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<document id="4443F03D4598F0E2D1EA9408F67C132E" ID-DOI="10.11646/zootaxa.1837.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5127540" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627042203922" checkinUser="felipe" docAuthor="Mader, Bryn J." docDate="2008" docId="03EB87C9FFB2DA6CEAFEFF42FE2A6CBF" docLanguage="en" docName="zt01837p085.pdf" docOrigin="Zootaxa 1837 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1837.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Metatelmatherium cristatum Granger and Gregory 1938" docType="treatment" docVersion="3" lastPageNumber="80" masterDocId="FFD2FFB1FFFDDA23EA69FFD5FFEB6F73" masterDocTitle="A species level revision of Bridgerian and Uintan brontotheres (Mammalia, Perissodactyla) exclusive of Palaeosyops" masterLastPageNumber="85" masterPageNumber="1" pageNumber="80" updateTime="1699015358681" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C7E34FE3FE426657D4DE0BC62B12BE4A">A species level revision of Bridgerian and Uintan brontotheres (Mammalia, Perissodactyla) exclusive of Palaeosyops</mods:title>
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<treatment id="03EB87C9FFB2DA6CEAFEFF42FE2A6CBF" ID-DOI="http://doi.org/10.5281/zenodo.5127480" ID-Zenodo-Dep="5127480" LSID="urn:lsid:plazi:treatment:03EB87C9FFB2DA6CEAFEFF42FE2A6CBF" httpUri="http://treatment.plazi.org/id/03EB87C9FFB2DA6CEAFEFF42FE2A6CBF" lastPageNumber="80" pageId="79" pageNumber="80">
<subSubSection id="C3586554FFB2DA6CEAFEFF42FBFB6FC2" box="[151,1040,151,177]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="8BFD36DFFFB2DA6CEAFEFF42FBFB6FC2" blockId="79.[151,1040,151,177]" box="[151,1040,151,177]" pageId="79" pageNumber="80">
<heading id="D0B581B3FFB2DA6CEAFEFF42FBFB6FC2" bold="true" box="[151,1040,151,177]" fontSize="11" level="1" pageId="79" pageNumber="80" reason="1">
<emphasis id="B936EACDFFB2DA6CEAFEFF42FBFB6FC2" bold="true" box="[151,1040,151,177]" pageId="79" pageNumber="80">
Species
<taxonomicName id="4C424D5CFFB2DA6CEA90FF42FBFB6FC2" authority="Granger and Gregory 1938" authorityName="Granger and Gregory" authorityYear="1938" box="[249,1040,151,177]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="cristatum">
<emphasis id="B936EACDFFB2DA6CEA90FF42FD5F6FC2" bold="true" box="[249,692,151,177]" italics="true" pageId="79" pageNumber="80">METATELMATHERIUM cristatum</emphasis>
<bibRefCitation id="EFD34B2EFFB2DA6CE8D5FF42FBFB6FC2" author="Granger, W. &amp; Gregory, W. K." box="[700,1040,151,177]" pageId="79" pageNumber="80" pagination="435 - 436" refId="ref44190" refString="Granger, W. &amp; Gregory, W. K. (1938) Addendum. A new titanothere genus from the upper Eocene of Mongolia and North America. In: Colbert, E. H., Fossil mammals from Burma in the American Museum of Natural History. Bulletin of the American Museum of Natural History, 74 (6), 435 - 436." type="journal article" year="1938">Granger and Gregory 1938</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586554FFB2DA6CEAFEFF37FD566E57" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="8BFD36DFFFB2DA6CEAFEFF37FD3B6F8F" blockId="79.[151,1436,226,972]" box="[151,720,226,252]" pageId="79" pageNumber="80">
<materialsCitation id="3B2A3C82FFB2DA6CEAFEFF37FD3B6F8F" box="[151,720,226,252]" collectionCode="AMNH" pageId="79" pageNumber="80" specimenCode="AMNH 26411" specimenCount="1" typeStatus="holotype">
<emphasis id="B936EACDFFB2DA6CEAFEFF37FEFB6F8F" bold="true" box="[151,272,226,252]" pageId="79" pageNumber="80">
<typeStatus id="54F9887DFFB2DA6CEAFEFF37FEE06F8F" box="[151,267,226,252]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBE49EA4FFB2DA6CEB7EFF37FE286F8F" box="[279,451,226,252]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="79" pageNumber="80">AMNH 26411</specimenCode>
, a skull and lower jaw.
</materialsCitation>
</paragraph>
<paragraph id="8BFD36DFFFB2DA6CEAAEFEDFFD566E57" blockId="79.[151,1436,226,972]" box="[199,701,266,292]" pageId="79" pageNumber="80">
<emphasis id="B936EACDFFB2DA6CEAAEFEDFFE2F6E57" bold="true" box="[199,452,266,292]" pageId="79" pageNumber="80">Referred specimens.</emphasis>
<typeStatus id="54F9887DFFB2DA6CEBA5FEDFFDED6E57" box="[460,518,266,292]" pageId="79" pageNumber="80">Type</typeStatus>
specimen only.
</paragraph>
</subSubSection>
<subSubSection id="C3586554FFB2DA6CEAAEFEE7FE8D6EB7" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="8BFD36DFFFB2DA6CEAAEFEE7FE8D6EB7" blockId="79.[151,1436,226,972]" pageId="79" pageNumber="80">
<emphasis id="B936EACDFFB2DA6CEAAEFEE7FEAC6E3F" bold="true" box="[199,327,306,332]" pageId="79" pageNumber="80">Diagnosis.</emphasis>
Species of
<taxonomicName id="4C424D5CFFB2DA6CEBB3FEE7FD596E38" authorityName="Granger and Gregory" authorityYear="1938" box="[474,690,306,331]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="B936EACDFFB2DA6CEBB3FEE7FD596E38" box="[474,690,306,331]" italics="true" pageId="79" pageNumber="80">Metatelmatherium</emphasis>
</taxonomicName>
, probably distinguished by an elongated and broadly sloping mandibular symphysis. The taxon may also be distinguished by a relatively high and wide posterior cusp on P1, relatively large P2, relatively thin cingula on P3 and P4, relatively broad occiput, and a relatively broad coronoid process.
</paragraph>
</subSubSection>
<subSubSection id="C3586554FFB2DA6CEAAEFE07FE2A6CBF" pageId="79" pageNumber="80" type="discussion">
<paragraph id="8BFD36DFFFB2DA6CEAAEFE07FF326DFF" blockId="79.[151,1436,226,972]" pageId="79" pageNumber="80">
<emphasis id="B936EACDFFB2DA6CEAAEFE07FEB96E9F" bold="true" box="[199,338,466,492]" pageId="79" pageNumber="80">Discussion.</emphasis>
Because the
<typeStatus id="54F9887DFFB2DA6CEB87FE07FDCB6E9F" box="[494,544,466,492]" pageId="79" pageNumber="80">type</typeStatus>
species of
<taxonomicName id="4C424D5CFFB2DA6CE8CCFE07FC966E98" authorityName="Granger and Gregory" authorityYear="1938" box="[677,893,466,491]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="B936EACDFFB2DA6CE8CCFE07FC966E98" box="[677,893,466,491]" italics="true" pageId="79" pageNumber="80">Metatelmatherium</emphasis>
</taxonomicName>
is an Asian taxon it is not properly within the scope of the present paper. Nevertheless, because it is the
<typeStatus id="54F9887DFFB2DA6CE92EFE2FFC926D67" box="[839,889,506,532]" pageId="79" pageNumber="80">type</typeStatus>
species of a taxon found in North America, it must be briefly discussed here. As noted above,
<bibRefCitation id="EFD34B2EFFB2DA6CE8B9FDF7FC5F6D4F" author="Mihlbachler, M. C." box="[720,948,546,572]" pageId="79" pageNumber="80" refId="ref45220" refString="Mihlbachler, M. C. (2005) Phylogenetic systematics of the Brontotheriidae (Mammalia, Perissodactyla). Columbia University, New York, 763 pp." type="book" year="2005">Mihlbachler (2005)</bibRefCitation>
has regarded this species as a junior synonym of
<taxonomicName id="4C424D5CFFB2DA6CEB6AFD9FFE676D10" baseAuthorityName="AMNH" baseAuthorityYear="2060" box="[259,396,586,612]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="ultimum">
<emphasis id="B936EACDFFB2DA6CEB6AFD9FFEF06D10" box="[259,283,586,611]" italics="true" pageId="79" pageNumber="80">M</emphasis>
.
<emphasis id="B936EACDFFB2DA6CEB45FD9FFE676D10" box="[300,396,586,611]" italics="true" pageId="79" pageNumber="80">ultimum</emphasis>
</taxonomicName>
from North America, but for the present, I prefer to continue to regard
<taxonomicName id="4C424D5CFFB2DA6CEE8DFD9FFA906D10" authorityName="Granger and Gregory" authorityYear="1938" box="[1252,1403,586,612]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="cristatum">
<emphasis id="B936EACDFFB2DA6CEE8DFD9FFB176D10" box="[1252,1276,586,611]" italics="true" pageId="79" pageNumber="80">M</emphasis>
.
<emphasis id="B936EACDFFB2DA6CEF65FD9FFA906D10" box="[1292,1403,586,611]" italics="true" pageId="79" pageNumber="80">cristatum</emphasis>
</taxonomicName>
as valid.
</paragraph>
<paragraph id="8BFD36DFFFB2DA6CEAAFFD4FFE2A6CBF" blockId="79.[151,1436,226,972]" pageId="79" pageNumber="80">
The type specimen of
<taxonomicName id="4C424D5CFFB2DA6CEBBFFD4FFCC36DC0" authorityName="Granger and Gregory" authorityYear="1938" box="[470,808,666,691]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="cristatum">
<emphasis id="B936EACDFFB2DA6CEBBFFD4FFCC36DC0" box="[470,808,666,691]" italics="true" pageId="79" pageNumber="80">Metatelmatherium cristatum</emphasis>
</taxonomicName>
was collected by the American Museum of Natural History's Central Asiatic Expedition of 1930 from the Camp Margetts Area (Huhebolhe Cliff),
<quantity id="4CBA9B3AFFB2DA6CEF5AFD17FA706DAF" box="[1331,1435,706,732]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="79" pageNumber="80" unit="mi" value="25.0">25 miles</quantity>
(approximately
<quantity id="4CBA9B3AFFB2DA6CEB38FD3FFE776C77" box="[337,412,746,772]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="79" pageNumber="80" unit="km" value="40.0">40 km</quantity>
) south-southwest of Erhlien (Iren Dabasu) in the Inner Mongolian Autonomous Region (
<bibRefCitation id="EFD34B2EFFB2DA6CEAF6FCC7FEB56C5F" author="Li, C. &amp; Ting, S." box="[159,350,786,812]" pageId="79" pageNumber="80" pagination="1 - 93" refId="ref44581" refString="Li, C. &amp; Ting, S. (1983) The Paleogene mammals of China. Bulletin of the Carnegie Museum of Natural History, 21, 1 - 93." type="journal article" year="1983">Li &amp; Ting 1983</bibRefCitation>
, p. 25). Since that time no other specimen of
<taxonomicName id="4C424D5CFFB2DA6CE915FCC7FBF96C58" authorityName="Granger and Gregory" authorityYear="1938" box="[892,1042,786,812]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="cristatum">
<emphasis id="B936EACDFFB2DA6CE915FCC7FC7F6C58" box="[892,916,786,811]" italics="true" pageId="79" pageNumber="80">M</emphasis>
.
<emphasis id="B936EACDFFB2DA6CE9CDFCC7FBF96C58" box="[932,1042,786,811]" italics="true" pageId="79" pageNumber="80">cristatum</emphasis>
</taxonomicName>
has been reported although, several authors (
<bibRefCitation id="EFD34B2EFFB2DA6CEB46FCEFFE576C27" author="Chow, M. M." box="[303,444,826,852]" pageId="79" pageNumber="80" pagination="201 - 214" refId="ref43758" refString="Chow, M. M. (1957) On some Eocene and Oligocene mammals from Kwangsi and Yunnan. Vertebrata Palasiatica, 1 (3), 201 - 214." type="journal article" year="1957">Chow 1957</bibRefCitation>
;
<bibRefCitation id="EFD34B2EFFB2DA6CEBA1FCEFFD596C27" author="Tang, X. &amp; Chow, M." box="[456,690,826,852]" pageId="79" pageNumber="80" pagination="119 - 133" refId="ref46293" refString="Tang, X. &amp; Chow, M. (1964) A review of vertebrate-bearing Lower Tertiary of south China. Vertebrata Palasiatica, 8 (2), 119 - 133." type="journal article" year="1964">Tang &amp; Chow 1964</bibRefCitation>
;
<bibRefCitation id="EFD34B2EFFB2DA6CE8D7FCEFFC946C27" author="Tang, Y. &amp; You, Y. &amp; Xu, Q. &amp; Qiu, Z. &amp; Hu, Y." box="[702,895,826,852]" pageId="79" pageNumber="80" pagination="279 - 292" refId="ref46327" refString="Tang, Y., You, Y., Xu, Q., Qiu, Z. &amp; Hu, Y. (1974) The lower Tertiary of the Bose and Yongle Basins of the Guangxi Zhuang Autonomous Region. Vertebrata Palasiatica, 12 (4), 279 - 292." type="journal article" year="1974">
Tang
<emphasis id="B936EACDFFB2DA6CE896FCEFFCD96C20" box="[767,818,826,851]" italics="true" pageId="79" pageNumber="80">et al</emphasis>
. 1974
</bibRefCitation>
;
<bibRefCitation id="EFD34B2EFFB2DA6CE9E2FCEFFBA16C27" author="Ding, S. &amp; Zheng, J. &amp; Zhang, Y. &amp; Tong, Y." box="[907,1098,826,852]" pageId="79" pageNumber="80" pagination="35 - 45" refId="ref43921" refString="Ding, S., Zheng, J., Zhang, Y. &amp; Tong, Y. (1977) The age and characteristics of the Liuniu and the Dongjun faunas, Bose Basin of Guangxi. Vertebrata Palasiatica, 15 (1), 35 - 45." type="journal article" year="1977">
Ding
<emphasis id="B936EACDFFB2DA6CE9A5FCEFFC146C20" box="[972,1023,826,851]" italics="true" pageId="79" pageNumber="80">et al</emphasis>
. 1977
</bibRefCitation>
) have referred material from the
<collectingRegion id="4986F83DFFB2DA6CEAAAFCB7FE726C0F" box="[195,409,866,892]" country="China" name="Guangxi" pageId="79" pageNumber="80">Guangxi Province</collectingRegion>
of southern
<collectingCountry id="F355764FFFB2DA6CE859FCB7FD9D6C0F" box="[560,630,866,892]" name="China" pageId="79" pageNumber="80">China</collectingCountry>
to
<taxonomicName id="4C424D5CFFB2DA6CE8F7FCB7FCF36C08" box="[670,792,866,892]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="browni">
<emphasis id="B936EACDFFB2DA6CE8F7FCB7FD5D6C08" box="[670,694,866,891]" italics="true" pageId="79" pageNumber="80">M</emphasis>
.
<emphasis id="B936EACDFFB2DA6CE8AFFCB7FCF36C08" box="[710,792,866,891]" italics="true" pageId="79" pageNumber="80">browni</emphasis>
</taxonomicName>
(here regarded as a nomen dubium) or
<taxonomicName id="4C424D5CFFB2DA6CEE98FCB7FEEC6CD7" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="undefined-i">
<emphasis id="B936EACDFFB2DA6CEE98FCB7FF256CD0" italics="true" pageId="79" pageNumber="80">Metatelmatherium</emphasis>
sp. I
</taxonomicName>
am not certain whether this material represents the genus
<taxonomicName id="4C424D5CFFB2DA6CE9C3FC5FFB696CD0" authorityName="Granger and Gregory" authorityYear="1938" box="[938,1154,906,931]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="B936EACDFFB2DA6CE9C3FC5FFB696CD0" box="[938,1154,906,931]" italics="true" pageId="79" pageNumber="80">Metatelmatherium</emphasis>
</taxonomicName>
but, if it does, it may be referable to
<taxonomicName id="4C424D5CFFB2DA6CEB4DFC67FE526CB8" authorityName="Granger and Gregory" authorityYear="1938" box="[292,441,946,972]" class="Mammalia" family="Brontotheriidae" genus="Metatelmatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="cristatum">
<emphasis id="B936EACDFFB2DA6CEB4DFC67FED76CB8" box="[292,316,946,971]" italics="true" pageId="79" pageNumber="80">M</emphasis>
.
<emphasis id="B936EACDFFB2DA6CEB22FC67FE526CB8" box="[331,441,946,971]" italics="true" pageId="79" pageNumber="80">cristatum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>