="[113,397,573,597]" pageId="68" pageNumber="464" pagination="1 - 379" refId="ref45783" refString="Manning RB &amp; Holthuis LB (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306: 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis, 1981</bibRefCitation>
</taxonomicName>
: fig. 34b), and the male abdominal somite 6 appearing hexagonal (
<bibRefCitation id="EFD34B3D3B7FE312FE05FD94FD139F33" author="Huang J-F &amp; Hsueh P-W &amp; Ng PKL" box="[451,656,605,629]" pageId="68" pageNumber="464" pagination="651 - 660" refId="ref45534" refString="Huang J-F, Hsueh P-W &amp; Ng PKL (2002) Crabs of the family Hexapodidae (Decapoda: Brachyura) from Taiwan, with description of a new genus and new species. Journal of Crustacean Biology, 22 (3): 651 - 660." type="journal article" year="2002">Huang et al., 2002</bibRefCitation>
: fig. 4G) (versus almost squarish in
<taxonomicName id="4C424D4F3B7FE312FE6AFDB4FE919FF3" authority=", Manning &amp; Holthuis, 1981" authorityName=", Manning &amp; Holthuis" authorityYear="1981" class="Malacostraca" family="Hexapodidae" genus="Lambdophallus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B936EADE3B7FE312FE6AFDB4FDDA9FD3" box="[428,601,637,661]" italics="true" pageId="68" pageNumber="464">Lambdophallus</emphasis>
,
<bibRefCitation id="EFD34B3D3B7FE312FDAAFDB4FE919FF3" author="Manning RB &amp; Holthuis LB" pageId="68" pageNumber="464" pagination="1 - 379" refId="ref45783" refString="Manning RB &amp; Holthuis LB (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306: 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis, 1981</bibRefCitation>
</taxonomicName>
: fig. 34C) easily distinguishes the two genera.
</paragraph>
</subSubSection>
</treatment>
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